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Амурский зоологический журнал, 2022, т. XIV, № 1
Amurian Zoological Journal, 2022, vol. XIV, no. 1
www.azjournal.ru
https://www.doi.org/10.33910/2686-9519-2022-14-l-48-60 http://zoobank.org/References/BCB59573-E835-4876-8FlD-B685A0ADDlC2
UDC 595.722
New records of Dolichopodidae (Diptera) from Primorsky Territory, Russia
I. Ya. Grichanov
All-Russian Institute of Plant Protection 3 Podbelskogo Rd., Pushkin, Saint Petersburg 196608, Russia
Abstract. A collection of long-legged flies was studied in the Laboratory of Entomology of the Federal Research Centre for Biodiversity of Terrestrial Biota of East Asia, Vladivostok. A new material was identified for the first time, collected mainly by the researchers of this laboratory in the Primorsky Territory since 1970s, as well as by Igor Ya. Grichanov during his visit to Vladivostok in 2021. Unidentified material from some other scientific institutions is also included in the list. In total, new records are given for 73 species of long-legged flies. Hercostomus rohdendorfi Stackelberg, 1933 is transferred to the genus Gymnopternus (comb.n.). Mesorhaga janata Negrobov, 1984 is placed in synonymy with Amblypsilopus pilosus (Negrobov, 1979) (syn.n.). Dolichophorus immaculatus, Dolichopus henanus, D. kumazawai, Hercostomus beijingensis, Rhaphium dorsiseta, and R. mediocre are found for the first time in Russia. Chrysotus caerulescens, Hydrophorus albiceps, Rhaphium fascipes and Syntormon violovitshi are reported from Primorye for the first time. The finding of Dolichopus griseifacies is the first reliable record from Primorye. In total, 215 species from 31 genera of Dolichopodidae are known in the regional fauna.
Keywords: Dolichopodidae, Russia, Primorsky Territory, new synonym, new combination, new records.
Новые находки Dolichopodidae (Diptera) в Приморском крае
России
И. Я. Гричанов
Всероссийский институт защиты растений, шоссе Подбельского, д. 3, г. Пушкин, 196608,
г. Санкт-Петербург, Россия
Аннотация. Изучена коллекция мух-зеленушек Лаборатории энтомологии Федерального научного центра биоразнообразия наземной биоты Восточной Азии ДВО РАН. Впервые определен материал, собранный главным образом сотрудниками этой лаборатории в Приморском крае с 1970-х гг., а также И. Я. Гричановым во время его пребывания во Владивостоке в 2021 г. Ранее неопределенный материал из некоторых других научных учреждений также включен в список. Всего приведены новые указания для 73 видов мух-зеленушек. Hercostomus rohdendorfi Stackelberg, 1933 перенесен в род Gymnopternus (comb. n.). Mesorhaga janata Negrobov, 1984 помещен в синонимы к Amblypsilopus pilosus (Negrobov, 1979) (syn. n.). Dolichophorus immaculatus, Dolichopus henanus, D. kumazawai, Hercostomus beijingensis, Rhaphium dorsiseta и R. mediocre впервые обнаружены в фауне России. Chrysotus caerulescens, Hydrophorus albiceps, Rhaphium fascipes и Syntormon violovitshi впервые обнаружены в Приморье. Находка Dolichopus griseifacies является первой достоверной находкой из Приморья. Всего в крае отмечено 215 видов из 31 рода приморской фауны Dolichopodidae.
Ключевые слова: Dolichopodidae, Россия, Приморский край, новый синоним, новая комбинация, новые указания.
Author
Igor Ya. Grichanov E-mail: grichanov@mail.ru SPIN: 1438-5370 Scopus Author ID: 8672518800 ResearcherID: A-1406-2013 ORCID: 0000-0002-7887-7668
Copyright: © The Author (2022). Published by Herzen State Pedagogical University of Russia. Open access under CC BY-NC License 4.0.
Сведения об авторе Гричанов Игорь Яковлевич E-mail: grichanov@mail.ru SPIN-код: 1438-5370 Scopus Author ID: 8672518800 ResearcherID: A-1406-2013 ORCID: 0000-0002-7887-7668
Права: © Автор (2022). Опубликовано Российским государственным педагогическим университетом им. А. И. Герцена. Открытый доступ на условиях лицензии СС БУ-ЫС 4.0.
Introduction
Recently, the data of the known dolicho-podid fauna of the Primorsky Territory, or Primorye, were summarised in form of the annotated checklist (Grichanov, Selivanova 2021). In total, 204 species from 30 genera of long-legged flies were listed. Dolichopus pen-natus Meigen, 1824 included in the Palaearc-tic Catalog (Negrobov, 1991) as known from Primorye was overlooked by that list (original material was not provided). Chrysotus orientalis Negrobov et Tsurikov, 2000 (Negrobov et al. 2000) was also overlooked by that list.
The cluster analysis of the dolichopodid genera and species composition was conducted for continental Chinese and Russian regions belonging to the East Palaearctic (Grichanov et al. 2021). This analysis along with the comparison of the the biodiversity index supports the uniqueness of Primorsky Territory as the biodiversity centre in East Asian Palaearctic. A decrease in the number of known species and biodiversity indices is observed northward, southward and westward of this Territory.
Recently, a rich collection of long-legged flies was studied in the Laboratory of Entomology of the Federal Research Centre for Biodiversity of Terrestrial Biota of East Asia, Vladivostok (FCBV). A new material collected mainly by the researchers of this laboratory in the Primorsky Territory since 1970s was identified for the first time. Also the material collected by Igor Ya. Grichanov during his visit to Vladivostok in 2021 and that from collection of the Zoological Museum of Moscow State University (ZMMU) is also included in the list. The material collected by Dr. Oksana Kosheleva (All-Russian Institute of Plant Protection) is fixed in 70% ethanol; it will be deposited at the Zoological Institute of the Russian Academy of Sciences (ZIN).
The information on the global distribution for each species collected follows Yang et al. (2018), Grichanov (2021) and Grichanov, Selivanova (2021). The type localities are provided and the country lists are arranged al-
phabetically. The words "Region" (oblast) and "Territory" (kray) are omitted from the list of Russian regions. It is worth noting that some species names were first published in keys, having priority against species descriptions and their distribution published later and referenced in this paper. It corresponds with the International Code of Zoological Nomenclature. Remarks are provided where deemed necessary.
New records
Genus Amblypsilopus Bigot, 1888
Amblypsilopuspilosus (Negrobov, 1979) Mesorhaga pilosa Negrobov 1979: 188. Mesorhaga janata Negrobov 1984: 32, syn. nov. Type locality: Japan, Ofune. Amblypsilopus janatus: Bickel 1994: 373. Material examined. 3^, 3$, Primorye, Khasansky Distr., Ryazanovka, 12, 14.08.1987, V. N. Makarkin (FCBV). Distribution. Type locality: Russia, Primo-rye, Komarovka. Palaearctic: Japan, Russia (Primorye).
