Научная статья на тему 'AN ANNOTATED CHECKLIST OF DOLICHOPODIDAE (DIPTERA) SPECIES FROM SAKHALIN, RUSSIA, WITH NEW RECORDS'

AN ANNOTATED CHECKLIST OF DOLICHOPODIDAE (DIPTERA) SPECIES FROM SAKHALIN, RUSSIA, WITH NEW RECORDS Текст научной статьи по специальности «Биологические науки»

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Ключевые слова
RUSSIA / SAKHALIN ISLAND / CHECKLIST / NEW RECORDS

Аннотация научной статьи по биологическим наукам, автор научной работы — Selivanova O. V., Grichanov I. Ya.

The results of long-term studies of the dolichopodid fauna of Sakhalin Island by Russian authors are presented in the form of an annotated list of species for the first time. For this paper, originally published localities were checked, if they were referenced, and the exact locations of some collection points were established according to the museum collections. The latter material is included into the checklist. In addition, an unpublished material of Dolichopodidae is included. New records for 36 species, including 9 species new for the Sakhalin, are presented. Campsicnemus picticornis andC. scambus, Diaphorus nigricans, Dolichopus bisetulatus, Hydrophorus brunnicosus, Nepalomyia tatjanae, Rhaphium firsovi, R. tridactylum and Syntormon monochaetus are reported from Sakhalin for the first time. Sixteen species are excluded from the fauna of the island. In total, 72 species from 18 genera of long-legged flies are listed.

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Текст научной работы на тему «AN ANNOTATED CHECKLIST OF DOLICHOPODIDAE (DIPTERA) SPECIES FROM SAKHALIN, RUSSIA, WITH NEW RECORDS»

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Амурский зоологический журнал, 2022, т. XIV, № 1

Amurian Zoological Journal, 2022, vol. XIV, no. 1

www.azjournal.ru

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https://www.doi.org/10.33910/2686-9519-2022-14-1-34-47 http://zoobank.org/References/7598EEAA-BC67-4CCE-A10B-CBBB09EC692A

UDC 595.722

An annotated checklist of Dolichopodidae (Diptera) species from Sakhalin, Russia, with new records

O. V. Selivanova1, I. Ya. Grichanov

2K

1 Voronezh State University, 1 Universitetskaya Sq., 394006, Voronezh, Russia 2 All-Russian Institute of Plant Protection, 3 Podbelskogo Rd., Pushkin, 196608, Saint Petersburg, Russia

Authors

Olga V. Selivanova E-mail: selivanova-o@list.ru SPIN: 7434-0968 Scopus Author ID: 14052979200 ResearcherlD: AAI-2156-2021 ORCID: 0000-0003-0408-4435 Igor Ya. Grichanov E-mail: grichanov@mail.ru SPIN: 1438-5370 Scopus Author ID: 8672518800 ResearcherID: A-1406-2013 ORCID: 0000-0002-7887-7668

Copyright: © The Authors (2022). Published by Herzen State Pedagogical University of Russia. Open access under CC BY-NC License 4.0.

Abstract. The results of long-term studies of the dolichopodid fauna of Sakhalin Island by Russian authors are presented in the form of an annotated list of species for the first time. For this paper, originally published localities were checked, if they were referenced, and the exact locations of some collection points were established according to the museum collections. The latter material is included into the checklist. In addition, an unpublished material of Dolichopodidae is included. New records for 36 species, including 9 species new for the Sakhalin, are presented. Campsicnemus picticornis and C. scambus, Diaphorus nigricans, Dolichopus bisetulatus, Hydrophorus brunnicosus, Nepalomyia tatjanae, Rhaphium firsovi, R. tridactylum and Syntormon monochaetus are reported from Sakhalin for the first time. Sixteen species are excluded from the fauna of the island. In total, 72 species from 18 genera of long-legged flies are listed.

Keywords: Dolichopodidae, Russia, Sakhalin Island, checklist, new records.

Аннотированный список видов Dolichopodidae (Diptera)

Сахалина и новые указания

О. В. Селиванова1, И. Я. Гричанов2

1 Воронежский государственный университет, Университетская пл., д. 1, 394006, г. Воронеж, Россия 2 Всероссийский институт защиты растений, шоссе Подбельского, д. 3, г. Пушкин, 196608,

г. Санкт-Петербург, Россия

Сведения об авторах

Cеливанова Ольга Владимировна E-mail: selivanova-o@list.ru SPIN-код: 7434-0968 Scopus Author ID: 14052979200 ResearcherlD: AAI-2156-2021 ORCID: 0000-0003-0408-4435 Гричанов Игорь Яковлевич E-mail: grichanov@mail.ru SPIN-код: 1438-5370 Scopus Author ID: 8672518800 ResearcherID: A-1406-2013 ORCID: 0000-0002-7887-7668

Аннотация. Впервые представлены в виде аннотированного списка видов итоги многолетних исследований фауны мух-зеленушек Сахалина российскими авторами. Для данной статьи были проверены первоначально опубликованные местонахождения и было установлено точное местоположение некоторых пунктов сбора в соответствии с музейными этикетками. Этот материал включен в список. Кроме того, включен неопубликованный материал по Dolichopodidae. Всего приведены новые указания для 36 видов мух-зеленушек. Campsicnemus picticornis и C. scambus, Diaphorus nigricans, Dolichopus bisetulatus, Hydrophorus brunnicosus, Nepalomyia tatjanae, Rhaphium firsovi, R. tridactylum и Syntormon monochaetus впервые обнаружены на Сахалине. Шестнадцать видов исключены из островной фауны. Всего перечислено 72 вида из 18 родов мух-зеленушек.

Права: © Авторы (2022). Опубликовано Российским государственным педагогическим университетом им. А. И. Герцена. Открытый доступ на условиях лицензии СС БУ-ЫС 4.0.

Ключевые слова: Dolichopodidae, Россия, Сахалин, список, новые указания.

Introduction

Almost the whole territory of the Sakhalin Island, the largest island of Russia, is covered with dense forests, mostly coniferous (Krestov et al. 2004). It is located mainly in the Sakhalin Island Taiga ecoregion (within the Taiga bio-me) and in the Hokkaido Deciduous Forests ecoregion (within the Temperate Broadleaf and Mixed Forests biome) in the south (Ecore-gions 2017).

The diversity of the natural and climatic conditions of Sakhalin Island forms rich species diversity of entomofauna. Monsoon climate and a dense river network favour the habitation of Dolichopodidae flies preferring hydrophilic plant communities. Proximity of the Asian mainland facilitates the penetration of continental fauna. Closeness of Sakhalin to the Japanese archipelago and their connection about 20,000 years ago suggest the presence of southern Palaearctic species on the island.

The long-legged flies were usually not a group of special interest of entomologists, who visited Sakhalin. A small but valuable collection of Dr. N. A. Violovitsh (the Zoological Institute of the Russian Academy of Sciences, Saint Petersburg; ZIN) in 1955 and 1956 was used by famous dipterologist Prof. O. P. Ne-grobov (Voronezh State University, Voronezh, Russia; VSU) for the description of the first species known from Sakhalin (Negrobov 1975). Subsequently, targeted collection of Dolichopodidae material was made there (Grichanov et al. 2021) by Drs. V. D. Logvi-novsky, V. V. Zlobin and I. V. Shamshev. Based on these materials, about 30 new species were described from Sakhalin published by O. P. Negrobov and his disciples. 27 species were recorded from Sakhalin by Negrobov (1991) without material provided. No material for 16 of these species was found in the VSU collection. They are excluded from the fauna of the island.

Negrobov, Sato (2006) counted, but not listed, 48 species discovered on the Sakhalin. However, Selivanova et al. (2020) listed 13 species and 2 subspecies as endemics of the

island, and also 16 additional species reported there (original material was not provided). Argyra takagii Negrobov et Sato, 2009 from that paper was in error considered endemic of the island. Rhaphium curvitarsus Negrobov, Maslova et Selivanova, 2020 was not included into that list. The unpublished material from VSU and ZIN collections is included into the checklist below.

The information on the global distribution for each collected species follows Negrobov et al. (2013), Yang et al. (2018) and Grichanov (2021). The type localities are provided and the country lists are arranged alphabetically. The words "region" (oblast) and "territory" (kray) are omitted from the list of Russian regions. The distribution is outlined in general for Holarctic and Trans-Palaearctic species. References dealing with Sakhalin only are given after a species name.

