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14TAXONOMIC AND NOMENCLATURAL NOTES ON CLINOPODIUM L. AND ZIZIPHORA L. (LAMIACEAE)
D. G. Melnikov
Komarov Botanical Institute of the Russian Academy of Sciences Professora Popova Str., 2, St. Petersburg, 197376, Russia DMelnikov@binran.ru
Nomenclatural combinations for three species: Clinopodium marifolium (Cav.) Melnikov, C. canescens (J. Presl) Melnikov, C. yakusimense (Masam.) Melnikov, and one variety C. multicaule (Maxim.) Kuntze var. shibetchense (H. Lev.) Melnikov are published. Replacement names for C. repens (Buch.-Ham. ex D. Don) Benth. and Ziziphora alpina (L.) Melnikov are given. The lec-totypes of Melissa alba Waldst. et Kit. and Calamintha heterotricha Boiss. et Reut., as well as the lectotype and the epitype of Nepeta marifolia Cav. are designated.
Keywords: new nomenclature combination, replacement name, lectotype, epitype.
Ongoing work on the revision of the genus Clinopodium L. made it necessary to publish some nomenclatural novelties.
1. Genus Clinopodium L.
In a previous publication (Melnikov, 2015), I published the replacement name Clinopodium hispanicum Melnikov for C. fruticosum (L.) Kuntze, nom. illeg. Now I treat this taxon as conspecific to a species originally published as Nepeta marifolia Cav. This requires a new combination in the genus Clinopodium.
Clinopodium marifolium (Cav.) Melnikov, comb. nova. = Nepeta marifolia Cav. 1800, Anales Ci. Nat. 2: 192, tab. 22. = Micromeria marifolia (Cav.) Benth. 1834, Labiat. Gen. Spec.: 382. = Melissa marifolia (Cav.) Pers. 1806, Syn. Pl. 2: 132. = Thymus marifolius (Cav.) Willd. 1809, Enum. Pl.: 624. = Calamintha marifolia (Cav.) Steud. 1840, No-mencl. Bot., ed. 2, 1: 251.
Described from Spain (Valencia): «Se cria con abundancia en los montes asperos de la Hoya de Castella, en el Carrascal de Alcoy, en Ma-riola, Penaglosa, Morella y otras partes».
Lectotype (Melnikov, designated here): [Icon:] Cavanilles, 1800: tab. 22. — Fig.
Epitype (Melnikov, designated here): «Fl. Hispanica: Valencia; Sierra de Espadan. 1500 m. VII [18]91. E. Reverchon (Dr. C. Bae-niz, Herbarium Europaeum, Micromeria marifolia Benth. s. n.)» (LE 01016338!). — Plate I.
Fig. The lectotype of Nepeta marifolia Cav.: Tab. 22 in Cavanilles (1800). Image from the Biodiversity Heritage Library (www.biodiversitylibrary.org). Digitized by Missouri Botanical Garden, Peter H. Raven Library.
= Melissa fruticosa L. 1753, Sp. Pl.: 593. = Clinopodium fruticosum (L.) Kuntze, 1891, Revis. Gen. Pl. 2: 516, nom. illeg., non Forssk. 1775, Fl. Aegypt.-Arab.: 107. = Satureja fruticosa (L.) Briq. 1898, Annuaire Conserv. Jard. Bot. Genève, 2: 192. = Micromeria fruticosa (L.) Druce, 1914, Bot. Exch. Club Soc. Brit. Isles, 3: 421. = Clinopodium serpyllifoli-um (M. Bieb.) Kuntze subsp. fruticosum (L.) Bràuchler, 2006, Taxon, 55: 980. = C. hispanicum Melnikov, 2015, Turczaninowia, 18(3): 109.
Among taxonomic synonyms of Clinopodium thymifolium (Scop.) Kuntze, I put Melissa alba Waldst. et Kit. (1811, Descr. Icon. Pl. Hung. 3: 227; = Calamintha officinalis Moench var. alba (Waldst. et Kit.) Nyman, 1881, Consp. Fl. Eur.: 588) described from Croatia: «Habitat in rupibus calcareis provinciae Liccanae Croatiae ad Plocha, pone Divo-Szello, atque in subalpinis et in ipsis alpibus montium Velebich. Floret a medio Julio in Augustum».
Lectotype of Melissa alba (Melnikov, designated here): «D. 68. — ad Plocham, in alpinis et subalpinis Velebit» (BP, № 8087).
Stored in the Herbarium LE is an original specimen (LE 01016435!) collected by F. Waldstein in Hungary in 1806, determined by him as Melissa alba, and provided with the authors' label containing the description of the species.
I consider Clinopodium raimondoi Spadaro described from Sicily con-specific with Calamintha canescens J. Presl described from the same island. Thence, I am publishing a new nomenclatural combination in the genus Clinopodium.
Clinopodium canescens (J. Presl) Melnikov, comb. nova. = Calamintha canescens J. Presl, 1826, Fl. Sicul.: 37. = Calamintha nepeta (L.) Savi var. canescens (J. Presl) Nyman, 1881, Consp. Fl. Eur.: 588. = Calamintha glandulosa (Req.) Benth. var. canescens (J. Presl) Silic, 1979, Monogr. Satureja Fl. Jugosl.: 142.
= Clinopodium raimondoi Spadaro, A. S. Faqi et Mazzola, 2016, in Fl. Medit. 25 (Spec. Iss.): 311, syn. nov.