Notes. Negrobov (1984) distiguished Amblypsilopus janatus (males described from Japan) from A. pilosus (males and females described from Primorye) by right vs. acute angle between M1+2 and M1. Descriptions of the two species including male genitalia figured have no significant differences. The material examined has right angle between Mx+2 and M1 in males and acute angle in females. It is worth noting that the line drawing of A. pilosus is inaccurate (Negrobov 1979). The wing of all Primorian male specimens examined is identical to wing pictured for A. janatus. Therefore, the two names are considered here as synonyms. The species seems not uncommon in both Primorye and Japan (Grichanov 2020; Grichanov, Selivanova 2021; Negrobov 1979; 1984).
Genus Argyra Macquart, 1834
Argyra arrogans Takagi, 1960 Material examined. 3^, Primorye, Khasansky Distr., Ryazanovka, 24.06., 19, 24.07.1987, V. N. Makarkin (FCBV).
Distribution. Type locality: Japan, Hokkaido, Aizan-Kei. Palaearctic: Japan, Russia (Primo-rye); Oriental: China (Zhejiang, Guizhou). Notes. This species was first recorded from Primorye by Selivanova et al. (2009). Argyra shamshevi Selivanova et Negrobov, 2006 Material examined. 1^, Primorye, Ussuri-isky Nature Reserve, Malaise trap, 30.06.10.07.1991, P. Vilkamaa (FCBV ex coll. Museum Zoology, Helsinki, Finland). Distribution. Type locality: Russia, Primorye, Vladivostok env.: Okeanskaya at Kedrovaya Pad Nature Reserve. Palaearctic: Russia (Pri-morye).
Argyra ussuriana Negrobov, 1973 Material examined. 1^, 1$, Primorye, Khasansky Distr., Ryazanovka, 16, 25.06.1987, V. N. Makarkin (FCBV). Distribution. Type locality: Russia, Primo-rye, Partizansky Distr., Tigrovaya. Palaearctic: Russia (Primorye).
Genus Campsicnemus Haliday, 1851
Campsicnemus versicolorus Negrobov et Zlobin, 1978
Material examined. 1^, Primorye, Khasansky Distr., Ryazanovka, 9.08.1987, V. N. Ma-karkin (FCBV).
Distribution. Type locality: Russia, Primorye, Yakovlevka. Palaearctic: Russia (Primorye).
Genus Chrysotimus Loew, 1857
Chrysotimus flavisetus Negrobov, 1978 Material examined. 1^, Primorye, Khasansky Distr., Ryazanovka, 23.07.1987, V. N. Ma-karkin (FCBV).
Distribution. Type locality: Russia, Primo-rye, Sichote-Alinsky Nature Reserve, 37 km from Ternei, River Serebryanka. Palaearctic: Russia (Primorye).
Chrysotimus spinuliferus Negrobov, 1978 Material examined. 8^, Primorye, Khasan-sky Distr., Ryazanovka, 19, 23.07., 9.08.1987, V. N. Makarkin; 3^, Primorye, Spassky Distr., [Gorno-Tayozhnoe vil.], Gornotayozhnaya station, 20.07.1990, V. N. Makarkin (all in FCBV). Distribution. Type locality: Russia, Primo-rye, Yakovlevka. Palaearctic: Russia (Kuriles, Primorye, Sakhalin, Yakutia).
Genus Chrysotus Meigen, 1824
Notes. The East Palaearctic Chrysotus species are usually indistinguishable by females; therefore, females collected are left unidentified.
Chrysotus caerulescens Negrobov, 1980 Material examined. 1$, Primorye, Oblach-naya Mt., 1700 m, 17.07.1993, V. N. Makarkin (FCBV).
Distribution. Type locality: Russia: Sob River, Bolshoy Ural, Obdorsk [= Salekhard]. Palaearctic: Russia (Altai Rep., Amur Region, Bury-atia, Khantia-Mansia, Krasnoyarsk, Yamalia). First record from Primorye.
Chrysotus cilipes Meigen, 1824 Material examined. 2$, Primorye, Spassky Distr., Novoselskoe, 28, 29.06.1985, V. N. Makarkin; 1$, Primorye, Arsenyev, agroland-scape, 30.06.1986, V. N. Makarkin; 1$, Primorye, Khasansky Distr., Ryazanovka, 19.07.1987, V. N. Makarkin; 1$, Primorye, Chernigovsky Distr., Gornyi Khutor, sweeping on Ambrosia, 24.06.1988, L. Esipenko; 1$, Primorye, Spassky Distr., Prokhory, sweeping, 6.07.1990, L. Esipenko; 1$, Primorye, Chernigovsky Distr., Dmitrievka, sweeping on Ambrosia, 6.07.1990, L. Esipenko; 2$, Pri-morye, Krasnoarmeisky Distr., Dersu, soya, 19-26.08.1991, V. N. Makarkin (all in FCBV); 32$, Primorye, Khankaisky Distr., 6 km N No-vokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Kosheleva; 2$, Pri-morye, Partizansk env., sweeping, 25.08.2019, mountain springs, Kosheleva (all in ZIN). Distribution. Type locality: Germany, Hamburg. Trans-Palaearctic species.
Chrysotus degener Frey, 1917 Material examined. 17$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Kosheleva (ZIN). Distribution. Type locality: Sri Lanka, Anu-radhapura. Palaearctic: China (Anhui, Beijing, Heilongjiang, Jiangsu, Liaoning, Shaanxi), Russia (Amur Region, Primorye, Yakutia); Oriental: China (Chongqing, Guangxi, Henan, Taiwan, Yunnan, Zhejiang), India, Myanmar, Pakistan, Sri Lanka.
Chrysotus glebi Negrobov et Maslova, 1995 Material examined. 1$, Primorye, Khasan-sky Distr., Ryazanovka, 24.06.1987, V. N. Ma-karkin (FCBV).
Distribution. Type locality: Russia, Primorye, Ussuriisky Nature Reserve. Palaearctic: Kyrgyzstan, Russia (Altai Rep., Amur Region, Karachai-Cherkessia, Leningrad, Murmansk, Primorye, Yakutia), Spain, Turkey, Ukraine. Chrysotus logvinovskii Negrobov et Tsuri-kov, 2000
Material examined. 1$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Ko-sheleva (ZIN).
Distribution. Type locality: Russia: Sakhalin, Anivsky Distr. Palaearctic: Russia (Sakhalin). First record from Primorye.
Chrysotus orientalis Negrobov et Tsurikov, 2000
Material examined. 3$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Ko-sheleva (ZIN).
Distribution. Type locality: [Inner] Mongolia: Khent. aimak, Kerulen River, 40 km WSW Bayan-Obo. Palaearctic: China (Inner Mongolia), Mongolia, Russia (Altai Rep., Primo-rye, Yakutia, Zabaikalye). Notes. The species was overlooked in the annotated checklist (Grichanov, Selivanova 2021).
Genus Diaphorus Meigen, 1824
Diaphorus nigricans Meigen, 1824 Material examined. 1$, Primorye, Khasan-sky Distr., Ryazanovka, 23.07.1987, V. N. Ma-karkin (FCBV).
