NEW EXPLORATION IN KORYAKIA - THE LICHENS OF THE CAPE GOVEN, BERING SEA COAST (NORTHERN KAMCHATKA, RUSSIA) Текст научной статьи по специальности «Биологические науки»

New exploration in Koryakia — the lichens of the Cape Goven, Bering Sea coast (Northern Kamchatka, Russia)

D. E. Himelbrant1, 2, I. S. Stepanchikova1, 2, T. Ahti3, V. Yu. Neshataeva2

'St. Petersburg State University, St. Petersburg, Russia 2Komarov Botanical Institute of the Russian Academy of Sciences, St. Petersburg, Russia 3Botanical Museum, Finnish Museum of Natural History, University of Helsinki, Finland Corresponding author. D. E. Himelbrant, [email protected]

Abstract. The lichen diversity of the Cape Goven within the Koryak State Reserve counts 394 species. 373 lichens, 18 lichenicolous fungi and 3 non-lichenized saprobic fungi related to lichens. Altogether 4 species are new to Russia (Miriquidica pulvinatula, Myriolecis andrewii, Ochrolechia alaskana, Rhizocarpon sublavatum), 1 — to Asiatic Russia (Collemopsidium foveola-tum), 29 other species are new to the Russian Far East, 4 — to the northern part of the Far East. Additionally, 51 other species are new to Kamchatka Territory, and 92 more are new to Koryakia. Among the new species to Russia or Russian Far East, 11 are also reported for the first time for Be-ringia. A total of 500 species of lichens and allied fungi are known from Koryakia now. The richest habitats in Cape Goven are rocky outcrops and tundras; unlike in the earlier explored Parapolsky Dale, shrublands, floodplain stands and bogs play relatively insignificant role in the lichen diversity. The lichens of seashore communities enrich the lichen flora of Cape Goven compared to inland areas. The lichen diversity of Cape Goven is significantly higher than in Parapolsky Dale due to its mountainous landscape and coastal position.

Keywords: lichen flora, new records, Far East, Beringia, Koryak State Reserve.

Новое исследование в Корякии — лишайники мыса Говена на побережье Берингова моря (Северная Камчатка, Россия)

Д. Е. Гимельбрант1, 2, И. С. Степанчикова1, 2, Т. Ахти3, В. Ю. Нешатаева2

1Санкт-Петербургский государственный университет, Санкт-Петербург, Россия 2Ботанический институт им. В. Л. Комарова РАН, Санкт-Петербург, Россия

3Ботанический музей, Финский музей естественной истории, университет Хельсинки,

Финляндия

Автор для переписки. Д. Е. Гимельбрант, [email protected]

Резюме. Выявленное разнообразие лишайников кластера «Мыс Говена» Корякского государственного заповедника насчитывает 394 вида. 373 вида лишайников, 18 видов лихенофиль-ных грибов и 3 вида нелихенизированных сапротрофных грибов. Из них 4 вида — новые для России (Miriquidica pulvinatula, Myriolecis andrewii, Ochrolechia alaskana, Rhizocarpon sublavatum), 1 — для Азии (Collemopsidium foveolatum), еще 29 видов — для Дальнего Востока, 4 — для северной части Дальнего Востока; всего 11 видов впервые обнаружены в пределах Берингии. Еще 51 вид является новым для Камчатского края, кроме того, 92 вида впервые отмечены для Корякии. К настоящему моменту для Корякии известно 500 видов лишайников и связанных с ними грибов. Самыми богатыми местообитаниями мыса Говена являются скальные обнажения

https://doii.org/1031111/Knr/2021551.121

121

и тундры; в отличие от Парапольского Дола, кустарниковые сообщества, пойменные древостой и болота играют относительно незначительную роль в разнообразии лишайников. Виды, характерные для приморских сообществ, обогащают лихенофлору мыса Говена. В отношении лишайников мыс Говена значительно богаче, чем Парапольский Дол, благодаря ландшафту и прибрежному положению.

Ключевые слова: лихенофлора, новые находки, Дальний Восток, Берингия, Корякский государственный заповедник.

Our new research is devoted to the further study on the lichen diversity in the understudied territory of Koryakia, which we started previously (Himelbrant et al., 2019). The most important information about Koryakia and the Koryak State Reserve can be found in the cited article on the "Parapolsky dol" (Parapolsky Dale), one of three parts of the reserve. Another recently visited part of the reserve is "Mys Govena" (Cape Goven), which is located in the southern part of the Goven Peninsula between Korf and Olutorsky Gulfs of the Bering Sea (Fig. 1). The investigated part, Cape Goven (452 km2), lies in the NE Koryakia and represents part of the low-mountain Pylginsky Range (part of Koryakskoe Highland) with maximal elevations up to 521 m a. s. l. (up to 1240 m in the main part of the range outside the study area) and coastal lowlands up to 20 m a. s. l. The main landscape types of Cape Goven are low mountains, heavily dissected by small rivers and creeks, as well as sea terraces and steep coasts 20-30 m high formed by sedimentary material, separate cliffs descending directly into the sea, and narrow (mainly up to 20 m wide) sand-pebble beaches. Cape Goven is situated within the hypoarctic zone. In the mountains different types of tundras prevail. Most common are lichen- and moss-dwarf shrub tundras with Arctous alpina (L.) Niedenzu, Betula divaricata Ledeb., B. exilis Sukacz., Campanula lasiocarpa Cham., Carex van-heurkii Muell., Chamaepericlymenum suecicum (L.) Aschers. et Graebn., Dianthus repens Willd., Diapensia obovata (Fr. Schmidt) Nakai, Hierochloe alpina (Sw.) Roem. et Schult., Empetrum nigrum L., Ledum decumbens (Ait.) Lodd. ex Steud., Loiseleu-riaprocumbens (L.) Desv., Luzula confusa Lindeb., Oxytropis nigrescens (Pall.) Fisch., Rhododendron camtschaticum Pall., Salix sphenophylla A. K. Skvortsov, Saxifraga cherlerioides D. Don, Trisetum spicatum (L.) K. Richt., Vaccinium uliginosum L., V. vi-tis-idaea L., and different species of terricolous lichens. Forest vegetation is absolutely absent, but shrublands are represented by Pinus pumila (Pallas) Regel (with Rhododendron aureum Georgi) distributed mainly on slopes of the local hills and mountains up to 190 m a. s. l., as well as Alnus fruticosa Rupr. and rarely Sorbus sambucifolia (Cham. et Schltdl.) M. Roem. on southern slopes and in downstream parts of the river or creek valleys. Salix alaxensis (Andersson) Coville, S. pulchra Cham., and S. udensis Trautv. et C. A. Mey. shrublands and subalpine meadows are also confined to downstream parts of valleys. Individual examples of Betula ermanii Cham. x B. divaricata can be found in the river valleys (Katranzhi, 2017; Neshataev et al., 2017; Neshataeva, Neshataev, 2018). Specific vegetation of seabird colonies is developed on rocky cliffs, especially in vicinities of the lighthouse at the tip of the Cape.

Fig. 1. Map of the study area. Northern boundary of Cape Goven part of the Koryak State Reserve

is shown as a dashed line.

Special lichenological studies at the Cape Goven of the Koryak State Reserve were not conducted ever before, but 64 most common and widespread lichens were men-

tioned in phytosociological papers (Neshataev et al., 2017; Neshataeva, Neshataev, 2018; Neshataeva et al., 2018) on the basis of our collections and identifications.

Material and Methods

The specimens were collected by Dmitry E. Himelbrant (DH) and Irina S. Ste-panchikova (IS) in August 2018 at the Cape Goven within the Koryak State Reserve and its conservation zone (Olutorsky District of Kamchatka Territory). Altogether 57 localities were investigated (Fig. 1): 34 standard sample plots (10 x 10 m, otherwise in natural boundaries of the community), where the lichen diversity on each substrate was described as detailed as possible, and 24 additional plots, where only individual substrates and species were recorded. Valentina Yu. Neshataeva provided phytosociological characteristics of sample plots and the study area in general. All geographical coordinates are given in the coordinate system WGS 1984.

Chromatography was performed by IS according to the standard techniques of high performance thin-layer chromatography using solvent systems A, B, C, and G (Orange et al., 2001). The nomenclature of taxa generally follows recently published checklists for Scandinavia and North America (Nordin et al., 2011; Esslinger, 2019) and special taxonomic papers and monographs (Dombrovskaya, 1996; Kukwa, 2011; Sheard et al., 2017; Spribille et al., 2020), as well as for lichenicolous fungi (Diederich et al., 2018; Lawrey, Diederich, 2018). The specimens were mainly identified by DH, IS, and Teuvo Ahti (Cladonia spp.), if otherwise, the identifiers' names are indicated in the species list. Representative specimens are deposited in the herbaria of the Botanical Museum of University of Helsinki (H), Senckenberg Natural History Museum in Frankfurt (FR), Komarov Botanical Institute RAS (LE), and Department of Botany, St. Petersburg State University (LECB).