Checklist and new records

In total, 72 species are recorded now from the Sakhalin Island that apparently make up about 50% of the actual Dolichopodidae fauna of this territory.

Genus Argyra Macquart, 1834

1. Argyra arrogans Takagi, 1960 References. Selivanova et al. 2009: 304. Distribution. Type locality: Japan, Hokkaido, Aizan-Kei. Palaearctic: Japan, Russia (Primo-rye, Sakhalin); Oriental: China.

2. Argyra igori Negrobov, Sato et Selivanova, 2012

References. Negrobov et al. 2012a: 2. Distribution. Type locality: Russia, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near settlement Urozhainoe. Palaearctic: Japan, Russia (Primorye, Sakhalin).

3. Argyra superba Takagi, 1960 References. Selivanova et al. 2009: 304. Material examined. 1^, 1$, Sakhalin Is., 50 km NE Yuzhno-Sakhalinsk, near Starodub-skoe, Shamshev, 7.08.1982. Distribution. Type locality: Japan, Hokkaido, Azian-Kei, Nukabira, Sapporo, Kamidaki, Toyama-ken, Honshu. Palaearctic: Japan, Russia (Sakhalin).

4. Argyra takagii Negrobov et Sato, 2009 References. Negrobov, Sato 2009: 73. Distribution. Type locality: Russia, Sakhalin Is., Anivsky District, Urozhainoe. Palaearctic: Japan (Hokkaido), Russia (Sakhalin, Taimyria).

5. Argyra zlobini Negrobov, Sato et Selivano-va, 2012

References. Negrobov et al. 2012a: 4. Distribution. Russia: Sakhalin Is., Anivsky District, right bank of Lyutoga River. Palae-arctic: Japan, Russia (Primorye, Sakhalin).

Genus Campsicnemus Haliday, 1851

6. Campsicnemus argyropterus Negrobov et Shamshev, 1985

References. Negrobov, Shamshev 1985: 74. Distribution. Type locality: Russia: Sakhalin Is., Yuzhno-Sakhalinsk. Palaearctic: Russia (Khabarovsk, Sakhalin).

7. Campsicnemus picticornis (Zetterstedt, 1843)

Material examined. 15^, Sakhalin Is., 38 km W Pyatirechye, Shamshev, 28.07.1982. Distribution. Type localities: Sweden: "Sue-cia meridionali et media, in Scania ad Raften; Ostrogothia ad Larketorp; Haradshammar; Holmiam." Trans-Palaearctic species. First record from Sakhalin.

8. Campsicnemus scambus (Fallen, 1823) Material examined. 2^, 4$, Sakhalin, near Yuzhno-Sakhalinsk, 29.09.1956, Violovich; 1$, Sakhalin Is., 44 km SW Yuzhno-Sakhalinsk, near Taranay, 4.08.1982, Shamshev; 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev; 1^, Sakhalin Is., 38 km W Pyatirechye, 28.07.1982, Shamshev; 2^, Sakhalin Is., 50 km NE Yuzhno-Sakhalinsk, near Starodubskoe, 7.08.1982, Shamshev.

Distribution. Type locality: Sweden, Esperod. Trans-Palaearctic species. First record from Sakhalin.

Genus Chrysotimus Loew, 1857

9. Chrysotimus spinuliferus Negrobov, 1978 References. Negrobov 1978: 1378. Distribution. Type locality: Russia, Primorye, Yakovlevka. Palaearctic: Russia (Primorye, Sakhalin, Yakutia).

Genus Chrysotus Meigen, 1824

10. Chrysotus cilipes Meigen, 1824 References. Maslova et al. 2011b: 465. Material examined. 1$, Sakhalin Is., Anivsky District, riverside Lyutoga, Zlobin, 14.07.1973; 1$, Sakhalin Is., Anivsky District, 38 km, Log-vinovskii, 17.07.1973; 1$, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk, Zlobin, 1.08.1973; 1$, Sakhalin Is., Anivsky District, Urozhainoe, Zlobin, 19.07.1978.

Distribution. Type locality: Germany, Hamburg. Trans-Palaearctic species.

11. Chrysotus cilitibia Maslova et Negrobov, 2015 References. Maslova, Negrobov 2015: 201. Distribution. Type locality: Russia: Sakhalin Is., Yuzhno-Sakhalinsk env. Palaearctic: Russia (Sakhalin, Kuriles: Kunashir).

12. Chrysotus kumazawai Negrobov, Maslova et Fursov, 2015

References. Negrobov et al. 2015: 11. Distribution. Type locality: Japan: Honshu, Ibaraki Pref., Tsukuba, Kannondai, Yatabe. Palaearctic: Japan (Honshu Island), Russia (Sakhalin).

13. Chrysotus logvinovskii Negrobov et Tsurikov, 2000

References. Negrobov et al. 2000: 231. Distribution. Type locality: Sakhalin Is., Aniv-sky District. Palaearctic: Russia (Sakhalin).

14. Chrysotus nudisetus Negrobov et Ma-slova, 1995

References. Negrobov, Maslova 1995: 460. Distribution. Type locality: Sakhalin Is., Aniv-sky District. Palaearctic: Russia (Chukotka, Khabarovsk, Magadan, Sakhalin, Primorye, Yakutia), Japan.

15. Chrysotus suavis Loew, 1857 References. Maslova et al. 2011b: 467. Distribution. Type locality: Germany: "Coln"; Austria: "Neusiedler See in Ungarn." Trans-Palearctic species.

16. Chrysotus zlobiniani Negrobov et Ma-slova, 1995

= Chrysotus zlobini Negrobov, 2000 References. Negrobov, Maslova 1995: 465. Distribution. Type locality: Sakhalin, Aniv-sky District, Lyutoga River. Palaearctic: Russia (Sakhalin, Yakutia).

Genus Diaphorus Meigen, 1824

17. Diaphorus nigricans Meigen, 1824 Material examined. 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, Shamshev, 13.07.1982.

Distribution. Type locality: "Germany." Afrotropical, Nearctic, Oriental, Palaearctic and Neotropical Regions. First record from Sakhalin.

18. Diaphorus zlobini Negrobov et Du-chanina, 1987

References. Negrobov, Duchanina 1987: 35. Distribution. Type locality: Russia: Sakhalin, Anivsky Distr., Urozhainoe. Palaearctic: Russia (Sakhalin).

Genus Diostracus Loew, 1861

19. Diostracus naegelei Negrobov, 1978 References. Negrobov 1978: 412. Material examined. 1$, Sakhalin Is., 16 km W Yuzhno-Sakhalinsk, near Chekhov Mt., 6.08.1982, Shamshev.

Distribution. Type locality: Russia: Sakhalin, Su-sunajaki Range. Palaearctic: Russia (Sakhalin).

Genus Dolichopus Latreille, 1796

20. Dolichopus amurensis Stackelberg, 1930 References. Negrobov 1991: 97 (no material provided); Maslova et al. 2011a: 91. Distribution. Type locality: Russia: "Amurlande: Banjbo, Port-Ajan" (= Khabarovsky Territory, Ayan, 56°45'N, 138°16'E). Palaearctic: Mongolia, Russia (Altai, Irkutsk, Khabarovsk, Krasnoyarsk, Primorye, Sakhalin).

21. Dolichopus bigeniculatus Parent, 1926 References. Negrobov 1991: 99 (no material provided).

Material examined. 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 27.06.1956, Violovich; 3$, Sakhalin Is., Anivsky District, Urozhainoe, 21, 27.06.1973, Logvinovskii; 1$, Sakhalin Is., 16 km W Yuzhno-Sakhalinsk, near Chekhov Mt., 19.08.1973, Logvinovskii; 1$, Sakhalin Is., 16 km W Yuzhno-Sakhalinsk, near Chekhov Mt., 15.07.1973, Zlobin; 1$, Sakhalin Is., Su-sunaisky mountain range, 9.09.1973, Zlobin; 1$, Sakhalin Is., 16 km W Yuzhno-Sakhalinsk, near Chekhov Mt., 7.09.1974, Zlobin; 1$, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk,

1.08.1973, Zlobin; 1$, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk, 15.08.1973, Zlobin; 3$, Sakhalin Is., near Novo-Aleksandrovsk, 26.07.1974, Zlobin; 1$, Sakhalin Is., 25 km SW Yuzhno-Sakhalinsk, riverside Lyutoga, 1.08.1982, Shamshev; 4$, Sakhalin Is., 9 km S Nevelsk, 23.07.1982, Shamshev; 3$, Sakhalin Is., near Yuzhno-Sakhalinsk, 5-6.07.1982, Shamshev; 22 $, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev; 11$, Sakhalin Is., 38 km W Pyati-rechye, 28.07.1982, Shamshev. Distribution. Type locality: China, Shanghai, "Zi-Ka-Wei" (= Xujiahui). Palaearctic: China (Beijing, Henan, Shaanxi, Shandong), Japan, Russia (Khabarovsk, Primorye, Sakhalin); Oriental: China (Anhui, Jiangsu, Sichuan, Zhejiang). First reliable record from Sakhalin.