Distribution. Type locality: "Germany." Afro-tropical, Nearctic, Oriental, Palaearctic and Neotropical Regions. Diaphorusparenti Stackelberg, 1928 Material examined. 5$, 2$, Primo-rye, Kedrovaya Pad, 20, 23.07., 3.08.1976, R. G. Soboleva; 1$, Primorye, Ussuriisky Distr., Kamenushka, Barsukovka River, 3.07.1979, R. G. Soboleva; 3$, 2$, Primorye, Khasansky Distr., Ryazanovka, 23, 23.07.1987, V. N. Ma-karkin (all in FCBV).
Distribution. Type locality: Russia, Primo-rye, "Suchansky" (= Partizansky) District, Ti-grovaya. Palaearctic: China (Ningxia, Hebei, Henan), Russia (Karachai-Cherkessia, Primo-rye, Zabaikalye).
Genus Dolichophorus Lichtwardt, 1902
Dolichophorus immaculatus Parent, 1944 Material examined. 2$, Primorye, Cherni-govsky Distr., Evseevka, sweeping, 10.07.1987, L. Esipenko; 3$, Primorye, Chernigovsky Distr., Dmitrievka, sweeping on Ambrosia, 6.07.1990, L. Esipenko (all in ZIN ex coll. FCBV). Distribution. Type locality: China: "Kinpeng, Ordos." Palaearctic: China (Inner Mongolia, Ningxia).
Notes. See Yang et al. (2011) for the redescription of this species. The genus is reported from Far Eastern Russia for the first time. The species is reported from Russia for the first time.
Genus Dolichopus Latreille, 1796
Notes. Several closely related Dolichopus species were identified by males from one sample (Khankaisky district, 6 km N Novokachalinsk, 1-4.09.2019) or one locality (e. g. Chernigovsky district, sweeping on Ambrosia). The Far Eastern species of the genus are usually indistinguishable by females; therefore, most females collected there are left unidentified.
Dolichopus agilis Meigen, 1824 Material examined. 9$, Primorye, Chernigovsky Distr., Evseevka, sweeping, 7.06., 4, 11, 15.07.1988, L. Esipenko; 11$, Primorye, Chernigovsky Distr., Dmitrievka, sweeping on Ambrosia, 7.06., 6, 11.07., 20.08.1988, L. Esipenko; 2$, Primorye, Chernigovsky Distr., Gornyi Khutor, sweeping on Ambrosia, 7, 28.06.1988, L. Esipenko; 2$, Primorye, Spassky Distr., Prokhory, sweeping, 7.06.1990, L. Esipenko (all in FCBV). Distribution. Type locality: not given (Germany?). Trans-Palaearctic species. Dolichopus amurensis Stackelberg, 1930 Material examined. 1$, Primorye, Ussuri-isky Distr., Kamenushka, Barsukovka River, 13.06.1979, R. G. Soboleva (FCBV).
Distribution. Type locality: Russia: "Amurlande: Banjbo, Port-Ajan" (= Khabarovsky Territory, Ayan, 56°45'N, 138°16'E). Palae-arctic: Mongolia, Russia (Khabarovsk, Primo-rye).
Dolichopus aubertini Parent, 1934 Material examined. 1$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Kosheleva (ZIN). Distribution. Type locality: China, "Tientsin" (= Tianjin). Palaearctic: China (Beijing, Hebei, Tianjin), Russia (Primorye).
Dolichopus bigeniculatus Parent, 1926 Material examined. 1$, Primorye, Vladivostok, 12th km, 19.07.1985, V. N. Makar-kin; 1$, Primorye, Arsenyev, agrolandscape, 28.06.1986, V. N. Makarkin; 5$, Primorye, Arsenyev, 10, 12.08.1990, V. N. Makarkin; 5$, Primorye, Krasnoarmeisky Distr., Dersu, soya, 19-26.08.1991, V. N. Makarkin (all in FCBV); 9$, Primorye, Khankaisky Distr., 6 km N No-vokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Kosheleva (ZIN). Distribution. Type locality: China, Shanghai, "Zi-Ka-Wei" (= Xujiahui). Palaearctic: China (Beijing, Henan, Shaanxi, Shandong), Japan, Russia (Kuriles, Primorye, Khabarovsk); Oriental: China (Anhui, Jiangsu, Sichuan, Zheji-ang).
Dolichopus calceatus Parent, 1927 Material examined. 1$, Primorye, Leso-zavodsky Distr., Innokentyevka, 10.07.1976, A. Berezantsev; 2$, Primorye, Chernigov-sky Distr., Gornyi Khutor, sweeping on Ambrosia, 4.07.1988, L. Esipenko; 2$, Primo-rye, Chernigovsky Distr., Evseevka, sweeping, 20.06.1988, L. Esipenko (all in FCBV). Distribution. Type locality: Russia, Zabaika-lye: "Transbaikale; Pjetschanka (= Peschanka), b. Tschita." Palaearctic: China, Kazakhstan, Russia (Altai Rep., Amur Region, Buryatia, Irkutsk, Kamchatka, Khabarovsk, Primorye, Tomsk, Yakutia, Zabaikalye).
Dolichopus cuneipennis Parent, 1926 Material examined. 1$, Primorye, Chernigovsky Distr., Gornyi Khutor, sweeping on Ambrosia, 15.07.1988, L. Esipenko (FCBV).
Distribution. Type locality: China: "Tchen-Kiang, Zi-Ka-Wei" (= Xujiahui, near Shanghai). Palaearctic: China (Heilongjiang, Jilin, Shaanxi), Russia (Kuriles, Primorye); Oriental: China (Shanghai).
Dolichopus discifer Stannius, 1831 Material examined. 1$, Primorye, Parti-zansky Mt. Range, Olkhovaya Mt., 1660 m, 8.08.1986, V. N. Makarkin (FCBV). Distribution. Type locality: Germany. Hol-arctic species.
Dolichopus disharmonicus Smirnov, 1948 Material examined. 1$, Primorye, Krasnoarmeisky Distr., Dersu, soya, 19-26.08.1991, V. N. Makarkin (FCBV). Distribution. Type locality: Russia, Okeanskaya, near Vladivostok. Palaearctic: Russia (Primorye). Dolichopus eurypterus Gerstacker, 1864 Material examined. 1$, Primorye, Ussuriisky Distr., Kamenushka, light trap, 13.07.1979, R. G. Soboleva; 1$, Primorye, [Ussuriisk, Gor-no-Tayozhnoe vil.], Gornotayozhnaya station, arboretum, 27.06.1980, M. Mikhailovskaya; 2$, Primorye, Spassky Distr., Novoselskoe, 29.06., 29.07.1985, V. N. Makarkin; 1$, Pri-morye, Khorol, 31.08.1986, V. N. Makarkin; 1$, Primorye, Khasansky Distr., Ryazanovka, light trap, 12.08.1987, V. N. Makarkin; 16$, Primorye, Chernigovsky Distr., Dmitrievka, sweeping on Ambrosia, 15, 19.07., 20.08.1988, L. Esipenko; 9$, Primorye, Chernigovsky Distr., Gornyi Khutor, sweeping on Ambrosia, 26.07.1987, 11, 15.07.1988, L. Esipenko; 14$, Primorye, Chernigovsky Distr., Evseevka, sweeping on Ambrosia, 7, 11.06., 4, 6, 15.07.1988, L. Esipenko (all in FCBV). Distribution. Type locality: Germany, Berlin. Palaearctic: Belgium, Czechia, Germany, Hungary, Kazakhstan, Latvia, Poland, Russia (Khabarovsk, Primorye). Dolichopus griseifacies Becker, 1917 Material examined. 14$, Primorye, Oblach-naya Mt., 1700 m, 17, 18.07.1993, V. N. Makarkin (FCBV).
