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ЭКОЛОГИЯ ЖИВОТНЫХ
УДК 595.762.12(47-12)
СРАВНИТЕЛЬНЫЙ АНАЛИЗ ВИДОВЫХ СОСТАВОВ ЖУЖЕЛИЦ ПРИБРЕЖНЫХ И ОСТРОВНЫХ ЭКОСИСТЕМ ЗАПАДНОГО КАСПИЯ
©2011 Абдурахманов Г.М., Нахибашева Г.М, Клычева С.М, Эльдерханова З.М.
Дагестанский государственный университет
Впервые изучен видовой состав жужелиц прибрежных и островных экосистем Западного Каспия. Приводится сравнительный анализ видового состава жужелиц прилегающих территорий.
For the first time studied the species composition of ground beetles of coastal and island ecosystems of the Western Caspian. The article provides a comparative analysis of species composition of ground beetles and adjacent areas.
Ключевые слова: жужелицы, прибрежные и островные территории, видовой состав.
Keywords: carabids, a coastal and an island territories, species composition.
Как известно, процессы флоро- и фауногенеза на определенных территориях протекают под воздействием, общих для них, экологических факторов. Иными словами, на определенных территориях в сообществах протекают своеобразные, свойственные им, характеризующие их процессы, хотя некоторые исследователи не всегда приходят к близким результатам в оценке путей становления фаун.
Разрушение сложившегося сообщества может быть следствием воздействия различных факторов, особенно климатических или обусловленных орографическими процессами. Следует подчеркнут, что контрастность смен условий в рассматриваемом регионе не была выражена резко на протяжении последних геологических эпох. Тем не менее, многократно повторяющиеся смены трансгрессий и регрессий Каспийского моря вызвали смены растительного мира и животного населения. Общим следствием проявления этой тенденции явилось постепенное обеднение третичного лесного энтомоценоза и замещение его элементами и целыми комплексами фаун аридных областей Европейской, Сибирской, Туран-ской, Средиземноморской и Переднеазиатской провинций.
Сравнение видового состава жужелиц прибрежных и островных территорий различных районов Западного Каспия приводится в табл. 1.
Таблица 1
Видовой состав жужелиц прибрежных экосистем и островов Западного Каспия
№ Видовой состав Прибрежные экосистемы Острова
Астраханская область Республика Калмыкия Республика Дагестан Республика Азербайджан Нордовый Тюлений Чечень
Брянская коса Крайновка Терско-Сунженский район Бархан Сарыкум Кизлярский район Каякент, Вихри Берикей, Самур Северный Азербайджан Апшеронский полуостров Талыш
Family CARABIDAE Subfamily CICINDELI-NAE
1. Genus MEGACEPHALA Latr. 1802
Subgenus Grammognatha Motsch. 1850
1. euphratica Dejean 1822 +
2. Genus CICINDELA Linnaeus 1758
Subgenus Eumecus Lin-
naeus 1758
2. germanica Linnaeus 1758 + + + + + + + + + + + + +
Subgenus Myriochile De-
jean 1825
3. orientalis Dejean 1825 + + + + +
4. melancholica Fabricius 1798 + + + + + +
Subgenus Cephalota Pal-
las 1776
5. atrata Pallas 1776 + +
6. besseri Dejean 1826 +
7. chiloleuca Fischer von Waldheim 1820 + + + + +
8. deserticola Faldermann + + + + +
1836
9. elegans Fischer von Waldheim 1822 + + + +
Subgenus Cicindina
Schrank 1781
10. arenaria viennensis Schrank 1781 +
11. contorta Fischer von Waldheim 1828 + + + + + + +
12. inscripta Zoubkoff 1833 + + +
13. sublacerata sublacerata Solsky 1874 + + + + +
14. litterifera Chaudoir 1842 + + +
Subgenus Lophiridia
Dokhtouroff 1887
15. littoralis Dokhtouroff 1887 + + + + + + + +
16. caucasica Adams 1817 +
17. fischeri Adams 1817 + + +
Subgenus Cicindela
Fischer von Waldheim
1825
18. asiatica Audoin et Brulle 1839 +
19. campestris pontica Fischer von Waldheim 1825 + + + + + + +
20. desertorum Dejean 1825 +
21. hybrida sahlbergi Fischer von Waldheim 1824 + +
22. lacteola lacteola Pallas 1776 + + +
23. maritima kirgisica Mandl. 1936 + +
24. coerulea lucifera Mandl. 1971 + +
25. monticola Menetries 1832 +
26. talychensis Chaudoir 1846 +
3. Genus OMOPHRON Fabricius 1776
Subgenus Omophron
Fabricius 1776
27. limbatus Fabricius 1776 + +
4.Genus LEISTUS Frolich 1799
28. fulvus Chaudoir 1846 + +
29. lenkoranus Reitter 1885 + +
5. Genus NEBRIA Latrelle 1825
Subgenus Eunebria Jean-
nel 1937
30. nigerrima Chaudoir 1846 + +
31. picicornis. luteipes Chaudoir 1850 + + +
Subgenus Nebria Latrelle 1825
32. brevicollis Fabricius. 1792 + +
Subgenus Alpaeus Bonelli 1809
33. faldermani Menetries 1832 +
6. Genus NOTIOPHILUS Chaudoir 1850
Subgenus Notiophilus Chaudoir 1850