Clinopodium heterotrichum (Boiss. et Reut.) Govaerts, 1999, World Checkl. Seed Pl. 3(1): 17. = Calamintha heterotricha Boiss. et Reut. 1852, Pugill. Pl. Afr. Bor. Hispan.: 93.
Described from Algeria: «Hab. in collibus propè Alger, Constantine, Bougie (Bové! et alii)».
Lectotype (Melnikov, designated here): «Alger et Bougie. Collines, XII 1839, Bové» (LE 01017615!; isolectotype — G 00402201).
Syntypes: «[Algérie] Constantine. Collines, X 1838, Bové, N.» (G 00424763), «Alger et Bougie. Collines, XII 1839, Bové, N.» (G 00424764), «Constantine. Collines. X 1838, Bové» (LE 01017618!, LE 01017619!).
Clinopodium multicaule (Maxim.) Kuntze var. shibetchense
(H. Lév.) Melnikov, comb. nova. = Calamintha umbrosa (M. Bieb.) Rchb. var. shibetchensis H. Lév. 1911, Repert. Spec. Nov. Regni Veg. 9: 322. = Clinopodium chinense Benth. var. shibetchense (H. Lév.) Koidz. 1929, Bot. Mag. (Tokyo), 43: 387. = Clinopodium umbrosum (M. Bieb.) K. Koch var. shibetchensis (H. Lév.) McKean, 1982, Notes Roy. Bot. Gard. Edinburgh, 40: 161.
Clinopodium yakusimense (Masam.) Melnikov, comb. nova. ^ Satureja yakusimensis Masam. 1930, J. Soc. Trop. Agric. 2: 35. = Clinopodium multicaule (Maxim.) Kuntze var. yakusimense (Masam.) Yahara, 1987, J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 14: 107.
Since the combination Clinopodium repens (Buch.-Ham. ex D. Don) Benth. is an illegitimate later homonym, I am giving it the replacement name Clinopodium hamiltonii.
Clinopodium hamiltonii Melnikov, nom. nov. = Thymus repens Buch.-Ham. ex D. Don, 1825, Prodr. Fl. Nepal.: 113. = C. repens (Buch.-Ham. ex D. Don) Benth. 1830, in N. Wallich, Pl. Asiat. Rar. 1: 66, nom. illeg., non Vell. 1829, Fl. Flumin.: 242.
2. Genus Ziziphora L.
When transferring the species of the genus Acinos to Ziziphora and publishing appropriate combinations (Melnikov, 2016), I missed an earlier homonym Ziziphora alpina Mill. (Miller, 1768) based on the other type than Thymus alpinus L. Now I am publishing the replacement name in the genus Ziziphora for this species.
Ziziphora alpicola Melnikov, nom. nov. = Thymus alpinus L. 1753, Sp. Pl.: 591. = Acinos alpinus (L.) Moench, 1794, Methodus: 407. = Ziziphora alpina (L.) Melnikov, 2016, Bot. Zhurn. [St. Petersburg], 101(1): 89, nom. illeg., non Z. alpina Mill. 1768, Gard. Dict., ed. 8, 3: sine pag.
References
Cavanilles A. J. 1800. Polvos vegetales contra la rabia, y descripcion de las plantas que los suministran (Polvos contra la rabia) II Anal. Hist. Nat. T. 2, № 4. P. 178-195.
Melnikov D. G. 2015. New sections of the genus Clinopodium L. (Lamiaceae) and their synopsis // Turczaninowia. Т. 18, № 3. P. 103-112. [In Russian with English abstract] (Мельников Д. Г. 2015. Новые секции рода Clinopodium L. (Lamiaceae) и их конспект // Turczaninowia. Т. 18, № 3. С. 103-112). doi: 10.14258/turczaninowia.18.3.6 Melnikov D. G. 2016. On the taxonomic status of the genus Acinos (Lamiaceae) // Bot. Zhurn. Vol. 101, № 1. P. 80-94. [In Russian with English abstract] (Мельников Д. Г. 2016. О таксономическом статусе рода Acinos (Lamiaceae) // Ботан. журн. Т. 101, № 1. С. 80-94). Miller Ph. 1768. The gardeners dictionary: Containing the best and newest methods of cultivating and improving the kitchen, fruit, flower garden, and nursery... Ed. 8. London. S. p.
СИСТЕМАТИЧЕСКИЕ И НОМЕНКЛАТУРНЫЕ ЗАМЕТКИ О РОДАХ CLINOPODIUML. И ZIZIPHORA L. (LAMIACEAE)
Д. Г. Мельников
Ботанический институт им. В. Л. Комарова РАН Гербарий высших растений ул. Профессора Попова, 2, Санкт-Петербург, 197376, Россия DMelnikov@binran.ru
Продолжающаяся работа над ревизией рода Clinopodium L. потребовала публикации некоторых таксономических и номенклатурных дополнений. Приводятся новые номенклатурные комбинации для трех видов: Clinopodium marifolium (Cav.) Melnikov, C. canescens (J. Presl) Melnikov, C. yakusimense (Masam.) Melnikov — и одной разновидности C. multicaule (Maxim.) Kuntze var. shibetchense (H. Lev.) Melnikov. Для C. repens (Buch.-Ham. ex D. Don) Benth. и Ziziphora alpina (L.) Melnikov обнародованы заменяющие названия. Выбраны лектотипы названий Melissa alba Waldst. et Kit. и Calamintha heterotricha Boiss. et Reut., а также лектотип и эпитип Nepeta marifolia Cav.
Ключевые слова: новые номенклатурные комбинации, заменяющие названия, лектотип, эпитип.