Distribution. Type locality: Russia, "Irkutsk, Sibirien." Palaearctic: Russia (Amur Region, Buryatia, Irkutsk, Khabarovsk, Primorye). First reliable record from Primorye.
Dolichopus henanus Yang, 1999 Material examined. 2$, Primorye, Arse-nyev, 27.07.1985, V. N. Makarkin (ZIN ex coll. FCBV).
Distribution. Type locality: China: Henan, Xixia. Palaearctic: China (Henan). Notes. The species is reported from Russia for the first time. Dolichopus hilaris Loew, 1862 Material examined. 1$, Primorye, Spassky Distr., Novoselskoe, 29.06.1985, V. N. Makarkin (FCBV); 12$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Ko-sheleva (ZIN); 1$, Primorye, Chernigovsky Distr., Dmitrievka, sweeping, 1.08.1983, Are-fin, Ivliev (ZIN ex coll. FCBV). Distribution. Type locality: Poland, Miçdzyrzecz. Trans-Palaearctic species. Dolichopus kumazawai Maslova, Negrobov et Fursov, 2014
Material examined. 6$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Ko-sheleva (ZIN).
Distribution. Type locality: Japan, Honshu, Aichi Pref., Nagoya, Nagakutecho Park. Palae-arctic: Japan.
Notes. The species is reported from Russia for the first time.
Dolichopus linearis Meigen, 1824 Material examined. 2$, Primorye, Arsenyev, agrolandscape, 28.06.1986, V. N. Makarkin; 3$, Chernigovsky Distr., Gornyi Khutor, sweeping on Ambrosia, 4.07.1988, L. Esipenko; 1$, Primorye, Chernigovsky Distr., Dmitrievka, sweeping on Ambrosia, 4.07.1988, L. Esipenko; 1$, Primorye, Chernigovsky Distr., Evseevka, sweeping on Ambrosia, 6.07.1988, L. Esipenko; 1$, Primorye, Spassky Distr., Prokhory, sweeping on Ambrosia, 6.07.1990, L. Esipenko; 1$, Primorye, Krasnoarmeisky Distr., Dersu, soya, 19-26.08.1991, V. N. Makarkin (all in FCBV). Distribution. Type locality: not given (Germany?). Trans-Palaearctic species. Dolichopus leucopus Smirnov, 1948 Material examined. 28$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk,
45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, Lake shore, Kosheleva (ZIN). Distribution. Type localities: Russia: "Okean-skaya, near Vladivostok; Suchan (= Parti-zansk); Kamen-Rybolov, Krivoi Klyuch, Gor-notayozhnaya station." Palaearctic: Russia (Khabarovsk, Primorye). Dolichopus longicornis Stannius, 1831 Material examined. 2$, Primorye, Kras-noarmeisky Distr., Dersu, soya, 19-26.08. 1991, V. N. Makarkin; 1$, Primorye, Krasnoarmeisky Distr., Dersu, Bol'shaya Ussurka (= Iman) River, 13.06.1990, V. N. Makarkin (all in FCBV).
Distribution. Type locality: not given (Germany: Hamburg?, Breslau?). Trans-Palaearc-tic species; Nearctic: Canada (Yukon), USA (Alaska).
Dolichopus martynovi Stackelberg, 1930 Material examined. 1$, Primorye, [Ussuri-isk, Gorno-Tayozhnoe vil.], Gornotayozhnaya station, arboretum, 3.07.1980, M. Mikhailovs-kaya; 3$, Primorye, Chernigovsky Distr., Ev-seevka, sweeping, 26.07.1987, L. Esipenko; 18$, Primorye, Chernigovsky Distr., Dmitrievka, sweeping on Ambrosia, 1, 7.06., 4, 6, 11, 15.07., 19.08.1988, 6.07.1990, L. Esipenko; 2$, Primorye, Chernigovsky Distr., Evseevka, sweeping, 6.07.1990, L. Esipenko; 14$, Pri-morye, Spassky Distr., Prokhory, sweeping, 6.07.1990, L. Esipenko (all in FCBV). Distribution. Type locality: Russia, Primo-rye, Ryabokon Peninsula. Palaearctic: China (Hebei, Heilongjiang, Inner Mongolia, Jilin, Ningxia, Shaanxi, Xinjiang), Russia (Primo-rye), Kazakhstan, Mongolia. Dolichopus microstigma Stackelberg, 1930 Material examined. 1$, Primorye, Ussuriisky Distr., Kamenushka env., forest, 10.07.1979, R. G. Soboleva (FCBV). Distribution. Type locality: Russia, Primo-rye, Sedanka. Palaearctic: Mongolia, Russia (Primorye, Sakhalin).
Dolichopus migrans Zetterstedt, 1843 Material examined. 28$, Primorye, Khankai-sky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Koshe-leva (ZIN).
Distribution. Type locality: Sweden: Gott-landia, Nahr, Hoburg et Furillen. Trans-Pal-aearctic species.
Dolichopus nataliae Stackelberg, 1930 Material examined. 1$, Primorye, [Us-suriisk, Gorno-Tayozhnoe vil.], Gornota-yozhnaya station, arboretum, 19.06.1980, M. Mikhailovskaya (FCBV). Distribution. Type localities: Russia, Primo-rye, "Spassk-Yakovlevka road at Ugodinza (= Pyatigorka) River; Tigrovaya." Palaearctic: Russia (Khabarovsk, Magadan, Primorye, Yakutia).
Dolichopus nitidus Fallen, 1823 Material examined. 5$, Primorye, Spassky Distr., Novoselskoe, 28, 29, 30.06.1985, V. N. Makarkin; 1$, Primorye, Vladivostok, 12th km, 26.08.1986, V. N. Makarkin; 1$, Primorye, Khorol, 29.08.1986, V. N. Ma-karkin; 1$, Primorye, Chernigovsky Distr., Gornyi Khutor, sweeping, 26.07.1987, L. Es-ipenko; 1$, Primorye, Arsenyev, 12.08.1990, V. N. Makarkin (all in FCBV). Distribution. Type locality: not given (Sweden?). Trans-Palaearctic species; Oriental: China (Shanghai).