22. Dolichopus bilamellatus Parent, 1929 References. Negrobov 1979: 647 (no material provided); Negrobov 1991: 99 (no material provided).

Material examined. 12$, Sakhalin Is., near Novo-Aleksandrovsk, 7.06.1973, Logvinovskii. Distribution. Type locality: Russia, "Province d'Amour." Palaearctic: Russia (Khabarovsk, Primorye, Sakhalin). First reliable record from Sakhalin.

23. Dolichopus bisetulatus Negrobov, 1977 Material examined. 1$, Sakhalin Is., near Novo-Aleksandrovsk, 22.07.1973, Zlobin. Distribution. Type locality: Russia: Primo-rye, Suputinski Reserve, Mokraya Pad. Palae-arctic: Russia (Primorye). First record from Sakhalin.

24. Dolichopus calceatus Parent, 1927 References. Negrobov 1991: 99 (no material provided).

Material examined. 1$, 1$, Sakhalin Is., 50 km NE Yuzhno-Sakhalinsk, near Starodub-skoe, 10.07.1982, Shamshev; 2$, 2$, Sakhalin Is., 38 km W Pyatirechye, 28.07.1982, Sham-shev; 6$, 3$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev; 3$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Dachnoye, 25.07.1982, Shamshev; 2$, Sakhalin Is., near Yuzhno-Sakhalinsk, 6, 7.07.1982, Shamshev; 1$, Sakhalin Is., 9 km S Nevelsk, 23.07.1982,

Shamshev; 6$, 2$, Sakhalin Is., 38 km E Aleksandrovsk-Sakhalinsky, near Tymovsky, 17-19.07.1982, Shamshev. Distribution. Type locality: Russia, Zabaika-lye: "Transbaikale; Pjetschanka (= Peschanka), b. Tschita." Palaearctic: China, Kazakhstan, Russia (Altai Rep., Amur Region, Buryatia, Irkutsk, Kamchatka, Khabarovsk, Primorye, Sakhalin, Tomsk, Yakutia, Zabaikalye). First reliable record from Sakhalin.

25. Dolichopus discifer Stannius, 1831 References. Negrobov, 1991: 109 (no material provided; as Dolichopus nigricornis Becker, 1917, nec Meigen, 1924).

Material examined. 1$, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk, 13.08.1973, Zlobin; 2$, Sakhalin Is., 38 km E Aleksandrovsk-Sakha-linsky, near Tymovsky, 17-19.07.1982, Sham-shev; 1$, Sakhalin Is., 44 km SW Yuzhno-Sakhalinsk, near Taranay 4.08.1982, Shamshev; 3$, 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Dachnoye, 25-27.07.1982, Shamshev; 2$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev.

Distribution. Type locality: Germany. Hol-arctic species known also from Chukotka, Kamchatka, Kuriles and Sakhalin Island. First reliable record from Sakhalin.

26. Dolichopus gubernator Mik, 1878 References. Negrobov 1991: 104 (no material provided).

Material examined. 1$, Sakhalin Is., 50 km NE Yuzhno-Sakhalinsk, near Starodubskoe, Shamshev, 10.07.1982; 2$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Dachnoye, 2526.07.1982, Shamshev; 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, Shamshev, 13.07.1982.

Distribution. Type locality: Austria: "bei Hammern in Mühlviertel in Oberösterreich." Palaearctic: Austria, Estonia, Finland, Latvia, Poland; Russia (Karelia, Khabarovsk, Kamchatka, Leningrad, Sakhalin, Kuriles). First reliable record from Sakhalin.

27. Dolichopus linearis Meigen, 1824 References. Negrobov 1991: 106 (no material provided).

Material examined. 2$, 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 10-12.07.1956, Violovich; 1$, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk,

I.08.1973, Zlobin; 3$, 2$, Sakhalin Is., near Novo-Aleksandrovsk, 22.07.1973, Zlobin; 4$, 8$, Sakhalin Is., near Novo-Aleksan-drovsk, 8, 22, 25-26.07.1973, Logvinovskii; 2$, 2$, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk, 1-8.08.1973, Logvinovskii; 2$, Sakhalin Is., Anivsky District, riverside Lyutoga, 13.07.1973, Logvinovskii; 12$, Sakhalin Is., near Novo-Aleksandrovsk, Zlobin, 6.09.1978; 3$, Sakhalin Is., near Novo-Aleksandrovsk, Zlobin, 14.07.1978; 1$, Sakhalin Is., Anivsky District, riverside Lyutoga, 14.07.1973, Zlobin; 3$, Sakhalin Is., Anivsky District, Urozhainoe, 19.07.1973, Zlobin; 4$, Sakhalin Is., 38 km W Pyatirechye, 28.07.1982, Shamshev; 14$, 15$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev; 2$, Sakhalin Is., near Yuzhno-Sakhalinsk,

II.08.1982, Shamshev; 1$, Sakhalin Is., 9 km S Nevelsk, 23.07.1982, Shamshev. Distribution. Type locality: not given (Germany?). Trans-Palaearctic species. First reliable record from Sakhalin.

28. Dolichopus longicornis Stannius, 1831 References. Negrobov 1991: 106 (no material provided).

Material examined. 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev; 1$, Sakhalin Is., Anivsky District, 38 km, Logvinovskii, 17.07.1973. Distribution. Type locality: not given (Germany: Hamburg?, Breslau?). Trans-Palae-arctic species. Nearctic: Canada (Yukon), USA (Alaska). First reliable record from Sakhalin.

29. Dolichopus longicostalis Negrobov et Barkalov, 1978

References. Negrobov, Barkalov 1978: 154. Material examined. 2$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev.

Distribution. Type locality: Russia: Sakhalin Is., Anivsky District, Urozhainoe. Palaearctic: Russia (Sakhalin).

30. Dolichopus microstigma Stackelberg, 1930 References. Negrobov et al. 2018: 268.

Material examined. 1$, Sakhalin Is., Ter-peniya Peninsula, Kotilovo, 70 km from Cape Terpeniya, 27.07.1956, Violovich. Type locality: Russia, Primorye, Sedanka. Palae-arctic: Mongolia, Russia (Primorye, Sakhalin).

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31. Dolichopus nigricercus Negrobov, Seli-vanova et Maslova, 2018

References. Negrobov et al. 2018: 268. Distribution. Type locality: Russia: Sakhalin Is., 38 km E Aleksandrovsk, vil. Tymovsky. Palearctic: Russia (Magadan, Kamchatka, Pri-morye, Sakhalin).

32. Dolichopus nitidus Fallen, 1823 References. Negrobov 1991: 109 (no material provided).

Material examined. 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 10.07.1956, Violovich; 1$, Sakhalin Is., Anivsky District, riverside Lyutoga, 8.08.1973, Logvinovskii; 1$, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk, 27.07.1973, Logvinovskii; 1$, Sakhalin Is., near Novo-Aleksandrovsk, 17.07.1974, Zlobin; 1? Sakhalin Is., 44 km SW Yuzhno-Sakhalinsk, near Taranay, 4.08.1982, Shamshev; 1?, Sakhalin Is., 50 km NE Yuzhno-Sakhalinsk, near Starodubskoe, 10.07.1982, Shamshev. Distribution. Type locality: not given (Sweden?). Trans-Palaearctic species; Oriental: China (Shanghai). First reliable record from Sakhalin.

33. Dolichopus plumipes (Scopoli, 1763) References. Negrobov, Rodionova 2004: 194. Distribution. Type locality: Slovenia, "Carni-oliae indigena." Mainly Holarctic species. Neotropical: Mexico; Oriental: India (Kashmir).