34. laticollis Chaudoir 1850 + +
35. palustris Duftschmid 1812 + + +
36. germinyi Fauv. 1863 +
37. aestuans Motschulsky 1864 + +
Subgenus Makarovius Barsevskis 1994
38. rufipes Curtis 1829 + +
Subgenus Latviaphilus Barsevskis 1994
39. biguttatus Fabricius 1779 + +
7. Genus CALOSOMA Linnaeus 1758
Subgenus Calosoma Linnaeus 1758
40. sycophanta Linnaeus 1758 + + + + + +
Subgenus Acalosoma Linnaeus 1758
41. inquisitor inquisitor Linnaeus 1758 + + + + + +
Subgenus Campalita Herbst 1784
42. auropunctatum Herbst 1784 + + + + +
43. maderae tectum Motschulsky 1846 + + +
Subgenus Caminara Gebler 1833
44. denticolle Gebler 1833 + + + + + + + + + +
45. imbricatum deserticola Semenov 1896 + +
46. reitteri Roeschke 1897 +
Subgenus Charmosta Illiger 1798
47. investigator Illiger 1798 + + +
8. Genus CARABUS Mannerheim 1827
Subgenus Carabus Linnaeus 1758
48. granulatus granulatus Linnaeus 1758 + + + +
49. sculpturatus Menetries 1832 +
Subgenus Trachycarabus Fischer von Waldheim 1823
50. estreicheri Fischer von Waldheim 1822 + +
51. campestris Fischer von Waldheim 1822 +
Subgenus Limnocarabus Linnaeus 1761
52. clathratus clathratus Linnaeus 1761 + + + + + +
Subgenus Procerus Dejean 1826
53. caucasicus Adams 1817 + +
Subgenus Megodontus Solier 1848
54. exaratus Quensel 1806 + + +
Subgenus Sphodristoca-
rabus Gehin 1885
55. adamsi hollbergi Mannerheim 1827 + + +
56. bohemanni Menetries 1832 +
Subgenus Chaetocarabus
Thomson 1875
57. intricatus Linnaeus 1761 +
Subgenus Lamprostus Motschuksky 1865
58. prasinescens Deuve 1994 +
Subgenus Tomocarabus
Fischer von Waldheim
1823
59. bessarabicus concretus Fischer von Waldheim 1823 + +
Subgenus Pachystus Motschulsky 1847
60. hungaricus mingens Quens 1806 + +
Subgenus Pachycarabus
Genhin 1876
61. staehlini Adams 1817 +
Subgenus Procrustes Bo-
nelli 1809
62. clypeatus Adams 1817 +
63. talyshensis Menetries 1832 +
9. Genus BLETHISA Zoubkoff 1829
64. eschscholtzi Zoubkoff 1829 +
10. Ge- ELAPHRUS Duftschmid
nus 1812
Subgenus Neoelaphrus Duftschmid 1812
65. cupreus Duftschmid 1812 + +
66. uliginosus Fabricius 1775 + +
Subgenus Elaphrus Se-
menov 1926
67. hypocrita Semenov 1926 +
68. riparius Linnaeus 1758 + +
11.Genus LORICERA Fabricius 1775
Subgenus Loricera
Fabricius 1775
69. pilicornis Fabricius 1775 +
12.Genus SIAGONA Latreille 1804
70. europaea Dejean 1826 + + + + + +
13.Genus SCARITES Chaudoir 1855
Subgenus Scarites
Chaudoir 1855
71. salinus Dejean 1859 + + + + + + + +
72. eurytus Fischer von Waldheim 1825 +
Subgenus Paradistichus
Makarov 2006
73. angustus Chaudoir 1855 + + + + +
Subgenus Parralelomor-
fus Motsch. 1850
74. terricola terricola Bonelli 1813 + + + + + + + + + + + + + +
Subgenus Scallophorites Pallas 1776
75. bucida Pallas 1776 + + + +
Subgenus Distichus Motsch. 1858
76. planus Bonelli 1813 + + + + + + + + + + +
14.Genus CLIVINA Herbst 1784
77. collaris Herbst 1784 + + + + + + + +
78. fossor Linnaeus 1758 + + + + + + + + + + + + +
79. laevifrons Chaudoir 1842 + + + + + + + + + +
80. ypsilon Dejean 1830 + + + + + + + + + + + +
15.Genus DYSCHIRIUS Stephens 1827
81. arenosus Stephens 1827 + +
82. humeratus Chaudoir 1850 + + +
83. humiolcus Chaudoir 1850 + + + +
84. zimini Znojko 1928 +
85. sp. +
86. sp. +
87. sp. +
16.Genus DYSCHIRIODES Herbst 1783
88. globosus Herbst 1783 + + +
89. dimidiatus osseticus Fedo-renko 1993 +
90. ferganensis ferganensis Znojko 1930 + + +
91. lafertei lafertei Putzeys 1846 + +
92. euxinus Znojko 1927 + +
93. rufimanus Fleischer 1898 +
94. strumosus Erichson 1837 + + + + +
95. tristis Stephens 1828 + +
96. chalceus Erichson 1937 + + + + + +
97. chalybeus gibbifrons Apf. 1899 + + +
98. cylindricus Fleischer 1898 + + + + + +
99. pusillus Dejean 1825 + + + + + + +
100. salinus striatopunctatus Putzeys 1846 + + + + + +
101. chalybeus Putz. 1846 +
102. impunctipennis Dawson 1854 +
103. neresheimeri Wagner 1915 +
104. nitidus nitidus Dejean 1825 + + +
105. politus politus Dejean 1825 +
106. aeneus aeneus Dejean 1825 + + + +
107. apicalis Putzeys 1846 +
108. auriculatus Wollaston 1867 +
109. scriptifrons Fleischer 1898 + +
110. luticola Chaudoir 1850 + + +
111. microthorax Motschulsky 1844 + +
112. persicus Fedonenko 1994 +
113. substriatus priscus Muller 1922 +
114. rufipes Dejean 1825 +
115. nigricornis Motsch. 1844 +
116. unisetosus +
17.Genus BROSCUS Panzer 1813
117. asiaticus Ballion 1870 +
118. semistriatus Dejean 1828 + + + + + + + + + +