Dolichopusplumipes (Scopoli, 1763) Material examined. 2$, Primorye, Spassky Distr., Novoselskoe, 28, 29.06.1985, V. N. Makarkin; 1$, Primorye, Arsenyev, 28.06.1986, V. N. Makarkin; 3$, Primorye, Vladivostok, 12th km, 19.07.1985, V. N. Makarkin; 8$, Primorye, Arsenyev, agrolandscape, 28, 30.06.1986, V. N. Makarkin; 1$, Primorye, Chernigovsky Distr., Dmitrievka, sweeping, 1.08.1983, Arefin, Ivliev; 1$, Primorye, Khasansky Distr., Ryazanovka, 24.06.1987, V. N. Makarkin; 4$, Primorye, Chernigov-sky Distr., Evseevka, sweeping on Ambrosia, 11.06., 4.07.1988, L. Esipenko; 2$, Primorye, Chernigovsky Distr., Gornyi Khutor, sweeping on Ambrosia, 28.06., 15.07.1988, L. Esip-enko (all in FCBV); 6$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Ko-sheleva (ZIN).
Distribution. Type locality: Slovenia, "Car-nioliae indigena." Mainly Holarctic species. Neotropical: Mexico; Oriental: India (Kashmir).
Dolichopusplumitarsis Fallen, 1823 Material examined. 1$, Primorye, Pogranich-nyi Distr., Barabash-Levada env., 24.06.1980, Krivolutskaya, Moroz; 1$, Primorye, Khorol env., oak forest, 16.06.1981, V. A. Mutin; 1$, Primorye, Chernigovsky Distr., Gornyi Khutor, sweeping on Ambrosia, 15.07.1988, L. Esipen-ko (all in FCBV).
Distribution. Type locality: "Sweden." Trans-Palaearctic species; Nearctic: Canada (Ontario), USA (Alaska). Dolichopus punctum Meigen, 1824 Material examined. 8$, Primorye, Spassky Distr., Novoselskoe, 30.06.1985, V. N. Makar-kin; 1$, Primorye, Spassky Distr., [Gorno-Tayozhnoe vil.], Gornotayozhnaya station, 19.07.1990, V. N. Makarkin (all in FCBV). Distribution. Type locality: Germany, "Gegend von Berlin." Palaearctic: Austria, Finland, Germany, Poland, Russia (Khabarovsk, Leningrad, Moscow, Primorye, Yakutia), Sweden. Dolichopus rezvorum Stackelberg, 1930 Material examined. 1$, Primorye, Ke-drovaya Pad, 21.07.1976, R. G. Soboleva; 1$, Primorye, Ussuriisky Distr., Kamenushka, Komarovka River, 27.06.1979, R. G. Soboleva; 1$, Primorye, Ussuriisky Distr., Kamenushka, Barsukovka River, 3.07.1979, R. G. Soboleva; 1$, Primorye, [Ussuriisk, Gorno-Tayozhnoe vil.], Gornotayozhnaya station, arboretum, 3.07.1980, M. Mikhailovskaya (all in FCBV). Distribution. Type localities: Russia, Primo-rye, Tigrovaya; Sedanka; road Spassk-Yakov-levka at River Ugodinza (= Pyatigorka). Palae-arctic: Mongolia, Russia (Khabarovsk, Krasnoyarsk, Primorye).
Dolichopus ringdahli Stackelberg, 1930 Material examined. 4$, Primorye, Arsenyev, 27.07.1985, V. N. Makarkin (FCBV). Distribution. Type locality: Russia, Primo-rye, Tigrovaya. Palaearctic: China (Jilin), Russia (Buryatia, Murmansk, Primorye, Yakutia).
Dolichopus robustus Stackelberg, 1928 Material examined. 1$, Primorye, Ke-drovaya Pad, Kedrovaya River, 11.06.1982, R. G. Soboleva; 1$, Primorye, 5 km NW Bara-bash, valley forest, clearing, 1.06.1982, G. Lafer; 2$, Primorye, middle flow of Nezhinka River, Kabanii spring, 8.06.1986, V. N. Makarkin; 2$,
Primorye, right tributary of Razdolnaya River, upper reaches of M. Ananyevka, 10.06.1986, V. N. Makarkin; 1$, Primorye, Khasansky Distr., Ryazanovka, 14.06.1987, V. N. Makarkin (all in FCBV).
Distribution. Type localities: Russia, Primorye, "Sud-Ussuri-Gebiet, Sutshan (= Parti-zansky) Distr., Tigrovaja und Sitza (now unpopulated Narechnoe village, ~43°08'00"N 133°08'00"E); Spassk Distr., Jakovlevka und Ugodinza (= Pyatigorka) River, 20 km nach W von Jakovlevka." Palaearctic: China (Shandong), Russia (Altai Rep., Amur Region, Bury-atia, Irkutsk, Kamchatka, Khabarovsk, Krasnoyarsk, Moscow, Primorye, Yakutia). Dolichopus shamshevi Negrobov, Selivanova et Maslova, 2014
Material examined. 1$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Ko-sheleva (ZIN).
Distribution. Type locality: Russia, Kamchatka, Petropavlovsk-Kamchatsky env., Nagornyi. Palaearctic: Russia (Amur Region, Kamchatka, Khabarovsk, Krasnoyarsk, Magadan, Primo-rye, Sakhalin, Zabaikalye). Dolichopus simius Parent, 1927 Material examined. 4$, Primorye, [Ussuri-isk, Gorno-Tayozhnoe vil.], Gornotayozh-naya station, arboretum, 27, 30.06., 3.07.1980, M. Mikhailovskaya; 1$, Primorye, Vladivostok, 12th km, 19.07.1985, V. N. Makar-kin; 1$, Primorye, Arsenyev, agrolandscape, 28.06.1986, V. N. Makarkin; 1$, Primorye, Arsenyev, 28.06.1986, V. N. Makarkin; 1$, Primorye, Anisimovka, Smolnyi spring, 8.07.1986, V. N. Makarkin (all in FCBV). Distribution. Type locality: Russia, Irkutsk Region: "Siberia: environs d'Irkutsk." Palaearctic: China (Heilongjiang, Inner Mongolia), Mongolia, Russia (Altai Rep., Bashkortostan, Amur Region, Buryatia, Commander Is., Irkutsk, Kamchatka, Khabarovsk, Khakassia, Krasnoyarsk, Kuriles, Magadan, Moscow, Novosibirsk, Primorye, Sakhalin, Sverdlovsk, Tomsk, Yakutia, Zabaikalye). Dolichopus stackelbergi Smirnov, 1948 Material examined. 2$, Primorye, Spassky Distr., Novoselskoe, 29, 30.06.1985, V. N. Ma-
karkin; 2$, Primorye, Krasnoarmeisky Distr., Dersu, soya, 19-26.08.1991, V. N. Makarkin (all in FCBV); 2$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Ko-sheleva (ZIN).