34. Dolichopusptenopedilus Meuffels, 1982 References. Maslova et al. 2012: 76. Material examined. 1$, Sakhalin Is., Kholmsk, 20.06.1966, Loktin; 1$, Sakhalin Is., Starodubskoe, 3.08.1991, Blagoderov. Distribution. Type locality: Japan, Hokkaido, Sounkyo. Palaearctic: Japan, Mongolia, Russia (Amur Region, Kamchatka, Khabarovsk, Krasnoyarsk, Magadan, Primorye, Sakhalin, Kuriles, Zabaikalye).

35. Dolichopus ringdahli Stackelberg, 1930 References. Negrobov 1991: 113 (no material provided).

Material examined. 2$, Sakhalin Is., Starodub-skoe, 24.07.1973, Logvinovskii; 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Dachnoye, 25.07.1982, Shamshev.

Distribution. Type locality: Russia, Primo-rye, Tigrovaya. Palaearctic: China (Jilin), Russia (Buryatia, Murmansk, Primorye, Sakhalin, Yakutia). First reliable record from Sakhalin.

36. Dolichopus shamshevi Negrobov, Seli-vanova et Maslova, 2014

References. Negrobov et al. 2014: 222. Distribution. Type locality: Russia, Kamchatka, Petropavlovsk-Kamchatsky env., Nagornyi. Palaearctic: Russia (Amur Region, Kamchatka, Khabarovsk, Krasnoyarsk, Magadan, Primorye, Sakhalin, Zabaikalye).

37. Dolichopus simius Parent, 1927 References. Negrobov 1979: 649 (no material provided); Negrobov 1991: 115 (no material provided); Kornev et al. 2013: 150. Material examined. 1?, Sakhalin Is., 50 km NE Yuzhno-Sakhalinsk, near Starodubskoe, 7.08.1982, Shamshev.

Distribution. Type locality: Russia, Irkutsk Region: "Siberia: environs d'Irkutsk." Palaearctic: China (Heilongjiang, Inner Mongolia), Mongolia, Russia (Altai Rep., Bashkortostan, Amur Region, Burya-tia, Commander Is., Irkutsk, Kamchatka, Khabarovsk, Khakassia, Krasnoyarsk, Kuriles, Magadan, Moscow, Novosibirsk, Primorye, Sakhalin, Sverdlovsk, Tomsk, Yakutia, Zabaikalye).

38. Dolichopus sinuatus Negrobov et Barka-lov, 1978

References. Negrobov, Barkalov 1978: 159; Negrobov et al. 2010: 523. Distribution. Type locality: Russia: Sakhalin, Novo-Aleksandrovsk. Palaearctic: Russia (Amur Region, Kuriles, Sakhalin).

39. Dolichopus spinuliformis Maslova, Negrobov et Selivanova, 2012

References. Maslova et al. 2012: 82. Material examined. 1$, [Sakhalin, Novo-Aleksandrovsk,] "Saghalien," Central Experimental Station, 16.07.1933, [in Japanese] (ZIN ex coll. FCBV).

Distribution. Type locality: Russia: Sakhalin Island, 32 km E Aleksandrovsk, vil. Tymovsky. Palaearctic: Russia (Sakhalin).

40. Dolichopus storozhenkoi Negrobov, Seli-vanova et Maslova, 2016

References. Negrobov et al. 2016: 34. Material examined. 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev; 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Dachnoye, 25.07.1982, Shamshev.

Distribution. Type locality: Sakhalin Region, Sakhalin Is., 25 km SW Yuzhno-Sakhalinsk, Lyutoga River. Palaearctic: Russia (Sakhalin).

41. Dolichopus ussuriensis Stackelberg, 1930 References. Negrobov 1991: 118 (no material provided).

Material examined. 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 22.06.1956, Violovich; 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 10.07.1956, Violovich; 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev.

Distribution. Type localities: Russia, Primo-rye: "Majkhe (= Shtykovo), near Shkotovo, Ti-grovaya, Spassk-Yakovlevka, Ugodinza (= Pyati-gorka) River." Palaearctic: Russia (Amur Region, Khabarovsk, Sakhalin, Primorye). First reliable record from Sakhalin.

42. Dolichopus vadimi Negrobov, Selivanova et Maslova, 2012

References. Negrobov et al. 2012: 308. Distribution. Type locality: Russia: Sakhalin, 29 km SW of Yuzhno-Sakhalinsk, Urozhai-noe. Palaearctic: Russia (Sakhalin).

43. Dolichopus verae Negrobov, 1977 References. Negrobov 1991: 118 (no material provided).

Material examined. 1$, Sakhalin Is., near Novo-Aleksandrovsk, 22.07.1973, Zlobin; 3$, Sakhalin Is., 50 km NE Yuzhno-Sakhalinsk, near Starodubskoe, 7.08.1982, Shamshev; 1$, Sakhalin Is., 38 km W Pyatirechye, 28.07.1982, Shamshev; 5$, Sakhalin Is., 25 km SW Yuzhno-Sakhalinsk, riverside Lyutoga, 1.08.1982, Shamshev.

Distribution. Type locality: Russia: Kuril Is., Iturup, Yasnyi nr. Kurilsk. Palaearctic: Russia

(Kuriles, Sakhalin). First reliable record from Sakhalin.

Genus Gymnopternus Loew, 1857

44. Gymnopternus aerosus (Fallen, 1823) References. Negrobov, Rodionova 2004: 201. Trans-Palaearctic species; Oriental: Taiwan.

Genus Hercostomus Loew, 1857

45. Hercostomus rivulorum Stackelberg, 1934 References. Nechai 2011: 108. Distribution. Type locality: Russia: "Station Tigrovaya, Sutshan-Distr., Ussuri Gebiet." Russia (Kuriles, Primorye, Sakhalin).

Genus Hydrophorus Fallen, 1823

46. Hydrophorus brunnicosus Loew, 1857 Material examined. 5$, 2$, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk, 13.08.1973, Zlobin. Distribution. Type locality: Poland: Poznan. Palaearctic: Austria, Belarus, Estonia, Finland, Poland, Russia (Crimea, Irkutsk, Krasnoyarsk, Leningrad, Lipetsk, Mordovia, Moscow, Novgorod, Novosibirsk, Orenburg, Ryazan, NW Siberia, Voronezh, ?Yakutia, Yaroslavl), Sweden. First record from Sakhalin.

Genus Medetera Fischer von Waldheim, 1819

47. Medetera sakhalinensis Negrobov et Naglis, 2015

References. Negrobov, Naglis 2015: 387. Distribution. Type locality: Russia, Sakhalin, 20 km S Yuzhno-Sakhalinsk, Dachnoye. Palae-arctic: Russia (Sakhalin).

Genus Nepalomyia Hollis, 1964

48. Nepalomyia tatjanae (Negrobov, 1984) Material examined. 2$, 9$, Sakhalin Is., 38 km W Pyatirechye, 28.07.1982, Sham-shev; 1$, 2$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev; 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 6.07.1982, Shamshev. Distribution. Type locality: Russia: Primorye, Kedrovaya Pad Reserve. Palaearctic: Russia (Primorye). First record from Sakhalin.

Genus Neurigona Rondani, 1856

49. Neurigona davshinica Negrobov, 1987 References. Negrobov 1987: 408.

Distribution. Type locality: Russia: Barguzin Reserve, Davshe. Palaearctic: Russia (Burya-tia, Sakhalin).

50. Neurigona kasparyani Negrobov, 1987 References. Negrobov 1987: 412. Distribution. Type locality: Russia: Sakhalin Is., 45 km N Yuzhno-Sakhalinsk, near Bykov. Palaearctic: Russia (Kuriles, Sakhalin), Japan.

51. Neurigona pullata Negrobov et Fursov, 1988

References. Negrobov, Fursov 1988: 407. Distribution. Type locality: Russia: Primo-rye, Kedrovaya Pad Reserve. Palaearctic: Russia (Amur Region, Primorye, Sakhalin).

Genus Rhaphium Meigen, 1803

52. Rhaphium albifrons Zetterstedt, 1843 References. Maslova et al. 2020: 140 (Mone-ron Is., Sakhalin Is.); Negrobov et al., 2020c: 90 (Moneron Is.).