119. karelini Zoubkoff 1837 +
18.Genus APOTOMUS Dejean 1825
120. testaceus Dejean 1825 + + + + + + +
121. rufithorax Pecchioli 1838 +
19.Genus PERILEPTUS Schaum 1860
Subgenus Perileptus Schaum 1860
122. areolatus Creutzer 1799 +
20.Genus TRECHOBLEMUS Ganglbauer 1896
123. micros Herbst 1784 +
21.Genus BLEMUS Dejean 1821
124. discus Fabricius 1792 +
22.Genus EPAPHIUS Stephens 1927
125. secalis Paykull 1790 +
23.Genus TRECHUS Schrank 1781
Subgenus Trechus Clairv. 1806
126. quadristriatus Schrank 1781 + + + + + + + +
127. quadrimaculatus Mot-schulsky 1850 + + +
128. liopleurus Chaudoir 1850 + + + + +
129. taghizadehi Morvan 1974 +
130. fasciatus Motschulsky 1851 +
24.Genus TACHYS Csiki 1928
Subgenus Tachys Csiki 1928
131. lencoranus Csiki 1928 + +
132. scutellaris Stephens 1829 + + + + +
133. vittatus Motschulsky 1850 + + + + + + +
25.Genus PARATACHYS Duftschmid 1812
134. bistriatus Duftschmid 1812 + + + + + + + +
135. centriustatus Reitter 1874 + +
136. fulvicollis Dejean 1831 + + + + + +
137. micros Fischer von Vald-heim 1828 + + + + + + +
138. turkestanicus Csiki 1928 + + + + + + + +
26.Genus ELAPHROPUS Motschulsky 1839
Subgenus Sphaerotachys J. Muller 1926
139. haemorroidalis Ponza 1805 + + + + +
Subgenus Tachyura Mots. 1862
140. diabrachys Kolenati 1845 +
27.Genus TACHYTA Kirby 1837
Subgenus Tachyta Kirby 1837
141. nana Gyllenhal 1810 + + + +
28.Genus ASAPHIDION Linnaeus 1761
142. flavipes Linnaeus 1761 + + +
143. austriacum Schweiger 1975 + + +
144. pallipes Duftschmid 1812 +
145. caraboides Schrank 1781 +
29.Genus OCYS Steph. 1828
Subgenus Oreoces Peye-rimhoff 1908
146. trechoides Reitter 1895 +
30.Genus BEMBIDION Motschulsky 1844
Subgenus Microserrullula Motschulsky 1844
147. quadricolle Motschulsky 1844 + + + +
Subgenus Bracteon Ahrens 1812
148. argenteolum Ahrens 1812 + +
149. velox Linnaeus 1761 + + +
Subgenus Odontonium Fabricius 1792
150. striatum Fabricius 1792 + +
Subgenus Metallina Herbst 1784
151. lampros Herbst 1784 + + + + +
152. properans Stephens 1829 + + + +
Subgenus Asioperyphus Vysoky 1986
153. lunatum Dustschmid 1812 +
154. amnicola J. Sahlberg 1900 +
155. kazakhstanicum Kryzha-novskij 1979 + + + + +
156. ustum Quensel 1806 + + + + + + + +
Subgenus Chlorodium Motsch. 1864
157. luteipes Motsch. 1844 +
Subgenus Princidium Motschulsky 1864
158. punctulatum Drapiez 1820 +
159. bracteonoides Reitter 1908 +
Subgenus Notaphus
160. varium Olivier 1795 + + + + + + + + + + + + +
Subgenus Eupetodromus Thunberg 1787
161. dentellum Thunberg 1787 + + +
Subgenus Philochtus Fabricius 1779
162. biguttatum Fabricius 1779 + +
163. judaicum J. Shahlberg 1907 +
164. inoptatum Schaum 1857 + + + + + +
165. lunulatum Geoffr. 1785 + +
166. zaitzevi Lutshnik 1937 + +
Subgenus Emphanes Dalla Torre 1877
167. latiplaga Chaudoir 1850 + + + + + + + + + + +
168. minimum Fabricius 1792 + + + + +
169. tenellum Erichson 1837 + + + + + + + + + + + +
170. rivulare Dejean 1831 + + + +
171. normannum Dejean 1831 + + + + + +
172. quadriplagiatum Motsch. 1844 + + + +
Subgenus Talanes Germar 1812
173. aspericolle Germar 1812 + + + + + + + + +
Subgenus Leja Panzer 1796
174. articulatum Panzer 1796 + + + + + +
175. octomaculatum Goeze 1777 + + + + + + +
Subgenus Semicampa Chaudoiri 1850
176. chaudoiri Chaudoiri 1850 +
177. dagestanum Jedlicka 1962 + + + + + + +
178. heydeni Gangl. 1891 + + + +
179. semicampa +
Subgenus Diplocampa Gyllenhal 1810
180. assimile Gyllenhal 1810 + + + + + + + +
181. bisulcatum Chaudoir 1844 + +
182. fumigatum Duftschmid 1812 + + + + +
Subgenus Bembidion Linnaeus 1761
183. quadrimaculatum Linnaeus 1761 + + + + +
184. quadripustulatum Serville 1821 + + + + + + + + +
185. sp. +
Subgenus Notaphocampa Netolitzky 1914
186. niloticum hamatum Kolena-ti 1845 + + + +
Subgenus Bembidioneto-litzkya Strand 1929
187. tibiale Duftschmid 1812 +
188. fasciolatum Duft. 1812 +
189. astrabadense Mennerheim 1844 +
190. relictum Apfelbeck 1904 +
191. peliopterum Chaudoir 1850 + +
192. gotschii Chaudoir 1846 +
Subgenus Euperyphus Jeannel 1941
193. combustum Menetries 1832 + +
Subgenus Ocydromus Muller 1830
194. multipunctatum Motschulsky 1850 +
195. decorum Zenker in Panzer 1801 +
196. xanthomum Chaudoir 1850 + + +
Subgenus Peryphus Dejean 1821
197. persicum Menetries 1832 + + + + +
198. atlanticum Wollaston 1854 +
199. andreae Fabricius 1787 +
200. subcostatum Motschulsky 1850 + + + +