Distribution. Type locality: Russia, Primorye, Nerpa (Bay), near Slavyanka. Palaearctic: Russia (Kuriles, Primorye). Dolichopus uniseta Stackelberg, 1929 Material examined. 1$, Primorye, Ussuri-isky Distr., Kamenushka, Komarovka River, 27.06.1979, R.G. Soboleva; 1$, Primorye, Ussuriisky Distr., Kamenushka, 13.07.1979, R. G. Soboleva; 2$, Primorye, Spassky Distr., Novoselskoe, 29, 30.06.1985, V. N. Makarkin; 4$, Primorye, Chernigovsky Distr., Evseevka, sweeping on Ambrosia, 7, 20.06., 4, 6.07.1988, L. Esipenko; 1$, Primorye, Chernigovsky Distr., Gornyi Khutor, sweeping on Ambrosia, 4.07.1988, L. Esipenko; 2$, Primorye, Spassky Distr., Prokhory, sweeping on Ambrosia, 6, 11.07.1990, L. Esipenko; 1$, Pri-morye, Arsenyev, 12.08.1990, V. N. Makarkin (all in FCBV); 2$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Ko-sheleva; 1$, Primorye, Partizansky Distr., Lozovyi env., station, 24.08.2019, mountain springs, Kosheleva; 1$, Primorye, Partizansk env., sweeping, 25.08.2019, mountain springs, Kosheleva (all in ZIN).
Distribution. Type localities: Russia, Yakutia and Primorye: "Kreis Jakutsk: Olom; Abyj, ungefahr 60°50' nordlicher Breite und 130° Ostlicher Lange zwischen der Lena und Amga, Amginiskaja Sloboda, amlinken Ufer des Flukes Amga; Sud-Ussuri-Gebiet: Jakov-levka, Kreis Spassk." Palaearctic: China (Heilongjiang, Hebei, Beijing, Shaanxi), Russia (Khabarovsk, Primorye, Yakutia).
Dolichopus vadimiani Negrobov et Barka-lov, 1978
Material examined. 1$, Primorye, Partizansky Mt. Range, Olkhovaya Mt., 1660 m, 8.08.1986, V. N. Makarkin (FCBV). Distribution. Type locality: Russia, Primo-rye, Sikhote-Alinsky Nature Reserve, Mayse cordon. Palaearctic: Russia (Primorye).
Dolichopus xanthopyga Stackelberg, 1930 Material examined. 1$, Primorye, Cherni-govsky Distr., Dmitrievka, soya, 3.08.1984, Arefin, Ivliev; 12$, Primorye, Chernigovsky Distr., Gornyi Khutor, sweeping on Ambrosia, 7, 28.06., 4, 11, 17.07.1988, L. Esipenko; 3$, Primorye, Chernigovsky Distr., Evseevka env., 26.07.1977, G. O. Krivolutskaya; 4$, Primorye, Chernigovsky Distr., Dmitrievka, sweeping on Ambrosia, 4, 11, 15.07.1988, L. Esipenko; 3$, Primorye, Chernigovsky Distr., Evseevka, sweeping on Ambrosia, 4.07.1988, 6.07.1990, L. Esipenko (all in FCBV); 12$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45°09'31.8"N, 131°59'58.2"E, 1-4.09.2019, lake shore, Kosheleva (ZIN). Distribution. Type localities: Russia, Pri-morye: "Yakovlevka env., Staraya Devitsa, Ryabokon Peninsula, Lefu (= Ilistaya) River mouth." Palaearctic: China (Heilongjiang), Russia (Khabarovsk, Primorye).
Genus Gymnopternus Loew, 1857
Gymnopternus daubichensis (Stackelberg, 1933)
Material examined. 1$, Primorye, Kedrovaya Pad, central station, 3.08.1976, R. G. Soboleva; 1$, Primorye, Ussuriisky Distr., Kamenushka, Barsukovka River, 3.07.1979, R. G. Soboleva; 1$, Primorye, [Ussuriisk, Gorno-Tayozhnoe vil.], Gornotayozhnaya station, Krivoi spring, 9.06.1980, M. Mikhailovskaya; 1$, Primorye, Ussuriisky Distr., Kamenushka, 7.08.1981, Belova; 2$, Primorye, Spassky Distr., Novo-selskoe, 29.06.1985, V. N. Makarkin; 1$, Pri-morye, Arsenyev, 28.06.1986, V. N. Makarkin; 6$, Primorye, Khasansky Distr., Ryazanovka, 18, 24, 25.06., 14.08.1987, V. N. Makarkin (all in coll. FCBV); 1$, Primorye, Partizansky Mt. Range, Olkhovaya Mt., 1660 m, 8.08.1986, V. N. Makarkin (ZIN ex coll. FCBV). Distribution. Type locality: Russia, Primo-rye: "Ussuri-Gebiet, Dorf Jakovlevka, Distrikt Spassk." Palaearctic: Russia (Kuriles, Primorye). Gymnopternus nemorum (Smirnov et Ne-grobov, 1977)
Material examined. 1$, Primorye, Parti-zansky Mt. Range, Olkhovaya Mt., 1660 m, 8.08.1986, V. N. Makarkin; 1$, Primorye,
Iman River, Melnichnoe vic., fir, 14.07.1991, Sidorenko; 2$, Primorye, Khasansky Distr., Ryazanovka, 23.07.1987, 25.07.1993, V. N. Ma-karkin (all in coll. FCBV); 1$, Primorye, Khan-ka Lake, 45.825°N, 131.02°E, 15-19.07.2018, N. Vikhrev; 2$, Primorye, Volchanets env., 42.908°N, 132.726°E, 1-4.08.2019, E. Erofeeva (all in ZMMU).
Distribution. Type locality: Russia, Primo-rye, Partisansk. Palaearctic: Russia (Kuriles, Primorye).
Gymnopternus pseudoceler (Stackelberg, 1933)
Material examined. 5$, Primorye, Kras-noarmeisky Distr., Dersu, soya, 19-26.08.1991, V. N. Makarkin (FCBV). Distribution. Type locality: Russia, "Primo-rye, Ussuri." Palaearctic: Russia (Kuriles, Pri-morye).
Gymnopternus rohdendorfi (Stackelberg, 1933), comb. nov.
Material examined. 1$, Primorye, Ani-simovka, 450 m, 43.13°N, 132.80°E, 2124.07.2018, N. Vikhrev (ZMMU). Distribution. Type locality: Russia: "Ussuri Gebiet, beim Dorf Jakovlevka, Distr. Spassk." Palaearctic: Russia (Khabarovsk, Primorye, Yakutia).