Material examined. 1$, Sakhalin Is., Anivsky District, riverside Lyutoga, 13.07.1973, Logvinovskii; 1$, Sakhalin Is., Anivsky District, riverside Lyutoga, 14.07.1973, Zlobin. Distribution. Type locality: Norway: "Scandinavia boreali-Norvegia Gamaes Vaerda-liae." Trans-Palaearctic species.

53. Rhaphium boreale (Van Duzee, 1923) References. Negrobov et al. 2020a: 51. Material examined. 1$, Sakhalin Is., 38 km W Pyatirechye, 28.07.1982, Shamshev; 2$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev; 8$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Dachnoye, 25-27.07.1982, Shamshev; 2$, Sakhalin Is., near Yuzhno-Sakhalinsk, Shamshev 5-7.07.1982; 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 5.08.1982, Shamshev; 1$, Sakhalin Is., 44 km SW Yuzhno-Sakhalinsk, near Taranay, 4.08.1982, Shamshev; 1$, Sakhalin Is., 25 km SW Yuzhno-Sakhalinsk, riverside Lyutoga, 1.08.1982, Shamshev; 1$, 1? (in copula), Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev.

Distribution. Type locality: USA: Alaska, Savonoski, Naknek Lake. Palaearctic: Korea, Russia (Altai Rep., Buryatia, Krasnoyarsk, Pri-morye); Nearctic: USA (Alaska).

54. Rhaphium curvitarsus Negrobov, Ma-slova et Selivanova, 2020

References. Negrobov et al. 2020b: 22. Distribution. Type locality: Russia: Sakhalin, 41 km N Yuzhno-Sakhalinsk, village Pokrov-ka. Palaearctic: Russia (Sakhalin).

55. Rhaphium discolor Zetterstedt, 1838 References. Negrobov et al. 2020a: 52. Material examined. 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, Shamshev 11.08.1982; 3$, Sakhalin Is., 44 km SW Yuzhno-Sakhalinsk, near Taranay, 4.08.1982, Shamshev. Distribution. Trans-Palaearctic species; Nearctic: USA (Alaska).

56. Rhaphium dispar Coquillett, 1898 References. Negrobov et al. 2020a: 52. Material examined. 2$, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk, 15.08.1973, Zlobin; 1$, Sakhalin Is., near Novo-Aleksandrovsk, 7.06.1973, Logvinovskii; 1$, Sakhalin Is., Anivsky District, Urozhainoe, 20.06.1973, Logvinovskii; 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, 13.07.1982, Shamshev; 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Dachnoye, 25.07.1982, Shamshev; 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 5.08.1982, Shamshev.

Distribution. Type locality: Japan. Palaearctic: Japan, Russia (Kamchatka, Magadan, Pri-morye, Sakhalin); Oriental: China (Guizhou, Sichuan, Taiwan, Zhejiang).

57. Rhaphium firsovi Stackelberg et Ne-grobov, 1976

Material examined. 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Urozhainoe, Shamshev, 13.07.1982

Distribution. Distribution. Type locality: Russia: Primorye, Suchansky (= Partizansky) District, Tigrovaya. Palaearctic: Russia (Pri-morye). Nearctic: USA (Alaska). First record from Sakhalin.

58. Rhaphium flavilabre Negrobov, 1979 References. Negrobov et al. 2020a: 53. Material examined. 1$, Sakhalin Is., Anivsky District, 38 km, 17.07.1973, Logvinovskii; 1$, Sakhalin Is., 50 km NE Yuzhno-Sakhalinsk, near Starodubskoe, 7.08.1982, Shamshev; 1$, Sakhalin Is., near Yuzhno-Sakhalinsk,

6.07.1982, Shamshev; 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 7.08.1982, Shamshev. Distribution. Type locality: Primorye, Komaro-vo-Zapovednoe, Ussuriysky Nature Reserve. Palaearctic: Russia (Khabarovsk, Primorye, Sakhalin).

59. Rhaphium macalpini Negrobov, 1986 References. Negrobov 1986: 162. Distribution. Palaearctic: Russia (Sakhalin); Nearctic: Canada.

60. Rhaphium nasutum (Fallen, 1823) References. Negrobov et al. 2020a: 54. Distribution. Holarctic species.

61. Rhaphium neolatifacies Yang et Wang, 2006

References. Negrobov 1986: 165. Distribution. Type locality: Russia: Sakhalin, 41 km N Yuzhno-Sakhalinsk, Pokrovka. Palae-arctic: Russia (Sakhalin).

62. Rhaphium patellitarse (Becker, 1900) References. Negrobov et al. 2020a: 55. Distribution. Type locality: Russia, Khan-taika [Taimyr]. Palaearctic: Russia (Altai Rep., Buryatia, Magadan, Sakhalin, Taimyr, Ural, Yakutia, Yamalia).

63. Rhaphium richterae Negrobov, 1977 References. Negrobov et al. 2020a: 55. Material examined. 1$, Sakhalin Is., Anivsky District, Urozhainoe, 20.06.1973, Logvinov-skii; 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 3.06.1973, Logvinovskii.

Distribution. Type locality: Russia, Kuril Is., Kunashir, Tretjakovo. Palaearctic: Russia (Kuriles, Sakhalin).

64. Rhaphium sachalinense Negrobov, 1979 References. Negrobov 1979: 524. Distribution. Type locality: Russia: Sakhalin: Yuzhno-Sakhalinsk. Palaearctic: Russia (Sakhalin).

65. Rhaphium terminale igorjani Negrobov, 1986 References. Negrobov 1986: 165. Distribution. Type locality: Russia: Sakhalin, Aleksandrovsk-Sakhalinsky, Tymovsky. Palae-arctic: Russia (Sakhalin).

66. Rhaphium tridactylum (Frey, 1915) Material examined. 1$, Sakhalin Is., Anivsky District, Urozhainoe, 21.06.1973, Logvi-novskii.

Distribution. Palaearctic: Finland, Mongolia, Russia (Murmansk, Kamchatka, Khabarovsk, Taimyr), Sweden. First record from Sakhalin.

Genus Sciapus Zeller, 1842

67. Sciapus paradoxus sachalinensis Negrobov et Shamshev, 1986

References. Negrobov, Shamshev 1986: 20. Distribution. Type locality: Russia: Sakhalin Is., 50 km NE Yuzhno-Sakhalinsk, near Starodubskoe. Palaearctic: Russia (Sakhalin).

Genus Syntormon Loew, 1857

68. Syntormon monochaetus Negrobov, 1975 Material examined. 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, Violovich, 23 May 1956; 1$, Sakhalin Is., Anivsky District, riverside Lyutoga, Logvinovskii, 8.08.1973; 1$, 6$, Sakhalin Is., 16 km W Yuzhno-Sakhalinsk, near Chekhov Mt., Logvinovskii, 24.08.1973; 2$ , Sakhalin Is., 8 km N Yuzhno-Sakhalinsk, Zlobin, 13.08.1973; 1$, 3$, Sakhalin Is., near Novo-Aleksandrovsk, Zlobin, 7.08.1973; 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Dachnoye, Shamshev, 25.07.1982 Distribution. Palaearctic: Japan, Russia (Buryatia, Khabarovsk, Primorye). First record from Sakhalin.

69. Syntormon pseudopalmarae Negrobov et Shamshev, 1985

References. Negrobov, Shamshev 1985: 75. Distribution. Type locality: Russia, Sakhalin Is., Yuzhno-Sakhalinsk. Palaearctic: Russia (Sakhalin).