201. femoratum Sturm 1825 +
202. distinguendum lindrothi De Monte 1957 + +
203. parallelipenne Chaudoir 1850 + + +
204. testaceum Duft. 1812 +
Subgenus Peryphanes Jeann. 1941
205. fraxator Menetries 1832 + +
206. lirykense Reitter 1908 +
Subgenus Terminophanes Mull.-Motzf. 1998
207. pulcherrimum Motsch. 1850 + +
Subgenus Notaphem-phanes Net. 1920
208. ephippium Marsh 1802 + + + + + +
Subgenus Phyla Motsch. 1844
209. obtusum Serville 1821 + +
Subgenus Nepha Motschulsky 1864
210. tetragrammum Chaudoir 1846 + +
211. tetrasemum Chaudoir 1846 + +
Subgenus Synechostictus Motsch. 1864
212. multisulcatum Reitter 1890 +
31.Genus CARDIODERUS Fischer von Waldheim 1823
213. chloroticus Fischer von Waldheim 1823 + + +
32.Genus POGONUS Lutshnik 1916
Subgenus Pogonoidius Lutshnik 1916
214. cumanus Lutshnik 1916 + +
215. meridionalis Dejean 1828 + +
216. punctulatus Dejean 1828 + + + + + + + +
Subgenus Pogonus Nicolai 1822
217. iridipennis Nicolai 1822 + + + + + + + + + +
218. luridipennis Germar 1822 + + + + + + + + +
219. submarginatum Reitter 1908 + + + + + + + +
220. orientalis Dejean 1828 + + + + +
221. transfuga Chaudoir 1870 + + + + + + + + + + + +
222. virens Menetries 1849 + +
33.Genus POGONISTES Gebler 1830
Subgenus Pogonistes Gebler 1830
223. angustus Gebler 1830 + +
224. convexicollis Chaudoir 1871 + +
225. rufoaeneus Dejean 1828 + + + + + +
Subgenus Syrdenus Wol-laston 1862
226. grayi Wollaston 1862 + +
34.Genus STOMIS Clairville 1806
Subgenus Stomis Clairville 1806
227. pumicatus Panzer 1796 +
228. tschitscherini Semenov 1904 +
35.Genus ABACETUS Dejean 1828
Subgenus Astigis Rambur 1838
229. inexepectatus Kr. et Abdur. 1983 +
36.Genus POECILUS Chaudoir 1868
Subgenus Poecilus Chaudoir 1868
230. cupreus Linnaeus 1758 + + + + + + + + + + + + +
231. anodon Chaud. 1868 +
232. lepidus Leske 1758 + + +
233. punctulatus Schaller 1783 + +
234. sericeus Fischer von Waldheim 1823 + + + +
235. versicolor Sturm 1824 + + + + + + +
236. subcoeruleus Quens. 1806 + + +
Subgenus Angoleus Chaudoir 1876
237. crenuliger Chaudoir 1876 + + + + + +
238. lissoderus Chaudoir 1876 + + + +
239. nitens Chaudoir 1850 + + +
240. puncticollis Dejean 1828 + + + + + + +
Subgenus Derus Quensel 1806
241. advena Quensel 1806 + + + + + + + + +
37.Genus PTEROSTICHUS Schaller 1783
Subgenus Platysma Schaller 1783
242. niger niger Schaller 1783 + + + + + + + + +
243. planicollis Tschischerine 1898 +
Subgenus Argutor Mot-schulsky 1865
244. chamaeleon Motschulsky 1865 + +
245. cursor Dejean 1828 + + + + + + + +
246. vernalis Panzer 1796 + + + + + + + + + +
247. leonisi Apfelbeck 1904 +
Subgenus Pedius Sturm 1824
248. inquinatus Sturm 1824 + +
249. longicollis Duftschmid 1812 + + + + + +
Subgenus Feronidius Jeannel 1941
250. fornicatus Kolenati 1845 + +
251. melas Greutzer 1799 +
Subgenus Adelosia Mar-sham 1802
252. macer Marsham 1802 + + + + + + + + + +
253. sp. +
254. sp. +
255. sp. +
Subgenus Phonias Sturm 1824
256. strenuus Panzer 1797 + + + +
257. taksonyis Csiki 1930 +
Subgenus Haptoderus Chaudoir 1868
258. iripennis Chaudoir 1868 +
Subgenus Melanius Illiger 1798
259. anthracinus Illiger 1798 + + +
260. minor Gyll. 1827 + + + + +
261. gracilis Dejean 1828 + + + + +
262. piceolus Chaudoir 1850 +
263. nigrita Paykull 1790 + + +
Subgenus Lyrothorax Chaudoir 1838
264. caspius Menetries 1832 +
Subgenus Omaseus Duftschmid 1812
265. aterrimus Herbst 1784 +
266. elongatus Duftschmid 1812 + + + + +
Subgenus Morphnosoma Illiger 1798
267. melanarius Illiger 1798 + + +
Subgenus Bothriopterus Chaudoir 1838
268. oblongopunctatus Fabricius 1787 +
269. substrenuus +
270. phonias +
271. pseudostrenuus +
38.Genus CALATHUS Chaudoir 1846
Subgenus Calathus Chaudoir 1846
272. distinguendus Chaudoir 1846 + +
273. syriacus Chaudoir 1863 +
274. longicollis Motschulsky 1864 + + + + +
275. fuscipes Goeze 1777 + +
Subgenus Neocalathus Paykull 1790
276. ambiguus Paykull 1790 + + + + + + + + + + + + + +
277. cinctus Motschuksky 1850 + + + +
278. melanocephalus Linnaeus 1758 + + + + + + + + +
279. ochropterus Duftschmid 1812 + + +
280. kollari Put. 1873 + +
281. erratus C.Sahlberg 1827 +
282. peltatus Kolenati 1845 + + +
Subgenus Dolichus Schaller 1783
283. halensis Schaller 1783 + + + + +
39.Genus PSEUDOTAPHOXENUS Tschitscherine 1895
284. rufitarsis major Tschitsche-rine 1895 + +
40.Genus TAPHOXENUS Fischer von Waldheim 1823
Subgenus Taphoxenus Fischer von Waldheim 1823