Notes. Hercostomus rohdendorfi characters correspond to the genus Gymnopternus (as defined by Brooks 2005), including nearly straight wing vein M, R4+5 and M subparallel, pleuron with a cluster of fine setulae in front of the posterior spiracle (not mentioned by Stackelberg 1934), fore tibia with an an-terodorsal comb-like row of strong setae, hy-popygium morphology etc. Gymnopternus ussurianus (Stackelberg, 1933)
Material examined. 1$, Primorye, Kedrovaya Pad, central station, 23.07.1976, R. G. Soboleva; 1$, Primorye, Shkotovsky Distr., Tikhookean-sky [= Fokino], 30.07.1979, Yu. D. Bodrova; 2$, Primorye, Ussuriisky Distr., Kamenushka, 7.08.1981, Belova; 5$, Primorye, Khasansky Distr., Ryazanovka, 23.07.1987, 25.07.1993, V. N. Makarkin; 5$, Primorye, Chernigovsky Distr., Gornyi Khutor, sweeping on Ambrosia, 7.06., 11.07.1988, L. Esipenko; 4$, Primo-
rye, Chernigovsky Distr., Evseevka, sweeping, 15.07.1988, L. Esipenko; 8$, Primorye, Arse-nyev, 25, 26, 27.07.1985, 10.08.1990, V. N. Ma-karkin; 1$, Primorye, Krasnoarmeisky Distr., Dersu, soya, 19-26.08.1991, V. N. Makarkin; 2$, Primorye, Iman River, Melnichnoe vic., BLP, 14.07.1991, Sidorenko (all in coll. FCBV); 1$, Primorye, Khanka Lake, 45.825°N, 131.02°E, 15-19.07.2018, N. Vikhrev; 1$, Andreevka, 42.7°N, 131.1°E, 26-31.07.2018, N. Vikhrev; 1$, Andreevka, 42.7°N, 131.1°E, 1-3.08.2018, N. Vikhrev; 3$, Ryazanovka, 42.827°N, 131.23°E, 1.08.2018, N. Vikhrev; 1$, Primorye, 10 km NE Vladivostok, 43.21°N, 132.07°E, 21-29.07.2019, E. Erofee-va (all in ZMMU).
Distribution. Type localities: Russia, "Ussuri-Gebiet, Tigrovaja, Sutshan (= Partizansky) Distr., Jakovlevka, Spassk-Distr., Basargin bei Wladiwostok, Rjabokonj am Chanka-See." Palaearctic: Japan, Russia (Amur Region, Khabarovsk, Kuriles, Primorye).
Genus Hercostomus Loew, 1857
Hercostomus beijingensis Yang, 1996 Material examined. 1$, Primorye, [Ussuri-isk, Gorno-Tayozhnoe vil.], Gornotayozhnaya station, 12.07.1990, Makarkin; 1$, Primorye, 10 km NE Vladivostok, 43.21°N, 132.07°E, 21-29.07.2019, E. Erofeeva (all in ZIN ex coll. FCBV and ZMMU).
Distribution. Type locality: China: Beijing, Yingtaogou. Palaearctic: China (Beijing, Henan, Shaanxi); Oriental: China (Hubei) Notes. The males from Primorye are very similar to H. beijingensis as described by Yang (1996), but smaller, 4.5 mm vs. 6 mm, and may belong to a new species. Hercostomus kedrovicus Negrobov et Logvi-novskii, 1977
Material examined. 8$, Primorye, Ke-drovaya Pad, central station, 21.07., 1, 3, 6.08.1976, R. G. Soboleva (FCBV). Distribution. Type locality: Russia, Primorye, Kedrovaya Pad Nature Reserve. Palaearctic: Russia (Primorye).
Hercostomus udeorum Stackelberg, 1933 Material examined. 2$, Primorye, Vladivostok, 12th km, [Akademicheskaya],
26.08.1986, V. N. Makarkin; 1$, Primorye, Khorol, 4.09.1986, V. N. Makarkin; 2$, 1?, Primorye, Krasnoarmeisky Distr., Dersu, soya, 19-26.08. 1991, V. N. Makarkin (all in coll. FCBV).
Distribution. Type locality: Russia, Primo-rye: "Ussuri-Gebiet bei der Station Tigrovaya, Distr. Sutshan (= Partizansky)." Palaearctic: Russia (Altai Rep., Amur Region, Khabarovsk, Primorye).
Genus Hydrophorus Fallen, 1823
Hydrophorus albiceps Frey, 1915 Material examined. 2?, Primorye, Vladivostok, Malaya Sedanka mouth, 43.21°N, 131.98°W, 26.09.2021, Grichanov (ZIN). Distribution. Type locality: "Eriksberg, Hifors, Viborg, Ylane, Uleaborg, Muonio, Kuusamo" [Finland, Russia]. Palaearctic: Austria, Belgium, Denmark, Finland, France, Ireland, Mongolia, Netherlands, Norway, Russia (Kamchatka, Krasnoyarsk, Leningrad, Murmansk, NW Siberia), Sweden, UK. First record from Primorye.
Genus Medetera Fischer von Waldheim, 1819
Medeterapenicillata Negrobov, 1970 Material examined. 1$, Primorye, Shkotovsky Distr., Peishula (= Suvorovka River), 1.06.1972, Arefin; 3$, Primorye, Shkotovsky Distr., imago on cedar trunk, infested by Y. sexdentatus, 1.06.1972, Arefin; 2$, Primorye, Shkotovsky Distr., from pupa in Y. sexdentatus holes, 1 .06. 1972, Arefin (all in coll. FCBV).
Distribution. Type locality: Russia, Primorye, Suputinsky (= Ussuriisky) Nature Reserve. Palaearctic: Japan, Russia (Altai Rep., Krasnoyarsk, Novosibirsk, Primorye, Tomsk).
Medetera victoris Negrobov, 1972 Material examined. 1$, 2?, Primorye, Vladivostok, Makovskogo Str. 11A, 43.20°N, 131.94°W, 22, 24.09.2021, Grichanov; 1?, Primorye, Vladivostok, 100 Let Vladivostoku Str. 159, 43.19°N, 131.92°W, 24.09.2021, Grichanov; 1?, Primorye, Vladivostok, Vereshchagina Str. 15, near Sedanka River, 43.21°N, 131.98°W, 26.09.2021, Grichanov (all in ZIN).
Distribution. Type locality: Russia: Ussuri, Sutshan Distr., Tigrovaja. Palaearctic: Russia (Primorye).
Genus Neurigona Rondani, 1856
Neurigona flavella Negrobov, 1987 Material examined. 2$, 21$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45009'31.8"N, 131059'58.2"E, 1-4.09.2019, lake shore, Kosheleva (ZIN). Distribution. Type locality: Russia, Primorye, Kedrovaya Pad Nature Reserve. Palaearctic: Russia (Kuriles, Primorye). Neurigona ninae Negrobov, 1987 Material examined. 1$, Primorye, Ussuri-isky Nature Reserve, Komarovka River valley, 5 .06. 1989, Sidorenko (FCBV). Distribution. Type locality: Russia, Primor-ye, Komarovo-Zapovednoe. Palaearctic: Russia (Primorye).
Genus Paraclius Loew, 1864
Paraclius argenteus Negrobov, 1984 Material examined. 3$, 3$, Primorye, Kedrovaya Pad, 21.07.1976, R. G. Soboleva (FCBV). Distribution. Type locality: Russia, Primo-rye, Kedrovaya Pad Nature Reserve. Palaearc-tic: Japan, Russia (Primorye).
Genus Poecilobothrus Mik, 1878
Poecilobothrus flaveolus (Negrobov et Cha-laya, 1987)
Material examined. 2$, Primorye, Kras-noarmeisky Distr., Dersu, soya, 1926.08.1991, V. N. Makarkin (FCBV); 1$, Primorye, Partizansky Distr., Lozovyi env., station, 24.08.2019, mountain springs, Koshe-leva (ZIN).
Distribution. Type locality: Russia, Primo-rye, Spassky Distr., Nakhimovka. Palaearctic: China (Beijing, Heilongjiang, Shaanxi), Japan, Russia (Amur Region, Khabarovsk, Primo-rye); Oriental: China (Hunan).