Sampling locations

Standard sample areas. Koryak State Reserve, Goven Peninsula: 1 — NW of Yuzhnaya Hill, ca. 200 m of the seashore, top of the local height, disintegrating butte, 59°49'15.2"N, 166°05'47.9"E, 58 m a. s. l., rocky outcrops, 8 VIII 2018, Kor-Goven-01; 2 — SW of Yuzhnaya Hill, Cape Goven, W shore, 59°48'12.9-33.3"N, 166°05'31.0-39.1"E, 12 m a. s. l., seashore rocks, 9 VIII 2018, Kor-Goven-02; 3 — ibid., lighthouse, 59°48'13.3"N, 166°05'37.9"E, 66 m a. s. l., concrete constructions, 9 VIII 2018, Kor-Goven-03; 4 — ibid, E of lightouse, slope of the stream valley, 59°48'13.5"N, 166°05'59.1"E, 82 m a. s. l., tundra with dwarf shrubs and lichens, 9 VIII 2018, Kor-Goven-04; 5 — S of Yuzhnaya Hill, top of local height, butte, 59°48'26.9"N, 166°07'36.5"E, 276 m a. s. l., open rocky outcrops surrounded by dwarf shrub tundra, 9 VIII 2018, Kor-Goven-05; 6 — W of Yuzhnaya Hill, upper part of local ridge, 59°48'44.9"N, 166°06'11.0"E, 132 m a. s. l., mosaic tundra on gravel, 11 VIII 2018, Kor-Goven-06; 7 — ibid, slightly inclined plane above seashore scarp, 59°48'44.9"N, 166°05'38.2"E, 26 m a. s. l., seashore tundra with dwarf shrubs and lichens, 11 VIII 2018, Kor-Goven-07; 8 — NW of Yuzhnaya Hill, N slope of stream valley, 59°49'22.8"N, 166°06'11.6"E, 26 m a. s. l., Pinus pumila community with mosses, Rhododendron aureum, and Chamaepericlymenum suecicum, 11 VIII 2018, Kor-Goven-08; 9 — NW of Yuzhnaya Hill, S slope of stream valley, 59°49'23.2"N, 166°06'02.6"E, 21 m a. s. l., Alnus fruticosa shrubs (up to 2-3 m high) with Calamagrostis purpurea (Trin.) Trin. and Spiraea beauverdiana Schneid., 11 VIII 2018, Kor-Goven-09; 10 — N of Yuzhnaya Hill, top of ridge, 59°49'42.4"N, 166°07'35.5"E, 214 m a. s. l., lichen community on rocky outcrops with Dia-pensia obovata and grasses, 12 VIII 2018, Kor-Goven-10; 11 — NE of Yuzhnaya Hill, near the top of

unnamed hill, SE slope, 59°49'48.6"N, 166°08'41.9"E, 260 m a. s. l., tundra with lichens and dwarf shrubs in mosaic with bare soil and short (to 10 cm high) Alnus fruticosa shrubs, 12 VIII 2018, Kor-Goven-11; 12 — ibid, top of unnamed hill (329.6 m), 59°49'18.0"N, 166°08'16.2"E, 329 m a. s. l., spots of dwarf shrub tundra on gravel, 12 VIII 2018, Kor-Goven-12; 13 — top of W ridge of Yuzhnaya Hill, 59°48'47.7"N, 166°06'53.3"E, 195 m a. s. l., Pinuspumila shrubs (35-40 cm high) on the upper part of S slope, 12 VIII 2018, Kor-Goven-13; 14 — between capes Goven and Primetny, top of local height close of the seashore, 59°50'44.8"N, 166°06'28.9"E, 135 m a. s. l., tundra with lichens, dwarf shrubs and grass, 13 VIII 2018, Kor-Goven-14; 15 — ibid., W slope of ridge along the seashore, 59°50'31.8"N, 166°06'28.6"E, 68 m a. s. l., dwarf shrub tundra with lichens and short (to 15 cm high) Alnus fruticosa shrubs, on slope faced to the sea, 13 VIII 2018, Kor-Goven-15; 16 — ibid, SE slope of ridge near the seashore, 59°50'22.6"N, 166°06'17.4"E, 45 m a. s. l., Alnus fruticosa shrubs (ca. 2 m high) with Sor-bus sambucifolia and Calamagrostispurpurea, 13 VIII 2018, Kor-Goven-16; 17 — same place, stream valley, 59°50'20.2"N, 166°06'21.5"E, 5 m a. s. l., steep rocky slope and bottom of the valley, nival community, 13 VIII 2018, Kor-Goven-17; 18 — NE of Yuzhnaya Hill, bottom of the stream valley, 59°49'29.1"N, 166°05'45.6"E, 5 m a. s. l., Salix alaxensis floodplain stand with Calamagrostis purpurea, 14 VIII 2018, Kor-Goven-18; 19 — S of Cape Galinvilan, S slope of the stream valley, 50 m of the seashore, 60°00'19.3"N, 166°10'18.8"E, ca. 0 m a. s. l., Alnus fruticosa shrubs (3.0-3.5 m high) with horsetails and ferns, 16 VIII 2018, Kor-Galin-19; 20 — ibid, 60°00'19.3"N, 166°10'18.5"E, 14 m a. s. l., dwarf shrub tundra with Vaccinium uliginosum and Empetrum nigrum, 16 VIII 2018, Kor-Galin-20; 21 — S of Cape Galinvilan, 100 m of the seashore, accumulative plane, 60°00'24.3"N, 166°10'16.8"E, 19 m a. s. l., seashore tundra with lichens and dwarf shrubs, 16 VIII 2018, Kor-Galin-21; 22 — SE of Cape Galinvilan, accumulative plane, 60°00'27.6"N, 166°10'37.1"E, 24 m a. s. l., hilly tundra with lichens, dwarf shrubs and Carex sp., 16 VIII 2018, Kor-Galin-22; 23 — N of the stream falling into the Valkavayam River mouth, accumulative plane, 59°58'39.1"N, 166°10'54.1"E, 16 m a. s. l., dwarf shrub tundra with lichens, 17 VIII 2018, Kor-Galin-23; 24 — S of the stream falling into the Valkavayam River mouth, slightly sloping accumulative plane, 59°58'41.0"N, 166°12'13.4"E, 31 m a. s. l., dwarf shrub tundra with Betula exilis, Vaccinium uliginosum, and lichens, 17 VIII 2018, Kor-Galin-24; 25 — valley of the stream falling into the Valkavayam River mouth, 59°58'41.4"N, 166°12'10.0"E, 27 m a. s. l., Salix pulchra floodplain stand with Calamagrostis purpurea, Chamerion angustifolium (L.) Holub, and Viratrum oxysepalum Turcz., 17 VIII 2018, Kor-Galin-25; 26 — N of the Kuyakynvayam River, accumulative plane, 59°59'18.9"N, 166°11'22.8"E, 19 m a. s. l., hilly tundra with dwarf shrubs, 17 VIII 2018, Kor-Galin-26; 27 — valley of the stream N of the Kuyakynvayam River, 60°00'24.7"N, 166°10'59.4"E, 27 m a. s. l., Alnus fruticosa and Sorbus sambucifolia shrubs with several trees of Betula ermanii, with Calamagrostis purpurea and ferns, 18 VIII 2018, Kor-Galin-27; 28 — E of Cape Galinvilan, accumulative plane, 60°00'49.7"N, 166°10'36.8"E, 18 m a. s. l., Betula divaricata shrubs (up to 1.5 m high) with Empetrum nigrum, Chamaepericlymenum suecicum, Calamagrostis purpurea, Spiraea beauverdiana, and horsetails, 18 VIII 2018, Kor-Galin-28; 29 — ibid., accumulative plane, 60°00'47.2"N, 166°11'44.1"E, 34 m a. s. l., open Pinus pumila community with Betula divaricata, Vaccinium uliginosum, and Empetrum nigrum, 18 VIII 2018, Kor-Galin-29; 30 — ibid, accumulative plane, 60°00'50.3"N, 166°11'35.6"E, 27 m a. s. l., tundra with lichens and dwarf shrubs, 18 VIII 2018, Kor-Galin-30. Conservation zone of Koryak State Reserve, Goven Peninsula: 31 — Cape Cha-yachy (Paryn), 60°04'09.4"N, 166°10'58.9"E, 19 m a. s. l., seashore cliff and rocky outcrops nearby, 19 VIII 2018, Kor-Chayachy-31; 32 — SE of Cape Peschany and W of lake Atijulgytgyn, regressive plane, 60°06'58.8"N, 166°13'47.2"E, 10 m a. s. l., tundra with lichens and dwarf shrubs, 19 VIII 2018, Kor- Chayachy-32.

Aditional collection points. Koryak State Reserve, Goven Peninsula: a1 — SWW of Yuzhnaya Hill, Cape Goven, W shore, 59°48'47.7"N, 166°05'23.6"E, 8 m a. s. l., seashore rocks (littoral), 9 VIII 2018, Kor-Goven-01ad; a2 — ibid, E of the lightouse, top of rigde between two streams, 59°48'13.6"N, 166°06'13.5"E, 120 m a. s. l., rocky outcrops, 9 VIII 2018, Kor-Goven-02ad; a3 — NW of Yuzhnaya