285. gigas Fischer von Waldheim 1823 + + + +
Subgenus Lychnifugus Motschulsky 1864
286. cellarum Adams 1817 + + + +
41.Genus LAEMOSTENUS Bonelli 1810
Subgenus Laemostenus Bonelli 1810
287. caspius Menetries 1832 + + + +
Subgenus Pristonychus Dejean 1828
288. gratus Faldermann 1836 +
289. lederi Reitter 1885 +
290. sericeus piceus Motschulsky 1850 + + + +
42.Genus AGONUM Duftschmid 1812
Subgenus Agonum Duftschmid 1812
291. atratum Duftschmid 1812 + + +
292. hypocrita Apt. 1904 + +
293. extensum Menetries 1849 + + + + + +
294. duftschmidi Schmidt 1994 +
295. gracilipes Duftschmid 1812 + +
296. lugens Duftschmid 1812 + + + + + + + + + + + + +
297. longicorne Chaudoir 1846 +
298. viduum Panzer 1797 + + + + +
299. versutum Sturm 1824 +
300. viridicupreum cuprinum Motschulsky 1844. + + + + + + + + + +
301. rugicolle Chaudoir 1846 +
302. impressum Panzer 1797 +
303. dolens S. 1827 +
Subgenus Europhilus Panzer1809
304. micans Nicolai 1822 + +
305. thoreyi thorei Dejean 1828 + + + + + + + + + +
43.Genus PLATYNUS Paykull 1790
Subgenus Platynus Paykull 1790
306. longiventre Mannerheim 1825 +
307. assimile Paykull 1790 + + + +
44.Genus OXYPSELAPHUS Herbst 1784
308. obscurum Herbst 1784 + +
45.Genus ANCHOMENUS Bonelli 1810
309. dorsalis Pontoppidan 1793 + + +
46.Genus ANCHAGONUM Kryz-hanovskij 1995
310. turkestanicum Ballion 1870 +
47.Genus SYNUCHUS Gyllenha 1810
Subgenus Synuchus Gyllenha 1810
311. vivalis Illiger 1798 +
48.Genus PLATYDERUS Stephens 1828
312. umbratus Menetries 1832 +
49.Genus ATHRANUS Leconte 1848
313. collaris Menetries 1832 +
50.Genus AMARA Putzeys 1858
Subgenus Zezea Putzeys 1858
314. chaudoiri chaudoiri Putzeys 1858 + + +
315. reflexicollis Motsch. 1844 +
316. tricuspidata Dejean 1831 +
Subgenus Amara De Geer 1774
317. aenea De Geer 1774 + + + + + + + + +
318. communis Panzer 1797 + + +
319. anthobia A. Villa et G.B. Villa 1883 + + +
320. curta Dejean 1828 + +
321. eurynota Panzer 1797 + + + + +
322. lunicollis Sch. 1837 + +
323. famelica Zimmermann 1832 +
324. familiaris Duftschmid 1812 + + +
325. lucida Duftschmid 1812 +
326. montivaga Sturm 1825 +
327. similata Gyllenhal 1810 + + + + + +
328. ovata Fabricius 1792 +
329. spreta Dejean 1831 + +
330. tibialis Paykull 1798 + + + +
Subgenus Celia Chaudoir 1848
331. ambulans Zimmermann 1832 + + +
332. bifrons Gyllenhal 1810 + + + + + +
333. sollicita Pantel 1888 +
334. praetermissa C. R. Sahlberg 1827 +
335. fusca Dejean 1828 + + + + +
336. ingenua Duftschmid 1812 + + + +
337. municipalis municipalis Duftschmid 1812 + + + +
338. saginata Menetries 1847 + + +
339. viridescens Rtt. 1883 +
Subgenus Paracelia Zim-
mermann 1831
340. saxicola Zimmermann 1831 + + + + + + + + +
Subgenus Bradytus Pay-
kull 1790
341. apricaria Paykull 1790 + + + + + + + + + + + +
342. fulva O.Muller 1776 + + +
343. consularis Duftschmid 1812 + + + + + +
344. crenata Dejean 1828 + + +
345. majuscula Chaudoir 1850 + + + +
Subgenus Amathitis
Motschulsky 1844
346. abdominalis Motschulsky 1844 +
347. parvicollis Gebler 1833 + + + + +
348. subplanata Putzeys 1866 +
Subgenus Percosia Zim-
mermann 1832
349. equestris Duftschmid 1812 + +
51.Genus HARPALODEMA Reitter 1888
350. lutescens Reitter 1888 + +
351. turkmenica Tschitscherine 1894 +
52.Genus CURTONOTUS Panzer 1797
Subgenus Curtonotus
Stephens 1828
352. aulicus Panzer 1797 + + + + +
353. castaneus Putzeys 1866 + +
354. convexiusculus Marsham 1802 + + + + + + + + +
355. cribricollis Chaundoir 1846 +
356. desertus Krynicki 1866 + +
357. propinquus Menetries 1832 + + + + +
Subgenus Ammoleirus
Gebler 1830
358. megacephalus Gebler 1830 + +
359. sp. +
53.Genus ZABRUS Reiche & Saulcy 1855
Subgenus Zabrus Reiche
& Saulcy 1855
360. tenebrioides longulus Reiche & Saulcy 1855 + + + + + + + + +
361. morio Menetries 1832 + + +
Subgenus Pelor Fischer
von Waldheim 1817
362. spinipes steveni Fischer von Waldheim 1817 + + + +
363. trinii Fischer von Waldheim 1817 +
54.Genus ANISODACTYLUS Fabricius 1787
Subgenus Anisodactylus
Fabricius 1787
364. binotatus Fabricius 1787 + + + + +
365. signatus Panzer 1797 + + + + + +
Subgenus Pseudodichirus
Lutshnik 1921
366. intermedius Dejean 1829 +
Subgenus Hexatrichus
Tschitscherine 1898
367. poeciloides pseudaeneus Dejean 1829 + + + + + + + + + + +
55.Genus GYNANDROMORPHUS Dejean 1829
368. etruscus Quensel 1806 +
56.Genus DIACHROMUS Erichson 1837
369. germanus Linnaeus 1758 + +