70. Syntormon violovitshi Negrobov, 1975 References. Negrobov 1975: 663. Material examined. 1$, Sakhalin Is., Yuzhno-Sakhalinsk, 18.06.1955, Violovich; 1$, Moneron Is., Violovich, 22.08.1956; 2$, Sakhalin Is., Yuzhno-Sakhalinsk, 30.06.1957, Violovich; 1$, Sakhalin Is., Yuzhno-Sakhalinsk, 5.07.1955, Violovich; 2$, Sakhalin Is., near Yuzhno-Sakhalinsk, 10.07.1956, Violovich; 3$, 2$, Sakhalin Is., Anivsky District, Urozhainoe, 9-11.06.1973, Logvinovskii; 10$, 9$, Sakhalin Is., Anivsky District, Urozhainoe, 18-19.06.1973, Logvinovskii; 4$, Sakhalin Is., Anivsky District, Urozhainoe, 27.06.1973, Logvinovskii; 2$, 1$, Sakhalin Is., Anivsky

District, Urozhainoe, 13-18.07.1973, Logvi-novskii; 19$, 24?, Sakhalin Is., Anivsky District, Urozhainoe, 20-24.06.1973, Logvinov-skii; 1$, Sakhalin Is., 16 km W Yuzhno-Sakhalinsk, near Chekhov Mt., 3.08.1973, Logvi-novskii; 1?, Sakhalin Is., Ogonki, 23.06.1973, Logvinovskii; 1$, Sakhalin Is., near Yuzhno-Sakhalinsk, 8.06.1973, Logvinovskii; 1?, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk, 6.07.1973, Logvinovskii; 1$, Sakhalin Is., An-ivsky District, riverside Lyutoga, 14.07.1973, Zlobin; 1?, Sakhalin Is., Starodubskoe, 24.07.1973, Zlobin; 1$, Sakhalin Is., Aniv-sky District, Urozhainoe, 07.1973, Zlobin; 5$, Sakhalin Is., 8 km N Yuzhno-Sakhalinsk, 13.08.1973, Zlobin; 2$, 1?, Sakhalin Is., near Novo-Aleksandrovsk, 7.08.1973, Zlobin; 1$, Sakhalin Is., near Novo-Aleksandrovsk, 22.07.1973, Zlobin; 1?, Sakhalin Is., near Yuzhno-Sakhalinsk, 5.08.1982, Shamshev; 1$, 14?, Sakhalin Is., near Yuzhno-Sakhalinsk, 4.07.1982, Shamshev; 1$, Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, near Dachnoye, 25.07.1982, Shamshev; 2$, 1?, Sakhalin Is., 38 km W Pyatirechye, 28.07.1982, Shamshev. Distribution. Type locality: Russia: Petropav-lovsk-Kamchatsky, Nagornyi. Palaearctic: Russia (Kamchatka, Sakhalin), Korea, Japan.

Genus Systenus Loew, 1857

71. Systenus sachalinensis Negrobov et Shamshev, 1985

References. Negrobov, Shamshev 1985: 77. Distribution. Type locality: Russia: Sakhalin Is., 29 km SW Yuzhno-Sakhalinsk, Dachnoye. Palaearctic: Russia (Sakhalin).

Genus Tachytrechus Haliday, 1851

72. Tachytrechus rubzovi Negrobov, 1976 References. Negrobov, Shamshev 1985: 80. Distribution. Type locality: China: "50 km from Mukden" [= Shenyang]. Palaearctic: China (Liaoning), Russia (Sakhalin).

SPECIES EXCLUDED FROM THE FAUNA OF SAKHALIN

Chrysotus femoratus Zetterstedt, 1843 Notes. The record of this species (Ne-grobov 1991: 72) without material provided may belong to one of the new species de-

scribed later. No material was found in the VSU collection.

Diaphorusparenti Stackelberg, 1928 Notes. No material was found in the VSU collection for the record of this species from Sakhalin (Negrobov 1991: 70). The species is spread in mainland East Asian Palaearctic.

Dolichopus basalis Loew, 1859 Notes. This species was recorded from Sakhalin by Negrobov (1979: 647) without material provided. Maslova et al. (2012) noted that this record belongs to Dolichopus spinuliformis. Dolichopus basalis is widespread in mainland East Asian Palaearctic.

Dolichopus cilifemoratus Macquart, 1827 Notes. This species was recorded from Sakhalin by Negrobov (1991: 100) without material provided. No material was found in the VSU collection.

Dolichopus divisus Becker, 1917 Notes. This species was also recorded from Sakhalin by Negrobov (1991: 102) without material provided. No material was found in the VSU collection. The species is spread in mainland East Asian Palaearctic. Dolichopus migrans Zetterstedt, 1843 Notes. This species was also recorded from Sakhalin by Negrobov (1991: 108) without material provided. No material was found in the VSU collection. This Trans-Palaearctic species can be found in Sakhalin. Dolichopus notatus Staeger, 1842 Notes. This species was also recorded from Sakhalin by Negrobov (1991: 110) without material provided. No material was found in the VSU collection. This Trans-Palaearctic species can be found in Sakhalin. Dolichopus pennatus Meigen, 1824 Notes. This species was recorded from Sakhalin by Negrobov (1991: 111) without material provided. No material was found in the VSU collection. Dolichopusplumitarsis Fallen, 1823 Notes. This species was recorded from Sakhalin by Negrobov (1979: 649, 1991: 112) without material provided. No material was found in the VSU collection. This Holarctic species can be found in Sakhalin.

Dolichopusportentosus Negrobov, 1973 Notes. This species was recorded from Sakhalin by Negrobov (1991: 112) without material provided. No material was found in the VSU collection. The species is spread in mainland East Asian Palaearctic. Dolichopus robustus Stackelberg, 1928 Notes. This species was also recorded from Sakhalin by Negrobov (1991: 113) without material provided. No material was found in the VSU collection. The species is spread mainly in mainland East Asian Palaearctic. Dolichopus sagittarius Loew, 1848 Notes. This species was also recorded from Sakhalin by Negrobov (1991: 114) without material provided. No material was found in the VSU collection. This East Palaearctic and West Nearctic species can be found in Sakhalin. Dolichopus setimanus Smirnov, 1948 Notes. This species was recorded from Sakhalin by Negrobov (1991: 114) without material provided. Describing their new species, Dolichopus storozhenkoi (Negrobov et al. 2016), the

authors used the material collected from Kuriles and Sakhalin and listed under D. setimanus (Selivanova et al. 2010). They noted that these records belong to D. storozhenkoi. So, D. setimanus is excluded from the list of Dolichopodidae recorded from Sakhalin. The latter species is spread in mainland East Asian Palaearctic. Dolichopus sharovi Smirnov, 1948 Notes. This species was recorded from Sakhalin by Negrobov (1991: 115) without material provided. It is known from Kamchatka and Primorye and can be found in Sakhalin. Dolichopus varians Smirnov, 1948 Notes. This species was recorded from Sakhalin by Negrobov (1991: 118) without material provided. It is known from Korea, Kamchatka, Khabarovsk Territory, Kuriles and Primo-rye and can be found in Sakhalin. Dolichopus xanthopyga Stackelberg, 1930 Notes. This species was also recorded from Sakhalin by Negrobov (1991: 119) without material provided. No material was found in the VSU collection. The species is spread in mainland East Asian Palaearctic.

References

Ecoregions. (2017) [Online]. Available at: https://ecoregions2017.appspot.com (accessed 30.12.2021). (In English)

Grichanov, I. Ya. (2021) Alphabetic list of generic and specific names of predatory flies of the epifamily Dolichopodoidae (Diptera). Saint Petersburg: All-Russian Research Institute of Plant Protection. [Online]. Available at: http://grichanov.aiq.ru/Genera3.htm/ (accessed 30.12.2021). (In English) Grichanov, I. Ya., Maslova, O. O., Selivanova, O. O., Kornev, I. I. (2021) Vklad shkoly professora O. P. Negrobova v izuchenie Dolichopodidae (Diptera) Dal'nego Vostoka Rossii [Contribution of the Professor O. P. Negrobov's school to the study of Dolichopodidae (Diptera) of the Russian Far East]. In: Itogi i perspekktivy razvitiya entomologii v Vostochnoj Evrope: sbornik statej IV Mezhdunarodnoj nauchno-prakkticheskoj konferentsii, posvyashchennojpamyati Aleksandra Mikhajlovicha Tereshkina (1953-2020), 1-3 dekabrya 2021 g. Minsk [Results and prospects for the development of entomology in Eastern Europe: Collection of articles of the IV International Scientific and Practical Conference dedicated to the memory of Alexander Mikhailovich Tereshkin (1953-2020), December 1-3, 2021, Minsk]. Minsk: A. N. Varaksin Publ., pp. 95-99. (In Russian) Kornev, I. I., Negrobov, O. P., Selivanova, O. V. (2013) Novye dannye po faune i sistematike Dolichopus simius Parent, 1927 (Dolichopodidae, Diptera) [New data on the distribution and systematic of Dolichopus simius Parent, 1927 (Dolichopodidae, Diptera)]. Amurskij zoologicheskij zhurnal — Amurian Zoological Journal, vol. V, no. 2, pp. 147-150. (In Russian) Krestov, P. V., Barkalov, V. Y., Taran, A. A. (2004) Botaniko-geograficheskoe rajonirovanie ostrova Sakhalin [Phytogeographical zoning of the Sakhalin Island]. In: Rastitelnyj i zhivotnyj mir ostrova Sakhalin [Flora and fauna of Sakhalin Island]. Vladivostok: Dal'nauka Publ., pp. 67-92. (In Russian) Maslova, O. O., Negrobov, O. P. (2015) Novyj vid Chrysotus Meigen, 1824 (Dolichopodidae: Diptera) s Sakhalina i Kuril'skikh ostrovov [New species of Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Sakhalin and Kuril Islands (Russia)]. Kavkazskij entomologicheskij byulleten' — Caucasian Entomological Bulletin, vol. 11, no. 1, pp. 201-203. (In Russian)