Hill, slopes and bottom of the stream valley, 59°49'01.5"N, 166°05'35.6"E, 16 m a. s. l., moss-lichen communities on slopes, driftwood in the stream and seashore tundra, 10 VIII 2018, Kor-Goven-03ad; a4 — ibid., slightly inclined plane above seashore scarp, 59°49'09.6"N, 166°05'49.0"E, 32 m a. s. l., mosaic of lichen tundra and primary lichen communities on bare soil, 11 VIII 2018, Kor-Goven-04ad; a5 — ibid., bottom of stream valley, 59°49'22.8"N, 166°06'02.9"E, 19 m a. s. l., dwarf shrub tundra with Potentilla fruticosa L., 11 VIII 2018, Kor-Goven-05ad; a6 — W of Yuzhnaya Hill, top of the local height, butte, 59°49'20.2"N, 166°06'06.8"E, 49 m a. s. l., open rocky outcrops covered with lichens, 11 VIII 2018, Kor-Goven-06ad; a7 — NE of Yuzhnaya Hill, E slope of unnamed hill, butte, 59°49'49.0"N, 166°08'48.5"E, 264 m a. s. l., open rocky outcrops on ridge near the top of hill, 12 VIII 2018, Kor-Goven-07ad; a8 — ibid, 59°49'26.2-31.2"N, 166°08'26.8-37.0"E, 237 m a. s. l., open rocky outcrops on ridge, 12 VIII 2018, Kor-Goven-08ad; a9 — between capes Goven and Primetny, top of ridge along the seashore, 59°50'39.7"N, 166°06'28.7"E, 101 m a. s. l., rocky outcrops, Alnus fruticosa and Pinuspumila shrubs and dwarf shrub tundra, 13 VIII 2018, Kor-Goven-09ad; a10 — ibid., W slope of stream valley, 59°50'14.7"N, 166°06'12.1"E, 9 m a. s. l., tundra with Empetrum nigrum, Vaccinium uliginosum, and lichens, 13 VIII 2018, Kor-Goven-10ad; a11 — ibid, slightly inclined plane above seashore scarp, 59°50'03.8"N, 166°05'49.3"E, 17 m a. s. l., seashore tundra with dwarf shrubs, lichens, and Carex spp., 13 VIII 2018, Kor-Goven-11ad; a12 — NE of Yuzhnaya Hill, slightly inclined plane above seashore scarp, 59°49'27.1"N, 166°05'37.8"E, 8 m a. s. l., seashore tundra with dwarf shrubs, 13 VIII 2018, Kor-Goven-12ad; a13 — SE of Cape Galinvilan, accumulative plane, 60°00'25.1"N, 166°10'31.2"E, 24 m a. s. l., Sphagnum swamp with dwarf shrubs and Carex spp., 16 VIII 2018, Kor-Galin-13ad; a14 — NE of Cape Galinvilan, mouth of stream falling into the sea, 60°01'04.5"N, 166°10'31.6"E, 3 m a. s. l., driftwood on the river bank (large log), 16 VIII 2018, Kor-Galin-14ad; a15 — ibid, seashore, 60°00'56.9"N, 166°10'09.1"E, ca. 0 m a. s. l., driftwood on supralittoral, 16 VIII 2018, Kor-Galin-15ad; a16 — N of Kuyakynvayam, mouth of stream falling into the sea, steep S slope of stream valley, 59°59'48.8"N, 166°10'23.8"E, ca. 0 m a. s. l., mosaic tundra with Salix spp., Empetrum nigrum, and grasses on bare soil, 17 VIII 2018, Kor-Galin-16ad; a17 — valley of the stream falling into the Valkavayam River mouth, 59°58'40.6"N, 166°12'08.0"E, 26 m a. s. l., tundra with Vaccinium uliginosum, 17 VIII 2018, Kor-Galin-17ad; a18 — N of the stream falling into the Valkavayam River mouth, accumulative plane, 59°58'49.4"N, 166°11'57.4"E, 24 m a. s. l., hilly tundra with dwarf shrubs and lichens, 17 VIII 2018, Kor-Galin-18ad; a19 — valley of the Kuyakynvayam River, 59°59'11.7"N, 166°11'32.4"E, 20 m a. s. l., nival lowland under the slope, lichen communities on gravel and sand, 17 VIII 2018, Kor-Galin-19ad; a20 — S of Cape Galinvilan, just above seashore scarp, 60°00'27.0"N, 166°10'11.9"E, 24 m a. s. l., tundra with dwarf shrubs and lichens, 17 VIII 2018, Kor-Galin-20ad; a21 — SE of Cape Galinvilan, accumulative plane, 60°00'57.9"N, 166°10'57.0"E, 20 m a. s. l., Sphagnum swamp with dwarf shrubs, 18 VIII 2018, Kor-Galin-21ad; a22 — Cape Galivilan, stream valley, 60°00'50.6"N, 166°09'59.9"E, ca. 0 m a. s. l., lichen tundra, 18 VIII 2018, Kor-Galin-22ad. Conservation zone of Koryak State Reserve, Goven Peninsula: a23 — N of Cape Chayachy (Paryn), between the seashore and small lake, 60°05'15.8"N, 166°12'13.9"E, 4 m a. s. l., wooden posts on seashore meadow, 19 VIII 2018, Kor-Chayachy-23ad; a24 — SE of Cape Peschany and W of lake Atijulgytgyn, regressive plane, vicinity of former fish factory no. 7, 60°08'11.7"N, 166°15'52.2"E, 10 m a. s. l., tundra with dwarf shrubs and lichens, 19 VIII 2018, Kor-Chayachy-24ad.

Results and Discussion

An annotated list of species of Cape Goven and vicinities is presented below. The species reported for the first time for the Kamchatka Territory are marked with !, for the Koryakia — with *, lichenicolous fungi with #, non-lichenized fungi with +, and species protected by both regional (Krasnaya... , 2018) and federal (Krasnaya... , 2008) Red Data Books — with R. Each species is accompanied with the list of sub-

strates and localities. The frequency of occurrence within the study area was estimated and is indicated in square brackets: [RJ "rare — single record" — the species was found once, [R] "rare" — recorded in 2-6 standard sample plots or more than in one additional plot, [O] "occasionally" — 7-12 standard sample plots, [F] "frequent" — 13-19, [C] "common" and [VC] "very common" — 20-34 (the latter categories are absent in the analyzed material). For the species already reported from the Cape Goven (Neshataev et al., 2017; Neshataeva, Neshataev, 2018; Ne-shataeva et al., 2018) the references are given. For the species not known before from the Koryakia or Kamchatka Territory information on presence in the neighboring regions is provided. Abbreviations of the neighboring regions are following: Ala — Alaska, Chu — Chukotka Autonomous Area, Kam — Kamchatka Territory (within limits of former Kamchatka Region including Commander Islands), Mag — Magadan Region, Yak — Republic of Sakha (Yakutia). References are given after each abbreviation (in brackets), one for each region. Lichen substances are given for TLC-analyzed specimens. For the species new to the Russian Far East or larger regions the most important identification characters are represented. Special comments are given when needed.

#Abrothallus parmeliarum (Sommerf.) Arnold — on thalli of Parmelia skultii over rocks, soil and plant debris; 1, 5, a6, a8 [R]; H, FR.

!Absconditella sphagnorum Vezda et Poelt — on mosses; 22 [RJ; H. — Yak (Chesnokov et al, 2016), Ala (Spribille et al, 2020). New to the Russian Far East. Differs from other species of the genus by growing over peat mosses and having one-septate spores (The lichens..., 2009).

Acarospora fuscata (Schrad.) Th. Fr. — on rocks; 1, a6 [R]; H, FR.

!A. moenium (Vain.) Räsänen — on concrete of a lighthouse; 3 [RJ; LECB (sub Candelariella sp.). — New to Beringia and Russian Far East. The nearest reported localities in Russia belong to Krasnoyarsk Territory (Sedelnikova, 2013). Widespread but probably overlooked sterile saxicolous lichen with squamulose-areolate, gray, often (also in our material) white-pruinose thallus bearing characteristic blackish soredia on margins of areoles.

*A. sinopica (Wahlenb.) Körb. — on rocks; 1 [RJ; FR (sub Amandinea cacuminum). — Kam (Kuznetsova, Himelbrant, 2006), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

!A. squamulosa (Schrad.) Trevis. — on rocks; 10 [RJ; LECB. — Ala (Talbot et al, 1997). New to the Russian Far East. The nearest known localities in Russia are Sokhondinsky Reserve within Trans-Baikal Territory (Yakovchenko, 2009) and Krasnoyarsk Territory (Sedelnikova, 2013). Saxicolous areolate lichen with thick, brown, epruinose, C+ red thallus and rugulate disc of roundish apothecia; for detailed description see Knudsen et al. (2019).

Adelolecia pilati (Hepp) Hertel et Hafellner — on rocks; 5 [RJ; H.

Alectoria ochroleuca (Hoffm.) A. Massal. — on soil; 1, 4-6, 10-12, 15, 23, a7 [O]; FR (Neshataev et al, 2017; Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

*Allantoparmelia alpicola (Th. Fr.) Essl. — on rocks; 5 [RJ; H. — Kam (Himelbrant et al,, 2014), Chu (Andreev et al,, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2010).

*A. sibirica (Zahlbr.) Essl. — on rocks; a8 [RJ; H (Plate I: 1a, 1b). — Kam (Mikulin, 1988), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2011).

Amandinea cacuminum (Th. Fr.) H. Mayrhofer et Sheard — on rocks; 1, 10, a6, a7 [R]; H, FR.

*A. coniops (Wahlenb.) M. Choisy ex Scheid. et H. Mayrhofer — on seashore rocks; 2, 31 [R]; H. — Kam (Trass, 1963), Chu (Andreev et al., 1996), Yak (Andreev et al., 1996), Ala (Geiser et al., 1998).

A. punctata (Hoffm.) Coppins et Scheid. — on plant debris, rocks, turf and soil over rocks, bark of Vaccinium uliginosum, driftwood; 1, 10, 14, 31, a2, a6, a23 [R]; H, FR, LECB.

!Anema jenisejense H. Magn. — on soil; 1 [R1]; H. — New to Beringia and Russian Far East. The species is rarely reported: besides the type locality in Krasnoyarsk Territory, it is known in Russia also from several Siberian regions (Sedelnikova, 2013) and Urals (Paukov, Trapeznikova, 2005); the records from North America are accounted as erroneous (Esslinger, 2019). Our material has relatively small (up to 7 mm in diam.), centrally attached, laciniate, fruticulose thalli without any generative structures, morphologically it corresponds with the protologue (Magnusson, 1936) and available specimens in H (Plate I: 2).

Arctocetraria andrejevii (Oxner) Kärnefelt et A. Thell — on soil; 7, 17, 21-23, 30, 32, a12, a13 [O]; H, FR, LECB (Neshataeva, Neshataev, 2018; Neshataeva et al,, 2018).

*Arctomia delicatula Th. Fr. — on soil; 5 [R1]; H. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

*Arctoparmelia centrifuga (L.) Hale — on rocks; 5, a8 [R]. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2010).