57.Genus BRADYCELLUS Erich-son 1837
Subgenus Bradycellus Erichson 1837
370. heinzi Jaeger 1990 +
58.Genus DICHEIROTRICHUS Motschulsky 1850
371. lacustris L.Redtenbacher 1858 + + + + + +
372. ustulatus Dejean 1829 + + + + +
373. gustavii Cr. 1871 + + + + +
374. dicheirotrichus +
375. trichocellus Ganglbauer 1892 +
376. discicollis Dejean 1829 + +
377. discolor Fald. 1836 + + +
378. placidus G. 1827 + + + + +
379. stenothorax Kabak & Ka-taev 1994 +
380. tscheresovae Komarov 1995 +
381. sp. + +
59.Genus STENOLOPHUS Fischer von Waldheim 1823
Subgenus Stenolophus Fischer von Waldheim 1823
382. discophorus Fischer von Waldheim 1823 + + + + + +
383. mixtus Herbst 1784 + + + + + + + + + + + + + +
384. persicus Mannerheim 1844 + + + + + + + + + + +
385. proximus Dejean 1829 + + + + + + + + + + + +
386. skrimshiranus Stephens 1828 + + + + + + + + +
387. teutonus Schrank 1781 + +
388. steveni Krynicki 1832 +
Subgenus Egadroma Mot-schulsky 1855
389. marginatus Dejean 1829 + + + + +
60.Genus HEMIAULAX Reitter 1894
390. nigripes Reitter 1894 +
391. morio Men. 1832 + + + +
61.Genus LOXONCUS Schmidt-Gobel 1846
392. procerus Schaum. 1858 + + + + + +
62.Genus ACUPALPUS Reitter 1884
Subgenus Acupalpus Dejean 1829
393. dubius Schilsky 1888 + + +
394. elegans Dejean 1829 + + + + + + + + + + + + +
395. exiguus Dejean 1829 + + + + +
396. flaviceps Motschulsky 1850 + + +
397. luteatus Duftschmid 1812 + + + + + +
398. maculatus Schaum 1860 + + + + + + +
399. meridianus Linnaeus 1767 + + + + +
400. paludicola Reitter 1900 +
401. parvulus Sturm 1825 + + + + + + + + + +
402. suturalis Dejean 1829 + + + +
Subgenus Ancylostria Schauberger 1930
403. interstitialis Reitter 1884 +
63.Genus ANTHRACUS Duftschmid 1812
404. consputus Duftschmid 1812 + + + + + +
405. longicornis Schaum 1857 + + +
64.Genus DAPTUS Fischer von Waldheim 1824
406. vittatus Fischer von Waldheim 1824 + + + + + + + + + + +
65.Genus PAROPHONUS Ganglbauer 1892
Subgenus Ophonomimus Schauberger 1923
407. hirsutulus Dejean 1829 + + + + + + + + + + +
Subgenus Parophonus Ganglbauer 1892
408. maculicornis Duftschmid 1812 + + +
Subgenus Tachyophonus Tschitscherine 1900
409. mendax Rossi 1790 + + +
410. laeviceps Menetries 1832 + +
411. planicollis Dejean 1829 + + +
66.Genus HARPALUS Fairmaire et Laboulbene 1854
412. cephalotes Fairmaire et Laboulbene 1854 + + + + +
413. griseus Panzer 1797 + + + + + + + + + +
414. rufipes De Geer 1774 + + + + + + + + + + +
415. calceatus Duftschmid 1812 + + + + + + + + + + +
416. melancholicus Dejean 1829 + + + + + + + +
417. tenebrosus Dejean 1829 + + + + + + + + +
418. rubripes Duftschmid 1812 + + + + + + +
419. politus politus Dejean 1829 + +
420. serripes serripes Quensel 1806 + + + + + + + + + + + +
421. picipennis Duftschmid 1812 + + + +
422. atratus Laterelle 1804 +
423. pumilus Sturm 1818 + + +
424. amplicollis Menetries 1848 + + + + + + + + + +
425. anxius Duftschmid 1812 + + + + + +
426. calathoides Motschulsky 1844 + +
427. kirgisicus Motschulsky 1844 + +
428. servus Duftschmid 1812 + + + +
429. subcylindricus Dejean 1829 + +
430. froelichi Sturm 1818 + + + + + + + + + +
431. hirtipes Panzer 1797 + + + + +
432. zabroides Dejean 1829 + + + + + + +
433. flavescens Piller et Mitter-pacher 1783 +
434. dimidiatus Rossi 1790 +
435. tardus Panzer 1797 + + + + + + + +
436. latus Linnaeus 1758 + + + + +
437. rufiscapus Gebler 1833 +
438. fuscicornis Menetries 1832 +
439. fuscipalpis Sturm 1818 + + +
440. inexpectatus Kataev 1989 + +
441. compressus Motschulsky 1844 +
442. sarmaticus sarmaticus Motschulsky 1850 + +
443. smaragdinus Duftschmid 1812 + + + + + +
444. pulvinatus lubricus Reitter 1900 + + +
445. salinus salinus Dejean 1829 +
446. autumnalis Duftschmid 1812 +
447. foveiger Tschitscherine 1895 +
448. caspius Steven 1806 + + + +
449. dispar splendens Gebler 1830 + +
450. pygmaeus Dejean 1829 + + +
451. circumpunctatus Chaudoir 1846 + + +
452. hospes hospes Sturm 1818 + + + + + +
453. steveni Dejean 1829 + + +
454. terrestris Motschulsky 1844 + +
455. affinis Schrank 1781 + + + + +
456. angulatus scytha Tschitscherine 1899 + +
457. distinguendus distinguen-dus Duftschmid 1812 + + + + + + + +
458. akinini Tschitscherine 1895 +
459. oblitus Dejean 1829 + +
460. signaticornis Duft. 1812 +
461. cisteloides Schouberti Tsch. 1828 + + +
462. metallinus Menetries 1838 + + +
463. honestus Duftschmid 1812 + +