Poecilobothrus pterostichoides (Stackelberg, 1934)
Material examined. 3$, Primorye, Kedrovaya Pad, central station, 6, 7.08.1976, R. G. Soboleva; 2$, Primorye, Shkotovsky Distr., Tikhookean-sky [= Fokino], 30.07.1979, Yu. D. Bodrova (all
in coll. FCBV); 1$, Primorye, Vladivostok, Malaya Sedanka mouth, 43.21°N, 131.98°W, 26.09.2021, Grichanov (ZIN). Distribution. Type locality: Russia, Primorye: "Ussuri-Gebiet, bei der Station Tigrovaja, Sutshan" (= Ussuriiskaya oblast, near Tigrovaya station, Suchan; now Primorsky Kray, Tigro-voi village, Partizansky Distr.). Palaearctic: China (Beijing, Shaanxi), Russia (Primorye).
Genus Rhaphium Meigen, 1803
Rhaphium albifrons Zetterstedt, 1843 Material examined. 4^, Primorye, Khasan-sky Distr., Ryazanovka, 19, 20.06.1987, V. N. Makarkin (ZIN ex coll. FCBV). Distribution. Type locality: Norway: "Scandinavia boreali-Norvegia Gamaes Vaerda-liae." Trans-Palaearctic species.
Rhaphium dispar Coquillett, 1898 Material examined. 1^, Primorye, Arse-nyev, 27.07.1985, V. N. Makarkin (ZIN ex coll. FCBV).
Distribution. Type locality: Japan. Palaearctic: Japan, Russia (Kamchatka, Magadan, Primorye, Sakhalin); Oriental: China (Guizhou, Sichuan, Taiwan, Zhejiang). Rhaphium dorsiseta Tang, Wang et Yang, 2016
Material examined. 1^, Primorye, Khasan-sky Distr., Ryazanovka, 16.06.1987, V. N. Makarkin (ZIN ex coll. FCBV). Distribution. Type locality: China: Inner Mongolia, Daqinggou, Primeval forest. Palae-arctic: China (Inner Mongolia). Notes. The species is reported from Russia for the first time.
Rhaphium fascipes (Meigen, 1824) Material examined. 1^, Primorye, Ussuri-isky Reserve, W border, 130 m, 22.05.1992, V. N. Makarkin (ZIN ex coll. FCBV). Distribution. Type locality: Germany: Hamburg. Holarctic species. First record from Pri-morye.
Rhaphium flavilabre Negrobov, 1979 Material examined. 1^, Primorye, Khasan-sky Distr., Ryazanovka, 15.06.1987, V. N. Makarkin; 1^, Primorye, Vladivostok, Sedanka station, 7.06.1991, V. N. Makarkin (all in ZIN ex coll. FCBV).
Distribution. Type locality: Primorye, Komarovo-Zapovednoe, Ussuriisky Nature Reserve. Palaearctic: Russia (Khabarovsk, Pri-morye, Sakhalin).
Rhaphium mediocre (Becker, 1922) Material examined. 1$, Primorye, An-dreevka, 42.70N, 131.10E, 26-31.07.2018, N. Vikhrev (ZMMU); 1$, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45009'31.8"N, 131059'58.2"E, 1-4.09.2019, lake shore, Ko-sheleva (ZIN).
Distribution. Type locality: China: Taiwan: Taihoku [= Taipei] [and Kankau]. Palaearctic: China (Hubei, Shanghai); Oriental: China (Guizhou, Hong Kong, Taiwan, Yunnan). Notes. The species is reported from Russia for the first time.
Rhaphium micans (Meigen, 1824) Material examined. 2$, Primorye, Khasan-sky Distr., Ryazanovka, 12.06., 18.07.1987, V. N. Makarkin; 1$, Primorye, Chernigovsky Distr., Dmitrievka, soya, 4.09.1984, Arefin, Iv-liev; 1$, Primorye, Spassky Distr., Prokhory, sweeping on Ambrosia, 6.07.1990, L. Esipen-ko (all in coll. FCBV); 22$, 19?, Primorye, Khankaisky Distr., 6 km N Novokachalinsk, 45009'31.8"N, 131059'58.2"E, 1-4.09.2019, lake shore, Kosheleva (ZIN).
Distribution. Type locality: Germany: "Hamburg." Trans-Palaearctic species.
Genus Syntormon Loew, 1857
Syntormon violovitshi Negrobov, 1975 Material examined. 1$, Primorye, Krasnoarmeisky Distr., 19-26.08.1991 (ZIN ex coll. FCBV).
Distribution. Type locality: Russia, Petropav-lovsk-Kamchatsky env., Nagornyi vil. Palae-arctic: Japan, Korea, Russia (Kamchatka, Kuriles, Sakhalin). First record from Primorye.
Conclusion
Grichanov, Selivanova (2021) suggested that the dolichopodid fauna of Primorye should increase by 10%. As a result of the study of a new material from the FCBV and some other collections, new records are given for 73 species of long-legged flies. Hercostomus rohdendorfi is transferred to the genus Gymnopternus. Mesorhaga janata Negrobov is placed in synonymy with Amblypsilopus pilosus (Negrobov). Dolichophorus immacu-latus, Dolichopus henanus, D. kumazawai, Hercostomus beijingensis, Rhaphium dorsi-seta, and R. mediocre are found for the first time in Russia. Chrysotus caerulescens, Hy-drophorus albiceps, Rhaphium fascipes and Syntormon violovitshi are reported from Primorye for the first time. In total, 215 species from 31 genera of Dolichopodidae are known in the regional fauna. The number has increased by 5% during the present research.
Acknowledgements
The author is sincerely grateful to Prof. A. S. Lelej and Dr. S. Yu. Storozhenko (FCBV, Vladivostok), Drs N. E. Vikhrev and A. L. Ozerov (ZMMU, Moscow), Dr. O. Ko-sheleva (VIZR, Saint Petersburg) for their kindness in providing specimens for study. The work was funded by RFBR and NSFC according to the research project No 2054-53005. The identification of new material from expeditions to Primorye was supported by the All-Russian Institute of Plant Protection project No. 0665-2019-0014. Drs. N. Vikhrev (ZMMU) and I. Shamshev (ZIN) kindly commented on earlier draft of the manuscript.
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For citation: Grichanov, I. Ya. (2022) New records of Dolichopodidae (Diptera) from Primorsky Territory, Russia. Amurian Zoological Journal, vol. XIV, no. 1, pp. 48-60. https://www.doi.org/10.33910/2686-9519-2022-14-1-48-60
Received 16 December 2021; reviewed 27 January 2022; accepted 1 February 2022.
Для цитирования: Гричанов, И. Я. (2022) Новые находки Dolichopodidae (Diptera) в Приморском крае России. Амурский зоологический журнал, т. XIV, № 1, с. 48-60. https://www.doi.org/10.33910/2686-9519-2022-14-1-48-60
Получена 16 декабря 2021; прошла рецензирование 27 января 2022; принята 1 февраля 2022.