A. separata (Th. Fr.) Hale — on rocks; 1, 5, 10 [R]; LECB.

*Arthonia apatetica (A. Massal.) Th. Fr. — on bark of Betula divaricata, Salix alaxensis, Sorbus sambucifolia; 18, 27, 28 [R]; LECB. — Kam (Neshataeva et al, 2004), Ala (Spribille et al, 2010).

!A. ligniariella Coppins — on driftwood; a24 [R1]; LECB. — Ala (Spribille et al, 2020). New to the Russian Far East. The nearest known localities in Russia belong to Komi Republic (Krasnaya..., 1998). Mostly lignicolous species, recognized by its non-amyloid hymenium and relatively small one-septate spores (The lichens..., 2009).

*A. mediella Nyl. — on bark of Alnus fruticosa and Betula ermanii; 9, 19, 27 [R]; LECB. — Kam (Neshataeva et al, 2004), Ala (Thomson, 1997).

!#A. varians (Davies) Nyl. — on thallus of Lecanora rupicola on rock; a2 [R1]; LECB. — Chu (Andreev et al, 1996), Yak (Zhurbenko, 2009b).

Arthopyrenia analepta (Ach.) A. Massal. — on bark of Alnus fruticosa, Betula ermanii, B. exilis, Salixpulchra; 9, 15-17, 19, 25, 27, a24 [O]; LECB.

*A. atomariella (Nyl.) Hulting — on bark of Vaccinium uliginosum; 14 [R1]; H. — Kam (Neshataeva et al., 2003).

!A. cinereopruinosa (Schaer.) A. Massal. — on bark of Salix sp.; 11 [R1]; H. — New to Beringia and Russian Far East. The nearest reported localities in Russia belong to Krasnoyarsk Territory (Sedelnikova, 2013). Close to A. analepta, but differs from that species in a few characters, first of all spores which are constricted at septum and have a median constriction in each cell (The lichens... , 2009).

!#cf. Arthrorhaphis aeruginosa R. Sant. et Tonsberg — on thallus of Cladonia cervicornis on soil; 10 [R1]; H. — Ala (Spribille et al, 2010). New to the Russian Far East. The nearest published locality in Russia belongs to Trans-Baikal Territory (Zhurbenko, Pino-Bodas, 2017). The species is probably widespread in boreal and Arctic regions, but is usually sterile (Zhurbenko, Pino-Bodas, 2017). Our material is sterile as well, however, the fungus seems to be easily recognized: parasitized basal squamules of Cladonia spp. obviously turn dark blue-green.

A. alpina (Schaer.) R. Sant. — on soil over rocks; 1, 12 [R]; LECB.

A. citrinella (Ach.) Poelt — on soil; 10, 17, a8 [R].

Asahinea chrysantha (Tuck.) W. L. Culb. et C. F. Culb. — on soil and rocks; 1, 5, 6, 11, 15 [R] (Neshataeva, Neshataev, 2018).

Aspicilia cinerea (L.) Körb. — on rocks; 5, 6, 10, 31 [R]; H.

!Aspilidea myrinii (Fr.) Hafellner — on rocks; a7 [R1]; LECB. — Mag (Kotlov, 1995), Yak (Chesnokov et al, 2016), Ala (Thomson, 1997).

Athallia pyracea (Ach.) Arup et al. — on bark of Betula ermanii; 27 [R1].

*#Bachmanniomyces punctum (A. Massal.) Diederich et Pino-Bodas — on thallus of Cladonia cornuta subsp. cornuta on soil; a12 [R1]; H. — Kam (Zhurbenko et al, 2012), Yak (Zhurbenko et al, 2005), Ala (Spribille et al, 2010).

!#B. uncialicola (Zopf) D. Hawksw. — on thalli of Cladonia amaurocraea, C. pyxidata, C. uncialis on soil; 10, 11, 14, a8, a12 [R]; H, FR. — Chu (Zhurbenko, Pino-Bodas, 2017), Ala (Spribille et al, 2020).

*Bacidia bagliettoana (A. Massal. et De Not.) Jatta — on plant debris; 5, 17, a8 [R]; H. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

Baeomyces carneus Flörke — on soil and gravel; 4, 6, 7, 12, 21 [R]; LECB (sub Carbonea vorticosa) (Neshataeva, Neshataev, 2018).

B. placophyllus Ach. — on soil; 5, 10 [R]; H (sub Catapyrenium cinereum).

B. rufus (Huds.) Rebent. — on soil; 7, 17 [R]; H (Neshataeva, Neshataev, 2018).

*Bellemerea cinereorufescens (Ach.) Clauzade et Cl. Roux — on rocks; 31 [R1]; H (sub Lecidea subpaupercula). — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

!B. sanguinea (Kremp.) Hafellner et Cl. Roux — on rocks; 17 [R1]; LECB. — New to Beringia and Russian Far East. The nearest reported locality in Russia is Pechora-Ilych Reserve in Komi Republic (Hermansson et al, 2006). Differs from B. cinereorufescens by thin thallus with abundant apothecia, higher hymenium, and larger spores (Foucard, 2001).

Biatora efflorescens (Hedl.) Räsänen — on bark of Alnus fruticosa and Betula ermanii; 19, 27 [R]; LECB.

B. pallens (Kullh.) Printzen — on bark of Pinuspumila and Potentilla fruticosa; 8, 24 [R]; LECB.

*Bilimbia microcarpa (Th. Fr.) Th. Fr. — on soil and plant debris; 17, 31 [R]; LECB. — Kam (Neshataeva et al, 2005), Ala (Spribille et al, 2020).

Blastenia ammiospila (Wahlenb.) Arup et al. — on soil and plant debris, bark of Alnus fruticosa, driftwood; 1, 10, 15, 21, a3, a7, a15, a16, a19 [R]; LECB (conf. I. Frolov).

!Brigantiaea fuscolutea (Dicks.) R. Sant. — on soil; a16 [R1]; H. — Chu (Andreev et al, 1996), Ala (Spribille et al, 2020).

!Bryobilimbia diapensiae (Th. Fr.) Fryday et al. — on dead cushions of Diapensia obovata; 6, 11 [R]; H, LECB. — Chu (Andreev et al, 1996), Yak (Poryadina, 2005), Ala (Thomson, 1997).

B. hypnorum (Lib.) Fryday et al. — on plant debris; 5, 26 [R]; LECB.

Bryocaulon divergens (Ach.) Kärnefelt — on soil and rocks; 1, 4-6, 10-12, 14, 15, 23, a2, a6, a8 [O]; H (sub Ramalina almquistii) (Neshataev et al, 2017; Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

*Bryonora castanea (Hepp) Poelt — on soil and plant debris; 11, 12, 17 [R]; H, FR. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2010).

*Bryoplaca jungermanniae (Vahl) Sochting et al. — on soil; 10 [RJ; LECB. — Kam (Mikulin, 1990), Chu (Frolov, Gagarina, 2020), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Thomson, 1997). The species was reported for Kamchatka by Mikulin (1990), but we couldn't find any relevant specimens in LE and VLA. There are also specimens from Kamchatka by Savicz (LE) which were misidentified as Placodium jungermanniae (Vahl) Tuck., however, they were never published under this name (see Khodosovtsev et al, 2004). The specimen from Bering Island by Almquist (S F-101920) reported by Nylander (1887) as Lecanora jungermanniae (Vahl) Nyl. refers to Caloplaca stillicidiorum.

*B. tetraspora (Nyl.) Sochting et al. — on mosses; 31 [RJ; LECB. — Kam (Khodosovtsev et al, 2004), Chu (Frolov, Gagarina, 2020), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

Bryoria nitidula (Th. Fr.) Brodo et D. Hawksw. — on soil and rocks; 1, 5, 10, 14, 21, 23, a6, a8 [R]; H (Neshataev et al, 2017; Neshataeva, Neshataev, 2018).

!Bryostigma muscigenum (Th. Fr.) Frisch et G. Thor — on bark of Alnus fruticosa and wood of Betula ermanii; 27 [R1]; LECB. — Chu (Andreev et al, 1996), Yak (Andreev et al,

1996), Ala (Spribille et al, 2010).

*Buellia aethalea (Ach.) Th. Fr. — on rocks and gravel; 1, 5, 10, 12, a2 [R]; H. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Thomson,

1997).

B. ectolechioides (Vain.) Erichsen — on rocks and gravel; 1, 3, 5, 6, 10, 14, 23, a2, a12 [O]; H (Neshataeva, Neshataev, 2018).

B. erubescens Arnold — on bark of Salix alaxensis and Sorbus sambucifolia; 18, 27 [R]; LECB.

!Caloplaca alcarum Poelt — on rocks, growing together with Lecanora sp.; 31 [R1]; LECB (conf. Frolov). — Chu (Kristinsson et al, 2010), Ala (Arup, 1995).

C. borealis (Vain.) Poelt — on bark of Alnus fruticosa, Betula divaricata, B. ermanii, B. exilis, Potentilla fruticosa, Salix alaxensis, S. pulchra, Sorbus sambucifolia, Vaccinium uliginosum; 9, 14-16, 18, 19, 24-27 [O]; LECB (det. or conf. Frolov).

*C. caesiorufella (Nyl.) Zahlbr. — on bark of Potentilla fruticosa and Salix alaxensis; 18, 21, 26, a14 [R]; LECB (det. Frolov). — Kam (Khodosovtsev et al, 2004), Chu (Frolov, Gagarina, 2020), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

!C. nivalis (Körb.) Th. Fr. — on mosses; a8 [R1]; LECB. — Ala (Spribille et al, 2020). New to the Russian Far East. The nearest known locality in Russia is Taimyr Peninsula in Krasnoyarsk Territory (Andreev et al, 1996). The species is easily recognized by having elongate (vermicular) spores with poorly developed septum (Zhurbenko, Sochting, 1993).