464. flavicornis Dejean 1829 + + +
465. dispar splendens Gebler 1830 + +
466. modestus Dejean 1829 +
467. saxicola Dejean 1829 +
468. litigiosus Dejean 1829 +
469. cupreus Dejean 1829 +
470. sp. +
67.Genus MICRODERES Dejean 1829
Subgenus Microderes Dejean 1829
471. brachypus Dejean 1829 + + + + + + + +
68.Genus PANGUS Dejean 1821
472. scaritides Sturm 1818 +
473. sp. +
69.Genus ACINOPUS Zoubkoff 1833
Subgenus Haplacinopus Zoubkoff 1833
474. striolatus Zoubkoff 1833 + + + +
Subgenus Oedematicus Bedel 1897
475. megacephalus Rossi 1794 + + + + +
Subgenus Acinopus Dejean 1821
476. laevigatus Menetries 1832 + + + + + + + +
477. picipes Olivier 1795 + + + + + +
Subgenus Osimus Dejean 1829
478. ammophilus Dejean 1829 + + + + +
70.Genus OPHONUS Duftschmid 1812
Subgenus Metophonus Duftschmid 1812
479. rupicola Sturm 1818 + + + + + +
480. nitidulus Stephens 1828 + +
481. cordatus Duftschmid 1812 + + +
482. melletii Heer 1837 + + + + +
483. rufibarbis Fabricius 1792 + + + + + + +
484. puncticeps Stephens 1828 + + + + +
485. puncticollis Paykull 1798 + + + + +
486. sciakyi Wrase 1990 + +
487. schaubergerianus Puel 1937 +
Subgenus Hesperophonus Fabricius 1775
488. azureus Fabricius 1775 + + + + + + + + +
489. subquadratus Dejean 1829 + + + + +
490. cribricollis Dejean 1829 + + + +
491. convexicollis Menetries 1832 + + + +
492. similis Dejean 1829 + + + +
493. minimus Motschulsky 1845 + +
494. sp. +
495. sp. +
496. sp. +
Subgenus Ophonus Panzer 1796
497. stictus Stephens 1828 + + + + +
498. ardosiacus Lutshnik 1922 + +
499. diffinis Dejean 1829 + +
500. sabulicola Panzer 1796 + + + + + +
71.Genus EUCARTERUS Reitter 1900
501. sparsutus Reitter 1898 + + +
72.Genus CARTERUS Dejean 1829
Subgenus Pristocarterus Wrase 1994
502. angustipennis Chaudoir 1852 + + +
503. angustus Menetries 1832 +
Subgenus Carterus Dejean 1829
504. rufipes Chaudoir 1843 + + +
73.Genus DITOMUS Fabricius 1792
505. tricuspidatus Fabricius 1792 + +
506. calydonius Rossi 1790 +
74.Genus DIXUS Dejean 1825
507. eremita Dejean 1825 + + +
508. obscurus Dejean 1825 + + + +
75.Genus AMBLYSTOMUS Erichson 1837
509. niger Helr. 1838 + + + +
510. metallescens Dejean 1829 + + +
76.Genus PERIGONA Castelnau 1835
Subgenus Trechicus Le-conte 1853
511. nigriceps Dejean 1831 +
77.Genus PANAGAEUS Fabricius 1775
512. bipustulatus Fabricius 1775 + +
513. cruxmajor Linnaeus 1758 + + + + + +
78.Genus CALLISTUS Fabricius 1775
514. lunatus Fabricius 1775 + + +
79.Genus EPOMIS Dejean et Boisd. 1830
515. dejeani Dejean et Boisduval 1830 + +
516. circumscriptus Duft. 1812 + + +
80.Genus DINODES Bonelli 1810
517. cruralis Fischer 1892 + + + +
518. dicipiens Dufour 1820 +
81.Genus CHLAENIUS Dejean 1826
Subgenus Chlaenites Mot-
schulsky 1850
519. inderiensis Motschulsky 1850 + + + +
520. spoliatus spoliatus Rossi 1790 + + + + + + + + +
Subgenus Trichochlaenius
Seidlitz 1887
521. aeneocephalus Dejean 1826 + + + + +
522. steveni Quensel 1806 + +
Subgenus Stenochlaenius
Rtt. 1908
523. coeruleus Steven 1809 + +
Subgenus Chlaenius Bo-
nelli 1810
524. festivus Panzer 1796 + + +
Subgenus Chlaeniellus
Krynicki 1832
525. nigricornis Fabricius 1787 + +
526. terminatus Dejean 1826 + +
527. tristis tristis Schaller 1783 + + + + + + + + + + + + +
528. chrysothorax Krunicki 1832 + +
529. vestitus Paykull 1790 + + + + + + + + + +
530. flavipes Menetries 1832 +
531. velutinus Duft. 1812 +
Subgenus Agostenus Gebler 1829
532. alutaceus Gebler 1829 + + + + + + + +
533. sulcicollis Paykull 1798 +
82.Genus OODES A.Villa et G.B.Villa 1833
Subgenus Oodes A.Villa
et G. B.Villa 1833
534. gracilis A.Villa et G.B.Villa 1833 + + + + + + + +
535. desertus Motsch. 1858 + + + + +
536. helopioides Fabricius 1792 + + + +
Subgenus Lachnocrepis
H.Bates 1873
537. prolixus H.Bates 1873 + + + +
83.Genus DIPLOCHEILA Brulle 1834
Subgenus Diplocheila Brulle 1834
538. transcaspica Semenov 1890 +
84.Genus LICINUS Latreille 1802
Subgenus Licinus La-
treille 1802
539. cassideus Fabricius 1792 +
85.Genus BADISTER Schrank 1798
Subgenus Badister
Schrank 1798
540. bullatus Schrank 1798 + + + + + + +
541. meridionalis Puel 1925 + + + + + + +
542. unipustulatus Bonelli 1813 + + + + +
543. sp. +
Subgenus Trimorphus Stephens 1828
544. sodalis Duftschmid 1812 +
Subgenus Baudia Motschulsky 1844
545. collaris Motschulsky 1844 + +
546. dilatatus Chaudoir 1837 + +
547. peltatus Panzer 1796 + + + +