Maslova, O. O., Negrobov, O. P., Kornev, I. I. (2011a) Nekotorye dannye po faune vidov roda Dolichopus Latr. (Diptera, Dolichopodidae) Sibiri i Dal'nego Vostoka Rossii [Some data on the fauna of species of the genus Dolichopus Latr. (Diptera, Dolichopodidae) of Siberia and the Russian Far East]. In: Sovremennye problemy entomologii [Modern problems of entomology]. Voronezh: Voronezh State University Publ., pp. 90-93. (In Russian) Maslova, O. O., Negrobov, O. P., Selivanova, O. V. (2011b) Fauna roda Chrysotus Meigen (Diptera, Dolichopodidae) Rossii. Chast' 1. Vidy gruppy Ch. cilipesMeigen i Ch. laesus Wied [Russian fauna of the genus Chrysotus Meigen (Diptera, Dolichopodidae). Part 1. Group of species Chrysotus cilipes Meigen and Ch. laesus Wied]. Entomologicheskoe obozrenie, vol. 90, no. 2, pp. 464-468. (In Russian)

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Maslova, O. O., Negrobov, O. P., Selivanova, O. V. (2012) The first records of Dolichopus ptenopedilus (Dolichopodidae, Diptera) from Russia and Mongolia with description of its female. Amurskij zoologicheskij zhurnal — Amurian Zoological Journal, vol. IV, no. 1, pp. 76-78. (In English) Maslova, O. O., Negrobov, O. P., Selivanova, O. V. (2020) New records of Rhaphium (Dolichopodidae, Diptera) from Russian protected areas. Nature Conservation Research, vol. 5, no. 3, pp. 139-144. https://www.doi.org/10.24189/ncr.2020.037 (In English) Nechai, N. A. (2011) Sistematika, fauna i zoogeografiya vidov roda Hercostomus Loew (Dolichopodidae, Diptera) Palearlüicheskoj fauny [Taxonomy, fauna, and zoogeography of the genus Hercostomus Loew (Dolichopodidae, Diptera) of the Palaearctic]. PhD dissertation (Biology). Saint Petersburg, All-Russian Institute of Plant Protection of Russian Academy of Agricultural Sciences, 178 p. (In Russian) Negrobov, O. P. (1975) Obzor mukh-zelenushek roda Syntormon Meig. (Dolichopodidae, Diptera) fauny Palearktiki [A review of the Palearctic species of the genus Syntormon (Diptera Dolichopodidae)]. Entomologicheskoe obozrenie, vol. 54, no. 3, pp. 652-664. (In Russian) Negrobov, O. P. (1977-1979a) Dolichopodidae, Unterfamilie Hydrophorinae, Unterfamilie Rhaphiinae. In: E. Lindner (ed.). Die Fliegen der Palaearktischen Region. Vol. 29. Lf. 316 (1977), 319 (1978), 321322 (1979a). Stuttgart: E. Schweizerbart Verlag, pp. 354-530. (In German) Negrobov, O. P. (1978) Rody gruppy Chrysotimus Fallen (Dolichopodidae, Diptera) fauny SSSR [Genera of group Chrysotimus Fallen (Dolichopodidae, Diptera) of the USSR fauna]. Zoologicheskii zhurnal, vol. 57, no. 9, pp. 1374-1381. (In Russian) Negrobov, O. P. (1979b) Dvukrylye sem. Dolichopodidae (Diptera) fauny SSSR. I. Podsemejstva Dolichopodinae i Medeterinae [Family Dolichopodidae (Diptera) of the fauna of the USSR. I. Subfamilies Dolichopodinae and Medeterinae]. Entomologicheskoe obozrenie, vol. 58, no. 3, pp. 646-659. (In Russian) Negrobov, O. P. (1986) Golarkticheskie svyazi fauny semejstva dolikhopodid [Holarctic relations of family Dolichopodidae (Diptera)]. In: Biogeografiya Beringiiskogo sektora Subarktiki: Materialy X Vsesoyuznogo simpoziuma, Magadan, 1983 [Biogeography of the Beringian sector of the Subarctic: Materials of the XAll-Union symposium, Magadan, 1983]. Vladivostok: Far Eastern Scientific Center of the Academy of Sciences of The USSR Publ., pp. 161-168. (In Russian) Negrobov, O. P. (1987) Novye palearkticheskie vidy dolikhopodid roda Neurigona Rond. (Diptera, Dolichopodidae) [New palaearctic species of the dolichopodid genus Neurigona Rond. (Diptera, Dolichopodidae)]. Entomologicheskoe obozrenie, vol. 66, no. 2, pp. 406-415. (In Russian) Negrobov, O. P. (1991) Family Dolichopodidae. In: A. Soos, L. Papp (eds.). Catalogue of Palaearctic Diptera. Vol. 7: Dolichopodidae — Platypezidae. Budapest: Akademiai Publ., pp. 11-139. https://doi. org/10.1016/B978-0-444-98731-0.50008-9 (In English) Negrobov, O. P., Barkalov, A. V. (1978) Novye vidy roda Dolichopus Latr. (Dolichopodidae, Diptera) Sibiri, Primor'ya i Sakhalina [Species of the genus Dolichopus Latr. (Diptera, Dolichopodidae) of Siberia, Primorye and Sakhalin]. In: Taksonomiya i ekologiya chlenistonogikh Sibiri [Taxonomy and ecology of Siberian Arthropoda]. Novosibirsk: Nauka Publ., pp. 154-162. (In Russian) Negrobov, O. P., Barkalov, A. V., Selivanova, O. V. (2010) New data on the fauna of the family Dolichopodidae (Diptera) from Russia, with a description of a new species of the genus Argyra Mcq. Euroasian Entomological Journal, vol. 9, no. 3, pp. 522-524. (In English) Negrobov, O. P., Dukhanina, E. V. (1987) Vidy roda Diaphorus Meigen (Dolichopodidae, Diptera) Dal'nego Vostoka SSSR [Species of the genus Diaphorus Meig. (Dolichopodidae, Diptera) of the Soviet Far East]. Nauchnye doklady vysshejshkoly. Biologicheskie nauki, no. 1, pp. 35-38. (In Russian) Negrobov, O. P., Fursov, V. N. (1988) Reviziya vidov roda Neurigona Rond. (Diptera, Dolichopodidae) Palearktiki. II [Revision of species of the genus Neurigona Rond. (Diptera, Dolichopodidae) of Palaearctic. II]. Entomologicheskoe obozrenie, vol. 67, no. 2, pp. 405-416. (In Russian) Negrobov, O. P., Maslova, O. O. (1995) Reviziya palearkticheskikh vidov roda Chrysotus Mg. (Diptera, Dolichopodidae). II [A revision of the Palaearctic species of the genus Chrysotus Mg. (Diptera, Dolichopodidae). II]. Entomologicheskoe obozrenie, vol. 74, no. 2, pp. 456-466. (In Russian)

Negrobov, O. P., Maslova, O. O., Chursina, M. A. (2020a) New records of Rhaphium (Dolichopodidae, Diptera) from Russia. Acta Biologica Sibirica, vol. 6, pp. 49-57. https://www.doi.org/10.3897/ abs.6.e53125 (In English)

Negrobov, O. P., Maslova, O. O., Fursov, V. N. (2015) New data on the genus Chrysotus Meigen, 1824 (Diptera:

Dolichopodidae) from Japan and Russia. Far Eastern Entomologist, no. 293, pp. 10-15. (In English) Negrobov, O. P., Maslova, O. O., Selivanova, O. V. (2020b) A new species of the genus Rhaphium (Diptera:

Dolichopodidae) from Sakhalin Island. Far Eastern Entomologist, no. 409, pp. 21-25. (In English) Negrobov, O. P., Naglis, S. (2015) Two new species of Medetera Fischer von Waldheim (Diptera, Dolichopodidae) from Russia and Mongolia. Zootaxa, vol. 3964, no. 3, pp. 386-390. https://doi. org/10.11646/zootaxa.3964.3.8 (In English) Negrobov, O. P., Obona, J., Manko, P., Maslova, O. O. (2020c) New faunistics notes on the fauna and variability of Rhaphium albifrons Zetterstedt, 1843 (Dolichopodidae: Diptera). In: Prostranstvenno-vremennye aspekkty funkktsionirovaniya biosistem. Sbornik materialov XVIMezhdunarodnoj nauchnoj ekologicheskoj konferentsii, posvyashchennoi pamyati Aleksandra Vladimirovicha Prisnogo [Spatiotemporal aspects of the functioning of biosystems. Collection of materials of the XVI International Scientific Environmental Conference dedicated to the memory of Alexander Vladimirovich Prisny]. Belgorod: Belgorod State University Publ., pp. 89-90. (In English) Negrobov, O. P., Rodionova, S. Y. (2004) New data on fauna ofsubfamily Dolichopodinae (Dolichopodidae, Diptera) in Russia and neighbouring territories (genus Hercostomus Lw.). An International Journal of Dipterological Research, vol. 15, no. 3, pp. 201-204. (In English) Negrobov, O. P., Sato, M. (2006) The comparative characteristic of fauna of family Dolichopodidae of Japan and Far East of Russia. In: 6th International Congress of Dipterology, Fukujoka, Japan, 2006. Fukujoka: s. n., p. 178. (In English) Negrobov, O. P., Sato, M. (2009) New species of the genus Argyra Macquart, 1834 (Diptera, Dolichopodidae)

from the Far East of Russia and Japan. Dipterists Digest, vol. 16, no. 1, pp. 73-79. (In English) Negrobov, O. P., Sato, M., Selivanova, O. V. (2012a) New species of the genus Argyra Macquart, 1834 (Diptera: Dolichopodidae) from the Russian Far East and Japan. Far Eastern Entomologist, no. 247, pp. 1-7. (In English)

Negrobov, O. P., Selivanova, O. V., Maslova, O. O. (2012b) A new species of Dolichopus (Diptera: Dolichopodidae) from Sakhalin and designation of lectotype of Dolichopus grunini Smirnov, 1948. Kavkazskij entomologicheskij byulleten' — Caucasian Entomological Bulletin, vol. 8, no. 2, pp. 308310. (In English)

Negrobov, O. P., Selivanova, O. V., Maslova, O. O. (2014) Novye dannye po sistematike palearkticheskikh vidov gruppy Dolichopus longisetus Negrobov, 1977 (Diptera, Dolichopodidae) [New data on systematics of Palaearctic species of the group Dolichopus longisetus Negrobov 1977 (Diptera, Dolichopodidae)]. Zoologicheskii zhurnal, vol. 93, no. 2, pp. 221-227. http://dx.doi.org/10.7868/ S0044513414020081 (In Russian) Negrobov, O. P., Selivanova, O. V., Maslova, O. O. (2016) Novyj vid iz roda Dolichopus Latr. (Diptera, Dolichopodidae) s Sakhalina i Kuril'skikh ostrovov [A new species of the genus Dolichopus Latr. (Diptera, Dolichopodidae) from Sakhalin and Kuril Islands]. Byulleten' Moskovskogo obshchestva ispytatelej prirody. Otdel biologicheskij — Bulletin of Moscow Society of Naturalists. Biological Series, vol. 121, no. 5, pp. 33-36. (In Russian). Negrobov, O. P., Selivanova, O. V., Maslova, O. O. (2018) Novye dannye po sistematike gruppy vidov Dolichopus lepidus Staeger, 1842 (Diptera: Dolichopodidae) [New data on the taxonomy of the group of species Dolichopus lepidus Staeger, 1842 (Diptera: Dolichopodidae)]. Kavkazskij entomologicheskij byulleten' — Caucasian Entomological Bulletin, vol. 14, no. 2, pp. 267-272. https://www.doi. org/10.23885/181433262018142-267272 (In Russian) Negrobov, O. P., Selivanova, O. V., Maslova, O. O., Chursina, M. A. (2013b) Check-list of predatory flies of the family Dolichopodidae (Diptera) in the fauna of Russia. In: I. Ya. Grichanov, O. P. Negrobov (eds.). Fauna i taksonomiya khishchnykh mukh Dolichopodidae (Diptera). Sbornik nauchnykh rabot [Fauna and taxonomy of Dolichopodidae (Diptera). Collection of papers]. Saint Petersburg: VIZR RAAS Publ., pp. 47-93. (Plant Protection News. Supplements). (In English) Negrobov, O. P., Shamshev, I. V. (1985) Novye svedeniya o faune dolikhopodid (Diptera, Dolichopodidae) Sakhalina [New data on the Dolichopodidae (Diptera) fauna of Sakhalin]. In: A. I. Cherepanov (ed.). Sistematika i biologiya chlenistonogikh i gel'mintov [Systematics and biology of arthropods and helminths]. Novosibirsk: Nauka Publ., pp. 74-80. (in Russian). Negrobov, O. P., Shamshev, I. V. (1986b) Novye vidy roda Sciapus Zeller (Dolichopodidae, Diptera) iz Sibiri [New species of the genus Sciapus Zeller (Dolichopodidae, Diptera) from Siberia]. Trudy Zoologicheskogo instituta Akadenii nauk SSSR, vol. 146, pp. 17-22. (In Russian)

Negrobov, O. P., Tsurikov, M. N., Maslova, O. O. (2000) Reviziya palearkticheskikh vidov roda Chrysotus Mg. (Diptera, Dolichopodidae), III [Revision of the Palaearctic species of the genus Chrysotus Mg. (Diptera, Dolichopodidae), III]. Entomologicheskoe obozrenie, vol. 79, no. 1, pp. 227-238. (In Russian)

Selivanova, O. V., Maslova, O. O., Negrobov, O. P. (2020) Obzor fauny semejstva Dolichopodidae Sakhalina [Review of the fauna of the family Dolichopodidae (Diptera) of Sakhalin]. In: XI Vserossiiskij dipterologicheskij simpozium (s mezhdunarodnym uchastiem), Voronezh, 24-29 avgusta 2020 g.: sbornik materialov [XI All-Russian Dipterological Symposium (with international participation), Voronezh, 24-29 August 2020: Collection of materials]. Saint Petersburg: LEMA Publ., pp. 204-207. (In Russian)

Selivanova, O. V., Negrobov, O. P., Maslova, O. O. (2009) Novye dannye o vidakh roda Argyra Macquart, 1834 (Dolichopodidae, Diptera) v faune Rossii [New data on species of the genus Argyra Macquart, 1834 (Dolichopodidae, Diptera) in the fauna of Russia]. Vestnik zoologii, vol. 43, no. 4, p. 304. (In Russian)

Yang, D., Zhang, L. L., Zhang, K. Y. (2018) Species catalogue of China. Vol. 2. Animals, Insecta (VI), Diptera (2), Orthorrhaphous Brachycera. Beijing: Science Press, 387 p. (In English)

For citation: Selivanova, O. V., Grichanov, I. Ya. (2022) An annotated checklist of Dolichopodidae (Diptera) species from Sakhalin, Russia, with new records. Amurian Zoological Journal, vol. XIV, no. 1, pp. 34-47. https://www. doi.org/10.33910/2686-9519-2022-14-1-34-47

Received 10 January 2022; reviewed 31 January 2022; accepted 15 February 2022.

Для цитирования: Селиванова, О. В., Гричанов, И. Я. (2022) Аннотированный список видов Dolichopodidae (Diptera) Сахалина и новые указания. Амурский зоологический журнал, т. XIV, № 1, с. 34-47. https://www. doi.org/10.33910/2686-9519-2022-14-1-34-47

Получена 10 января 2022; прошла рецензирование 31 января 2022; принята 15 февраля 2022.

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