C. stillicidiorum (Vahl) Lynge — on bones, plant debris, and mosses; 1, 2, 5, 10, 31, a6-a8 [R]; LECB (conf. Frolov).

!C. tornoensis H. Magn. — on mosses; 17 [R1]; LECB. — Ala (Spribille et al, 2010). New to the Russian Far East. The nearest reported localities in Russia belong to Krasnoyarsk (Zhurbenko, Sochting, 1993) and Trans-Baikal (Chesnokov et al, 2016) territories. The species is morphologically close to C. nivalis, but has different spores; for detailed description see Zhurbenko, Sochting (1993).

*Calvitimela aglaea (Sommerf.) Hafellner — on rocks; 1, 5, 6 [R]; H, LECB. — Kam (Mikulin, 1988), Chu (Hertel, Andreev, 2003), Mag (Kotlov, 1995), Yak (Hertel, Andreev, 2003), Ala (Spribille et al, 2020).

!C. armeniaca (DC.) Hafellner — on rocks; 5, a8 [R]; H. — Chu (Hertel, Andreev, 2003), Mag (Kotlov, 1995), Yak (Poryadina, 2005), Ala (Spribille et al, 2011).

Candelariella arctica (Körb.) R. Sant. — on rocks; 31 [R1]; H.

*С. аиге11а (НоАт.) гаЫЬг. — оп гоек8; 2 [^]; Н. — Кат (Himelbгant ег а1, 2014), СЬи (Лпагееу ег а1, 1996), Mag (Kotlov, 1995), Уак (/1пагееу ег а1, 1996), А1а (Geiseг ег а1, 1998).

С. еогаШга (Ку1.) Н. Magn. — оп гocks; а6 [И1].

*С. p1aеodizans (Ку1.) Н. Magn. — оп soi1 апа р1аП deЬгis; 1, 5, 10, 11, а2, а8 [И]; Н, БИ, ЬБСБ I. Уа^Ье^о). — Кат ^киНп, 1988), СЬи (Andгeev ег а1, 1996), Л1а (Thomson, 1997).

С. vite11ina (Hoffm.) Ми11. Лгg. — оп soi1 апа гocks; 1, 10, 14, 31, а6 [И]; Н, БИ, ЬБСБ.

С. xanthostigma (ЛсЬ.) Lettau — оп р1ап; deЬгis; 31 [И1]; Н.

!#СагЬопеа aggгegantu1a (Ми11. А^.) Diedeгich et TгieЬe1 — оп Ша11ш of Ьеоапота gangaleoides оп госк; 31 [И1]; LECБ. — СЬи ^Ьи^пко, 2009Ь), Уак ^Ьи^пко, 2009Ь), Л1а (SpгiЬi11e ег а1, 2020).

!С. voгtiеosa (Fl6гke) Heгte1 — оп гocks; 12, 31 [И]; Н, LECB. — СЬи (ШЛ^, Лndгeev, 2003), Mag (Койоу 1995), Уак (Лndгeev ег а1, 1996), Л1а (SpгiЬi11e ег а1, 2020).

!Catapyгenium еineгeum (Peгs.) К6гЬ. — оп soi1 oveг гocks; 5 [И1]; Н. — СЬи (Andгeev ег а1, 1996), Уак (Andгeev ег а1, 1996), Л1а (SpгiЬi11e ег а1, 2020).

СаШапа atгopurpuгea (ScЬaeг.) Vëzda et Poe1t — оп p1ant deЬгis and Ьагк of Уаоапшш и^тозит; 14, а6 [И]; Н.

!#Ceеidonia xenophana (К6гЬ.) TгieЬe1 et RamЬo1d — оп ШаШ of Porpidia тасгосагра and Р. гиЬегЫоза оп gгave1; 23, 26 [И]; Н (Р^ I: 5). — СЬи (Ш^Д Лndгeev, 2003), Л1а (SpгiЬi11e ег а1, 2020).

*Cetгaгia aеu1eata (ScЬгeЬ.) Бг. — оп soi1; 1, 10, 14, 23 [И]; Н. — Kam (Hime1bгant ег а1, 2014), СЬи (Лndгeev ега1, 1996), Уак (Лndгeev ега1, 1996), Л1а (SpгiЬi11e ег а1, 2020).

С. is1andiеa (L.) ЛcЬ. s. 1at. — оп soi1, гaгe1y оп Ьагк of Веги1а divaricaгa; 4, 7, 15, 20, 23, 26 [subsp. islandica £ sorediaгa (Schaeг.) ЛгпоЫ], 28-30, 32, а10, а12, а16 [О]; LECB (NesЬataeva, NesЬataev, 2018; NesЬataeva ег а1., 2018).

КС. kamеzatiеa Savicz — оп soi1; 4, 6, 7, 21-23, 30, 32 [О]; Н, БИ (Neshataeva, NesЬataev, 2018).

С. 1aevigata Rassad. — оп soi1; 1, 4-7, 11, 14, 15, 21-23, 26, 30, 32, а18 [Б]; Н (Neshataeva, NesЬataev, 2018; Neshataeva ег а1., 2018).

С. muricata (ЛcЬ.) Eckfe1dt — оп soi1; 1, 6, 12 [И]; LECB. (NesЬataeva, NesЬataev, 2018).

С. nigгiеans Ny1. — оп soi1 and гocks; 5, 6, 11, 12 [И] (Neshataeva, NesЬataev, 2018; Neshataeva ег а1., 2018).

С. sepinco1a (EЬгЬ.) ЛcЬ. — оп Ьагк of Alnus fruгicosa, Веги1а divaricaгa, В. егтапп, В. ехШз, Ртш ритНа, РогеМШа fruгicosa, БаИхрикИга, БогЬт затЬиа^Иа, Уаттит uliginosum; 8, 9, 13, 15, 20, 24-30, а9, а24 [О]; LECB.

*Cetгaгie11a еommixta (Ny1.) Л. TЬe11 et Kaгnefe1t — оп гocks; 31 [И1]; Н. — Kam ег al., 2014), СЬи (Andгeev ег al., 1996), Уак (Andгeev ег al., 1996), Л1а ^гМ^

ег al., 2020).

С. de1isei (Бoгy ex ScЬaeг.) Kaгnefe1t et Л. TЬe11 — оп soi1; 7, 17, 21-23, 26, 30, 32, а16 [О]; Н (Neshataeva, NesЬataev, 2018; NesЬataeva ег а1, 2018).

RCetгe1ia a1askana (Ш. и Си1Ь. et С. Б. Си1Ь.) Ш. L. Си1Ь. et С. Б. Си1Ь. — оп soi1; а8 [И1]; Н.

*Cheiгomyеina petгi D. Hawksw. et Poe1t — оп Ьагк of АЫш^игкоза; 27 [И1]; LECБ. — Kam (Hime1Ьгant ег а1, 2009), Л1а (SpгiЬi11e егal., 2020).

Cladonia amauгoегaea (Fl6гke) ScЬaeг. — оп soi1; 1, 4, 10, 14, 15, 22, 23, 32, а7, а8, а13 [О]; Н (NesЬataeva, NesЬataev, 2018).

Cladonia arbuscula (Wallr.) Flot. subsp. arbuscula (incl. subsp. beringiana Ahti) — on soil, rarely on bark of Betula divaricata and Pinus pumila; 1, 4-6, 10, 11, 13-15, 17, 20-23, 26, 29, 30, 32, a10, a12, a13 [F]; H (Neshataev et al., 2017; Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

C. bacilliformis (Nyl.) Sarnth. — on soil; 13 [R1]; LECB. C. bellidiflora (Ach.) Schaer. — on soil; 6, 13, 17 [R]; H.

C. borealis S. Stenroos — on soil; 1, 10-12, 14, 17, 26, 32, a12 [O] (Neshataeva, Neshataev, 2018).

*C. botrytes (K. G. Hagen) Willd. — on soil; a13 [R1]. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Poryadina, 2005), Ala (Spribille et al, 2010).

C. carneola (Fr.) Fr. — on soil; 14 [R1]. C. cenotea (Ach.) Schaer. — on soil; a3, a13 [R]; H.

*C. cervicornis (Ach.) Flot. — on soil; 10, 14 [R]; H (sub Arthrorhaphis aeruginosa). — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2010).

C. chlorophaea (Florke ex Sommerf.) Spreng. s. str. — on soil; 13, 28, 32 [R]. Thalli contain fumarprotocetraric acid (Neshataeva, Neshataev, 2018).

!C. cineracea Ahti — on soil; 17, 26 [R]; H. — Yak (Ahti, 2007). Earlier known from Chukotka, Khabarovsk, Trans-Baikal, and Krasnoyarsk territories, Buryatia, Alaska, and Northwest Territories, Canada (H, LE). The species is close to C. verticillata, but contains atranorin, is whitish (usually without any brown colouring), more slender and has many narrow, concave scyphi. In a phylogenetic tree (Stenroos et al, 2019) it is also very distinct. C. coccifera (L.) Willd. — on soil; 17 [R1]; H.

C. coniocraea (Florke) Spreng. — on bark of Alnus fruticosa, Betula divaricata, B. ermanii; 19, 27, 28 [R].

C. cornuta (L.) Hoffm. — [O]; LECB (Neshataeva, Neshataev, 2018).

subsp. cornuta — on soil and on bark of Alnus fruticosa; 4, 7, 15, 19, 26, 27, a12, a13.

subsp. groenlandica (E. Dahl) Ahti — on soil; 13, 22, a13.

C. crispata (Ach.) Flot. — [R]; H (Neshataeva, Neshataev, 2018).

var. crispata — on soil; 4, 7, 21, 22, 30, 32.

var. cetrariiformis (Delise) Vain. — on soil; 7.