86.Genus MASOREUS Gyllenhal 1813
548. wetterhalli Gyllenhal 1813 + + + +
87.Genus CORSYRA Steven in Dejean 1825
549. fusula Steven in Dejean 1825 + +
88.Genus PENTAGONICA Schm-Goebel 1846
550. sp. +
89.Genus ODACANTHA Linnaeus 1767
551. melanura Linnaeus 1767 + + + + + + + + +
90.Genus LEBIA Hoffmannsegg 1803
Subgenus Lebia Linnaeus 1758
552. crux-minor Linnaeus 1758 + + +
553. menetriesi Ballion 1869 + + +
554. turkestanica Jedlicka 1966 +
555. trimaculata Villers 1789 + + + +
556. holomera Chaudoir 1870 +
557. humeralis Dejean 1825 +
558. scapularis Fourcroy 1785 + +
Subgenus Lamprias Bonelli 1809
559. cyanocephala Linnaeus 1758 + + + + +
91.Genus CYMINDOIDEA Castel-nau 1832
Subgenus Platytarus Fairmaire 1850
560. famini Dejean 1826 + + + +
92.Genus DEMETRIAS Samouelle 1819
Subgenus Demetrias Samouelle 1819
561. monostigma Samouelle 1819 + + + + +
Subgenus Aetophorus Germar 1824
562. imperialis Germar 1824 + + + +
93.Genus DROMIUS Bonelli 1810
563. laeviceps Motsch. 1850 +
94.Genus PARADROMIUS Dejean 1826
Subgenus Paradromius Dejean 1826
564. suturalis Motschulsky 1844 +
565. linearis Ol. 1795 + + +
95.Genus PHILORHIZUS Hope 1838
Subgenus Philorhizus Hope 1838
566. notatus Stephens 1827 +
567. sigma Rossi 1790 +
96.Genus SYNTOMUS Duftschmid 1812
568. obscuroguttatus Duftschmid 1812 + + + + + + +
569. parallelus Ballin 1870 +
570. pallipes Dejean 1825 + + + +
571. truncatellus Linnaeus 1761 +
572. fuscomaculatus Motschulsky 1844 + +
97.Genus CHAROPTERUS Motschulsky 1839
573. paracenthesis Motschulsky 1839 + +
98.Genus MICROLESTES Komarov 1990
574. badulini Komarov 1990 +
575. corticalis Dufour 1820 + + +
576. romanschov +
577. fissuralis Reitter 1901 + +
578. fulvibasis Reitter 1900 + + + + + +
579. maurus Sturm 1827 + + +
580. minutulus Goeze 1777 + + + +
581. negrita Wollaston 1854 + + + + +
582. plagiatus Duftschmid 1812 + + + + + +
99.Genus CYMINDIS Gyllenhal 1810
Subgenus Cymindis Gyllenhal 1810
583. decora Fischer von Waldheim 1829 + +
584. lineata Quensel 1806 + + + +
585. picta picta Pallas 1771 + + + + +
586. scapularis Schaum. 1857 + +
587. ornata Fischer von Waldheim 1824 + +
588. andreae Men. 1832 +
589. axillaris Fabricius 1794 + +
590. accentifera Zubkoff 1833 +
591. intermedia Chaudoir 1873 +
Subgenus Iscariotes Emetz et Kryzhanovskij 1973
592. kiritschenkoi Emetz et Kryzhanovskij 1973
593. sabulosa Motschulsky 1850 + +
Subgenus Menas Motschulsky 1844
594. variolosa Fabricius 1794 + + + + + +
595. violacea Chaudoir 1873 + + +
596. cylindrica Motsch. 1844 +
Subgenus Tarsostinus Gebler 1825
597. lateralis Fischer von Waldheim 1821 + +
598. equestris Gebler 1825 +
Subgenus Neomenas Emetz 1974
599. antonowi Semenov 1891 + +
100.Genus AGATUS Gebler 1843
60 cingulatus Gebler 1843 +
101.Genus DRYPTA Latreiile 1796
601. dentata Rossi 1790 + + +
102. Genus POLYSTICHUS Fourcroy 1785
602. connexus Fourcroy 1785 + + + + + + +
103.Genus ZUPHIUM Rossi 1790
Subgenus Zuphium Rossi 1790
603. olens Rossi 1790 + + + + + + + + + +
Subgenus Parazuphium Jeneal 1942
604. chevrolati Castelnau 1833 + + + +
104. Genus BRACHINUS Quensel 1806
Subgenus Brachinus F. Weber 1801
605. bipustulatus Quensel 1806 + + + + + + + + +
606. bodemeyeri Apfelbeck 1904 + +
607. costatulus Quensel 1806 + + + + + + + + + +
608. crepitans Linnaeus 1758 + + + + + + +
609. cruciatus Quensel 1806 + + + + + + + + +
610. elegans Chaudoir 1842 + + +
611. hamatus Fischer von Waldheim 1828 + + + + + + + + + +
612. exhalans Rossi 1792 + + + + + +
613. explodens Duftschmid 1812 + + + + + +
614. klapperichi Jedlicka 1955 +
615. plagiatus Reiche 1868 + + + + +
616. psophia Serville 1821 + + + + + + + +
617. quadriguttatus Gebler 1829 + +
618. alexandri Battoni 1984 + + + +
619. ejaculans Fischer von Waldheim 1829 + + + + +
620. brevicollis Motschulsky 1844 + + + + + +
621. bayardi Dejean 1831 +
622. nigricornis Gebler 1829 +
623. berytensis Reiche 1855 + + + +
624. sp. +
105.Genus MASTAX Steven 1806
625. thermarum thermarum Steven 1806 + + + + +
ИТОГО: 8 2 5 00 2 2 1 0 TT 1 6 m 1 3 m 1 9 m 1 9 m 1 0 1 6 00 1 8 TT 1 7 1 W4 <s 00 00
Работа выполнена при поддержке ФЦП «Иследования и разработки по приоритетным направлениям развития научно-технического комплекса России на 2007-2013 годы», ГК №16.552.11.7051
Работа выполнена при поддержке ФЦП «Научные и научно-педагогические кадры инновационной России» на 2009-2013 гг»., ГК №16.740.11.0051