C. cyanipes (Sommerf.) Nyl. — on soil, rarely on bark of Pinus pumila; 4, 8, 13, 15, a13 [R]; H.

C. deformis (L.) Hoffm. — on soil; 28, a13 [R]; H (Neshataeva, Neshataev, 2018). C. ecmocyna Leight. — on soil; a10 [R1]; H.

C. fimbriata (L.) Fr. — on soil, bark of Betula divaricata, B. ermanii, Potentilla fruticosa, Salix alaxensis, Sorbus sambucifolia; 18, 26-28 [R]; LECB. Thalli contain fumarprotocetraric acid.

C. gracilis (L.) Willd. — [O] (Neshataev et al, 2017; Neshataeva, Neshataev, 2018).

subsp. gracilis s. lat. — on soil; 14.

subsp. turbinata (Ach.) Ahti — on soil; 1, 32.

subsp. vulnerata Ahti — on soil; 4, 15, 21, 26, 30.

C. grayi G. Merr. ex Sandst. — on plant debris; a13 [R1]; LECB. Thallus contains grayanic acid.

C. kanewskii Oxner — on soil; 6, 7, 11, 17, 26, 30, a20 [R]; H (Neshataeva, Neshataev, 2018).

C. macilenta Hoffm. — on soil; 32, a13 [R] (Neshataeva, Neshataev, 2018).

C. macroceras (Delise) Hav. — on soil; 4, 6, 7, 10, 12, 14, 17, 21, a16 [O] (Neshataeva, Neshataev, 2018).

C. macrophylla (Schaer.) Stenh. — on soil; 32, a3 [R]; H (Neshataeva, Neshataev, 2018).

C. macrophyllodes Nyl. — on soil; 6, 17, 26 [R]; H (Neshataeva, Neshataev, 2018).

C. maxima (Asahina) Ahti — on soil; 7, 15, 17, 20-22, 26, 28, 30, 32 [O]; H (Neshataeva, Neshataev, 2018).

*C. mitis Sandst. — on soil; 7, 10-12, 17, 22, 32, a10 [O]; H. — Kam (Himelbrant et al., 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

C. nipponica Asahina — on soil; 7, 11, 21, a20 [R]; H (Neshataeva, Neshataev, 2018).

C. pleurota (Florke) Schaer. — on soil; 4, 6, 7, 15, 26, a13 [R]; H.

C. pyxidata (L.) Hoffm. — on soil; 1, 5, 6, 10, 14, 17, 26, 31, 32, a2, a8, a12 [O]; H (Neshataeva, Neshataev, 2018).

C. rangiferina (L.) F. H. Wigg. — on soil; 1, 4, 6, 7, 10, 11, 13-15, 20-23, 26, 29, 30, 32, a6, a11-a13 [F]; H (Neshataev et al, 2017; Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

C. rei Schaer. — on soil; a13 [R1].

C. scabriuscula (Delise) Nyl. — on soil; a5 [R1]; H.

C. squamosa Hoffm. — on soil; 4-6, 11, 21, 26, a3, a8, a13 [R]; H (Neshataeva, Neshataev, 2018).

C. stellaris (Opiz) Pouzar et Vezda — on soil; 4, 7, 11, 13, 15, 21-23, 29, 30, 32 [O]; H (Neshataev et al, 2017; Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

C. straminea (Sommerf.) Florke — on soil; 4, 6, 15 [R].

C. stygia (Fr.) Ruoss — on soil; 4, 7, 11, 15, 20-23, 30, a10, a12 [O]; H (Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

C. subfurcata (Nyl.) Arnold — on soil; 7, 30, 32 [R]; H (Neshataeva, Neshataev, 2018).

C. sulphurina (Michx.) Fr. — on soil; a12 [R1]; H.

*C. trassii Ahti — on soil; 10, 11, 17, 26, a10 [R]; H. — Kam (Himelbrant et al, 2014), Chu (Makarova, Katenin, 2009), Ala (Spribille et al, 2020).

C. turgida Hoffm. — on soil; 17 [R1]; H.

C. uliginosa (Ahti) Ahti — on soil; 7, 10, 26 [R]; H (Neshataeva, Neshataev, 2018).

C. uncialis (L.) F. H. Wigg. — on soil; 4-7, 10, 11, 14, 15, 17, 21-23, 26, 29, 30, 32, a8, a11, a18 [F]; H (Neshataev et al, 2017; Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

C. wainioi Savicz — on soil; 4, 15, a6, a18 [R]; H.

!Clauzadeana macula (Taylor) Coppins et Rambold — on rocks; 5 [R1]; H (sub Calvitimela armeniaca) (Plate I: 4). — Chu (Hertel, Andreev, 2003), Mag (Kotlov, 1995), Yak (Hertel, Andreev, 2003), Ala (Talbot et al, 1997).

!Collemopsidium foveolatum (A. L. Sm.) F. Mohr — on barnacles over rocks in the sublittoral zone; a1 [R1]; H. — Ala (Dillman et al, 2012). New to the Russian Far East and Asian Russia. Differs from related species by having completely immersed thallus and relatively poorly developed involucrellum which is either lid-like or absent (Mohr et al, 2004).

Dibaeis baeomyces (L. f.) Rambold et Hertel — on soil; 4, 7, 11, 17 [R] (Neshataeva, Neshataev, 2018).

*Epilichen scabrosus (Ach.) Clem. — lichenicolous on thallus of Baeomyces sp. on soil over rocks; 1 [R1]; LECB. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Poryadina, 2020), Ala (Spribille et al, 2020).

Euopsis granatina (Sommerf.) Nyl. — on rocks; a8 [R1]; H.

Flavocetraria cucullata (Bellardi) Karnefelt et Thell — on soil; 1, 4-6, 10, 11, 14, 15, 22, 23 [O]; FR (Neshataev et al., 2017; Neshataeva, Neshataev, 2018).

F. minuscula (Elenkin et Savicz) Ahti et al. — on soil; 4, 14, 21 [R] (Neshataeva, Neshataev, 2018).

F. nivalis (L.) Karnefelt et Thell — on soil, rarely on bark of Pinus pumila; 1, 4, 6, 10, 11, 13, 15, 21, 23 [O] (Neshataev et al, 2017; Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

IFlavoplaca flavocitrina (Nyl.) Arup et al. — on concrete of lighthouse; fertile specimen; 3 [R1]; LECB (det. Frolov). — Ala (Spribille et al, 2020). New to the Russian Far East. The nearest known localities in Russia belong to Republic of Altai and Trans-Baikal Territory (Vondrak et al, 2016). Similar to F. citrina (Hoffm.) Arup et al., from which differs by the flat areoles, farinose soredia which are bright yellow and paler than the cortex, and smooth margins of apothecia (The lichens..., 2009). Our specimen is fertile.

Frutidella furfuracea (Anzi) M. Westb. et M. Svensson — on bark of Alnus fruticosa, Betula divaricata, B. ermanii, Pinus pumila, Potentilla fruticosa, Sorbus sambucifolia, on wood of dwarf shrubs; 8, 9, 13, 16, 19, 26-29, a6 [O]; FR, LECB. Thalli contain sphaerophorin.

IFuscidea circumflexa (Nyl.) V. Wirth et Vezda — on rocks; 5, 10, a2, a12 [R]; LECB (Plate I: 3). — Chu (Andreev et al, 1996). Thalli contain alectorialic acid. Rarely reported species, probably endemic of Beringia.

Fuscopannaria praetermissa (Nyl.) P. M. Jorg. — on soil; a8 [R1]; H (sub Cladonia pyxidata).

!#Geltingia associata (Th. Fr.) Alstrup et D. Hawksw. — on thalli of Ochrolechia spp. on rocks and mosses; 10, a13 [R]; H, LECB. — Ala (Spribille et al, 2020). New to the Russian Far East. The nearest reported localities are in Krasnoyarsk Territory (Zhurbenko, 2007). Highly visible fungus with black apothecia and broadly ellipsoid spores; for detailed description see Diederich et al. (2010).

Gowardia nigricans (Ach.) Halonen et al. — on soil, rocks, bark of Pinus pumila; 1, 4-7, 10-15, 21-23, 30, a6, a13 [F]; H (Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

*Helocarpon crassipes Th. Fr. — on plant debris; 5 [R1]; LECB. — Kam (Mikulin, 1988), Ala (Spribille et al, 2020).

IHydropunctaria maura (Wahlenb.) Keller et al. s. lat. — on seashore rocks in littoral zone; 2 [R1]; H. — Chu (Andreev et al, 1996), Ala (Spribille et al, 2020).

IHypogymnia beringiana (Krog) McCune — on soil over rocks; 5, a8 [R]; H, FR, LECB. — Yak (McCune, 2008), Ala (McCune, 2008).

IH. fistulosa McCune et Krog — on soil; 5, a6-a8 [R]; H, FR, LECB. — Chu (Zhurbenko, 2009a), Ala (McCune, 2008).

H. pseudophysodes (Asahina) Rassad. — on bark of Betula divaricata and Pinus pumila; 8, 28, 29 [R]; LECB.

H. subobscura (Vain.) Poelt — on soil over rocks; 1, 5 [R]; LECB.

Immersaria athroocarpa (Ach.) Rambold et Pietschm. — on gravel; 11, 12 [R]; LECB.

Ilonaspis ceracea (Arnold) Jatta — on pebbles; 23 [R1]; H (sub Rhizocarpon infernulum and R. sublavatum). — New to Beringia and Russian Far East. The nearest reported localities in Russia belong to Republic of Altai and Kemerovo Region (Sedelnikova, 2013). Characterized by thin brownish-gray thallus with trebouxioid photobiont, and concave (first immersed) pale-brownish apothecia without dark pigments (Foucard, 2001).

I. obtecta (Vain.) R. Sant. — on gravel; 11, 17 [R]; LECB.

II. odora (Ach.) Th. Fr. — on gravel; a3 [R1]; LECB. — Chu (Andreev et al, 1996), Ala (Spribille et al, 2020).

Japewia tornoensis (Nyl.) Tensberg — on soil and plant debris, also on thallus of Umbilicaria torrefacta on rock, bark of Pinuspumila, Potentilla fruticosa, Vaccinium uliginosum, wood of Alnus fruticosa, Pinus pumila, and Salix sp.; 4-6, 8, 10, 11, 13, 15, 21, 23, 24, 32, a7, a9 [O]; FR, LECB.

!Lambiella furvella (Nyl. ex Mudd) M. Westb. et Resl — lichenicolous on thalli of crustose lichens (Lecanora sp. and Rhizocarpon sp.) on rock; 31 [R1]; H. — Ala (Spribille et al., 2010). New to the Russian Far East. The nearest known locality is in Krasnoyarsk Territory (Zhdanov, 2010). The lichen grows over crustose saxicolous species and forms thick brown isidiate thalli resembling Placynthiella icmalea (The lichens... , 2009).

L. insularis (Nyl.) T. Sprib. — lichenicolous on thalli of Lecanora sulphurea and L. rupicola; a6 [R1]; H.

*Lecanora aitema (Ach.) Hepp — on bark of Alnus fruticosa and driftwood; 4, 9, 19, a14, a15 [R]; LECB. — Kam (Himelbrant et al,, 2014).

L. albellula (Nyl.) Th. Fr. — on bark of Pinus pumila; 8 [R1].

*L. atrosulphurea (Wahlenb.) Ach. — on rocks; 5, 6, 10, a7 [R]; H, LECB. — Kam (Himelbrant et al., 2014), Chu (Andreev et al., 1996), Yak (Andreev et al., 1996), Ala (Thomson, 1997).

L. boligera (Norman ex Th. Fr.) Hedl. — on bark of Alnus fruticosa, Betula divaricata, B. ermanii, B. exilis, Pinus pumila, Potentilla fruticosa, Rhododendron aureum, Salix pulchra, Sorbus sambucifolia, on wood; 8, 9, 13, 16, 19, 20, 24-29, a6, a9, a15, a24 [O]; LECB.

L. cadubriae (A. Massal.) Hedl. — on bark of Betula ermanii, Pinus pumila, Vaccinium uliginosum; 8, 15, 27, a9 [R]; LECB (sub Tetramelas chloroleucus).

L. chlarotera Nyl. — on bark of Alnus fruticosa, Betula ermanii, Pinus pumila, Salix alaxensis, Vaccinium uliginosum; 8, 9, 15, 16, 18, 19, 27, a8 [O]; H, LECB.

L. circumborealis Brodo et Vitik. — on bark of Alnus fruticosa; 9, 15, 16 [R]; LECB.

!L. epibryon (Ach.) Ach. — on plant debris; 1, 5, 10, 14, 17, a8 [R]; H, FR, LECB. — Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2010).

L. fuscescens (Sommerf.) Nyl. — on bark of Alnus fruticosa, Betula divaricata, B. ermanii, Pinus pumila, Potentilla fruticosa, Sorbus sambucifolia; 8, 9, 13, 16, 19, 24, 27-29 [O]; LECB.

!L. gangaleoides Nyl. — on seashore rocks; 5, 31 [R]; H. — Chu (Andreev et al, 1996).

L. intricata (Ach.) Ach. — on rocks and gravel; 1, 5, 10-12, 31 [R]; LECB (sub Carbonea vorticosa).

!L. orae-frigidae R. Sant. — on driftwood; a15 [R1]; LECB. — Yak (Poryadina, 2005), Ala (Spribille et al, 2020).

L. polytropa (Ehrh. ex Hoffm.) Rabenh. — on rocks and gravel; 1-3, 5, 6, 10-12, 14, 23, 31, a3, a6, a7 [O]; H, FR, LECB.

*L. pulicaris (Pers.) Ach. — on bark of Sorbus sambucifolia; 27 [R1]; LECB. — Kam (Mikulin, 1987), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

L. rupicola (L.) Zahlbr. — on rocks; 1, 5, 10, a2, a6 [R]; LECB (sub Arthonia varians).

!L. sulphurea (Hoffm.) Ach. — on rocks; 1, 5, 10, a2, a6, a7 [R]; H, LECB. — Ala (Hertel, Andreev, 2003). New to the northern part of the Russian Far East; known from Khabarovsk Territory (Yakovchenko et al, 2013).

L. symmicta (Ach.) Ach. — on plant debris, bark of Alnus fruticosa, Betula ermanii, B. exilis, Pinus pumila, Potentilla fruticosa, Salix alaxensis, Sorbus sambucifolia, Vaccinium uliginosum, on wood; 6-9, 11-16, 18, 21, 23, 24, 27, a7-a9, a15, a24 [F]; LECB.

Plate I. 1a, 1b — Allantoparmelia sibirica (H, Kor-Goven-08ad); 2 — Anema jenisejense (H, Kor-Goven-01); 3 — Fuscidea circumflexa (LECB, Kor-Goven-02ad); 4 — Clauzadeana macula (H, Kor-Goven-05); 5 — Cecidoniaxenophana (H, Kor-Galin-26). Scale bars: 1a, 1b, 3, 4 — 1 mm, 2, 5 — 0.5 mm.

Lecidea auriculata Th. Fr. subsp. auriculata — on rocks; 5, 10, 31 [R]; H. *L. confluens (Weber) Ach. — on gravel; 12 [RJ; LECB. — Kam (Mikulin, 1988), Chu (Andreev et al., 1996), Mag (Kotlov, 1995), Ala (Talbot et al., 1997).

*L. epiphaea Nyl. — on mosses; 5, a8 [R]; H. — Kam (Neshataeva et al, 2006), Chu (Andreev et al, 1996), Yak (Zhurbenko, 2009b), Ala (Spribille et al, 2010).

Plate II. 1 — Lecidea subpaupercula (H, Kor-Chayachy-31); 2 — Micarea inquinans (H, Kor-Goven-11); 3 — Rinodina calcigena (H, Kor-Goven-03); 4 — Santessoniella arctophila (H, Galin-16ad). Scale bars: 1 — 1 mm, 2-4 — 0.5 mm.

L. erythrophaea Flörke ex Sommerf. — on bark of Alnus fruticosa and Salix alaxensis; 18, 19, 27 [R]; LECB.

*L. inops Th. Fr. — on gravel; 12 [R1]; LECB. — Kam (Nylander, 1887).

*L. lapicida (Ach.) Ach. — [R]; H, FR, LECB. — Kam (Himelbrant et al., 2014), Chu (Hertel, Andreev, 2003), Mag (Kotlov, 1995), Yak (Hertel, Andreev, 2003), Ala (Spribille et al., 2020).

var. lapicida — on rocks; 1, 5, 6, 10, 31, a2, a6, a7.

var. pantherina Ach. — on rocks; 1, 5, 6, 10, 12.

*L. praenubila Nyl. — on rocks; 31 [R1]; LECB (sub Sagedia zonata). — Kam (Nylander, 1887), Chu (Andreev et al., 1996), Ala (Spribille et al., 2020).

!L. promiscens Nyl. — on rocks; 6 [R1]; LECB. — Chu (Hertel, Andreev, 2003), Ala (Hertel, Andreev, 2003). 1

!L. subpaupercula Mas. Inoue (syn. L. fuscoatrata Nyl., L. fuscatoatra Nyl.) — on rocks; 31 [R1]; H, FR, LECB. — New to Beringia and Russian Far East; in Russia known from Kola

Peninsula, also reported from Japan and North America. The species has thin brownish thallus of atrobrunnea-type, and dark brown hypothecium (Plate II: 1). It is similar to L. paupercula Th. Fr., but has lower hymenium, smaller spores, and medulla and exciple P- (Andreev, 1998).

*Lecidea swarzioidea Nyl. — on rocks; 5, 10 [R]; H, LECB. — Kam (Hertel, Andreev, 2003), Ala (Spribille et al., 2020).

ILecidella asema (Nyl.) Knoph et Hertel — on rocks; 31 [R1]; H (sub Rusavskia sorediata). — Ala (McCune et al, 2018). New to the Russian Far East. Known in Russia from Irkutsk and Leningrad regions (Himelbrant et al, 2019). Characterized by granular to verrucose and rimose thick yellowish thallus with thiophanic acid in cortex (C+ orange) and blue-green epithecium (The lichens... , 2009).

IL. carpathica Körb. — on rocks; 31 [R1]; H (sub Lecidea subpaupercula). — Chu (Andreev et al, 1996), Ala (Spribille et al, 2020).

IL. effugiens (Nilson) Knoph et Hertel — on rocks; 1, 5 [R]; H, LECB. — Chu (Andreev et al, 1996), Ala (Spribille et al, 2010).

L. euphorea (Flörke) Hertel — on plant debris, bark of Alnus fruticosa, wood; 5, 9, a13, a15 [R]; LECB.

IL. scabra (Taylor) Hertel et Leuckert — on rocks; 2 [R1]; H. — Ala (McCune et al, 2018). New to the Russian Far East. Probably widespread, but overlooked species; in Russian Arctic known from Dolgy Island in Arkhangelsk Region, also cited from Novosibirskie islands, Krasnoyarsk Territory (Urbanavichus et al, 2009). Characterized by rimose to granular thallus with C+ red-orange cortex, and fine soredia formed in roundish, more or less discrete soralia (McCune, 2017).

IL. stigmatea (Ach.) Hertel et Leuckert — on concrete of lighthouse; 3 [R1]; LECB. — Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

IL. wulfenii (Hepp) Körb. — on plant debris, turf, and soil over rocks; 1, 10, 14, 31, a2, a8 [R]; H, FR. — Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

ILecidoma demissum (Rutstr.) Gotth. Schneid. et Hertel — on soil; 11 [R1]; LECB. — Chu (Andreev et al, 1996), Ala (Spribille et al, 2020).

Lepra borealis (Erichsen) I. Schmitt et al. — on bark of Alnus fruticosa, Betula divaricata, B. ermanii, Pinus pumila, Potentilla fruticosa, Rhododendron aureum, Sorbus sambucifolia; 8, 13, 19, 20, 24, 27, 28 [O]; LECB. Thalli contain protocetraric, fumarprotocetraric, and succinprotocetraric acids.

L. dactylina (Ach.) Hafellner — on soil and plant debris; 1, 7, 17 [R]; H.

L. panyrga (Ach.) Hafellner — on soil and plant debris; 4, 6, 10, 11, 15, 21, a8 [R] (Neshataeva, Neshataev, 2018).

ILepraria borealis Lohtander et Tonsberg — on soil; 31 [R1]; LECB. Thallus contains atranorin and jackinic/rangiformic acid. — New to Beringia and Russian Far East. The nearest reported localities are in the Republic of Buryatia (Kharpukhaeva, Khanin, 2012) and Taimyr Peninsula, Krasnoyarsk Territory (Kristinsson et al, 2010). The member of L. neglecta group (see Saag et al, 2009), accounted also as a chemotype, together with L. caesioalba and L. gelida (Lendemer, 2013).

IL. caesioalba (de Lesd.) J. R. Laundon — on mosses; 17 [R1]; LECB. Thallus contains atranorin, jackinic/rangiformic, and stictic acids. — Ala (Spribille et al., 2010). New to the Russian Far East. The nearest known localities are in Taimyr Peninsula and Central Siberian Reserve in Krasnoyarsk Territory (Kristinsson et al, 2010; Zhdanov, 2013).

*L. elobata Tensberg — on bark of Alnus fruticosa, plant debris and mosses; 5, 17, 19, 26 [R]; LECB. — Kam (Neshataeva et al., 2004), Ala (McCune et al., 2018).

IL. gelida Tensberg et Zhurb. — on mosses; 2 [R1]; LECB. Thallus contains porphyrilic, alectorialic acids, and unidentified fatty acid. — Chu (Kristinsson et al., 2010).

L. neglecta (Nyl.) Erichsen — on soil; 1, 5 [R]; LECB. Thalli contain alectorialic acid.

Leptogium saturninum (Dicks.) Nyl. — on plant debris; 5 [R1].

I#Lichenodiplis lecanorae (Vouaux) Dyko et D. Hawksw. — on apothecia of Caloplaca borealis on twigs of Vaccinium uliginosum; 14 [R1]; H. — Kam (Zhurbenko et al, 2012).

*Lichenomphalia alpina (Britzelm.) Redhead et al. — on soil over rocks; a8 [R1]. — Kam (Himelbrant et al, 2008), Ala (Geml et al, 2012).

*RL. hudsoniana (H. S. Jenn.) Redhead et al. — on soil over rocks; 5 [R1]; LECB. — Kam (Trass, 1963), Yak (Poryadina, 2005), Ala (Spribille et al, 2010).

L. umbellifera (L.) Redhead et al. — on soil; 3 [R1].

Lobaria linita (Ach.) Rabenh. — on soil; 5, 14, 17, 23, 31, a16, a17 [R]; H (Neshataeva, Neshataev, 2018).

L. scrobiculata (Scop.) DC. — on soil over rocks; a9 [R1]; LECB.

Lopadium coralloideum (Nyl.) Lynge — on plant debris; 31, a8 [R]; H, FR.

L. pezizoideum (Ach.) Körb. — on soil and plant debris; 11, 26, a16 [R]; H, LECB.

Melanelia hepatizon (Ach.) Thell — on soil over rocks and boulders; 1, 5, 6, 10, 12, 23, a6, a8 [R]; H, FR, LECB.

M. stygia (L.) Essl. — on rocks and gravel; 1, 5, 6, 11, 12, 23, a2, a3 [R]; H, FR, LECB.

Melanohalea infumata (Nyl.) O. Blanco et al. — on rocks; 1, 10, a2 [R]; H.

M. olivacea (L.) O. Blanco et al. — on bark of Alnus fruticosa, Betula ermanii, Salixalaxensis, Vaccinium uliginosum; 9, 15, 18, 27, a8 [R]; H.

M. septentrionalis (Lynge) O. Blanco et al. — on bark of Alnus fruticosa and Betula ermanii; 27, a8 [R]; LECB.

IMicarea incrassata Hedl. — on plant debris; 26 [R1]; LECB. — Chu (Andreev et al, 1996), Yak (Zhurbenko et al, 2005), Ala (Thomson, 1997).

I#M. inquinans (Tul.) Coppins — on thallus of Dibaeis baeomyces on disturbed soil; 11 [R1]; H (conf. L. A. Konoreva, S. V. Chesnokov). — Yak (Zhurbenko, 2020), Ala (Fryday, 2017). New to the Russian Far East. The species forms aggregated black apothecia on thalli of Dibaeis and Baeomyces (Plate II: 2); for description and world distribution see Zhurbenko (2020).

M. prasina Fr. — on plant debris; 26 [R1]; LE (det. Chesnokov). Thallus contains micareic acid.

IMiriquidica griseoatra (Flot.) Hertel et Rambold — on rocks; 5 [R1]; H. — Chu (Andreev et al, 1996).

IM. lulensis (Hellb.) Hertel et Rambold — on rocks; 1, a2 [R]; LECB. — Chu (Andreev et al, 1996), Ala (Hertel, Andreev, 2003).

IM. pulvinatula (Arnold) Hertel et Rambold — on gravel; 11 [R1]; LECB (sub Immersaria athroocarpa). — Ala (Hertel, Andreev, 2003). New to Russia. The nearest reported locality is Mystery Creek in Alaska. The species is close to more common M. lulensis, from which it differs in containing stictic acid (K+ yellow, not forming crystals) instead of norstictic acid (Hertel, Andreev, 2003).

Montanelia disjuncta (Erichsen) Divakar et al. — on rocks; 1, 5, a6 [R]; LECB.

M. panniformis (Nyl.) Divakar et al. — on rocks; a8 [R1]; H.

I#Muellerella pygmaea (Körb.) D. Hawksw. — on thallus of Lecidea subpaupercula on rock; 31 [R1]; H. — Chu (Zhurbenko, 2009b), Ala (Spribille et al, 2020).

Mycobilimbia carneoalbida (Müll. Arg.) S. Ekman et Printzen — on soil; 26 [RJ; LECB.

M. epixanthoides (Nyl.) Vitik. et al. ex Hafellner et Türk. — on mosses; 17 [RJ; LECB.

*Mycoblastus affinis (Schaer.) T. Schauer — on soil, plant debris, and wood of dwarf shrubs; 1, 5, 6, 11, 12, a8 [R]; H, FR. — Kam (Himelbrant et al, 2014), Chu (Andreev et al., 1996), Ala (Spribille et al., 2020).

M. sanguinarius (L.) Norman — on plant debris; 11 [RJ; H.

!Myriolecis andrewii (B. de Lesd.) Sliwa et al. — on seashore rock near seabird colony; 2 [RJ; H. — New to Beringia and Russia. Known from Europe and North America. Coastal species with thin thallus and more or less dispersed apothecia with well-developed margin and disc K+ yellow, C+ orange, Pd+ orange (see Sliwa, 2007; Bertrand et al, 2018).

!M. semipallida (H. Magn.) Sliwa et al. — on concrete of lighthouse; 3 [RJ; LECB. — Ala (Sliwa, 2007). New to the Russian Far East. In Asian Russia known from Altai Territory (Davydov, 2001). This is a common but overlooked species of the M. dispersa group inhabiting concrete structures and characterized by UV+ orange-yellow apothecia with epihymenial granules soluble in K (Sliwa, 2007).

M. straminea (Ach.) Sliwa et al. — on seashore and mountain rocks manured by birds; 1, 2, 10, 31, a6 [R]; H, FR, LECB.

!M. torrida (Vain.) Sliwa et al. — on concrete of lighthouse; 3 [RJ; LECB. — Chu (Andreev et al, 1996), Ala (Spribille et al, 2020).

*M. zosterae (Ach.) Sliwa et al. — [R]; H, LECB. — Kam (Nylander, 1887), Chu (Andreev et al, 1996), Ala (Spribille et al, 2010).

var. zosterae — on plant debris; 1, 10.

var. behringii (Nyl.) Sliwa — on seashore rocks near seabird colony; 2.

var. palanderi (Vain.) Sliwa — on plant debris; 10.

*Myriospora smaragdula (Wahlenb. ex Ach.) Nägeli ex Uloth — on rocks; 1 [RJ; H. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Poryadina, 2005), Ala (Thomson, 1997).

*Naetrocymbe punctiformis (Pers.) R. C. Harris — on bark of Pinus pumila and Sorbus sambucifolia; 8, 27, 29 [R]; LECB. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Ala (McCune et al, 2009).

Nephroma arcticum (L.) Torss. — on soil; 21, 29, 30, a10 [R]; LECB (Neshataeva, Neshataev, 2018).

N. bellum (Spreng.) Tuck. — on bark of Salix alaxensis; 18 [RJ; LECB.

!Ochrolechia alaskana (Verseghy) Kukwa — on plant debris and rocks; a2 [RJ; LECB. Thalli contain lecanoric, gyrophoric, and variolaric acids, and two "microstictoides-unknowns". — Ala (Kukwa, 2009). New to Russia. The species resembles a morph of O. frigida, but unlike that species O. alaskana has variolaric acid (C+ yellow) in lower part of the cortex additionally to gyrophoric acid which is C+ red, and also presence of "microstictoides-unknowns" detected by TLC (Kukwa, 2009, 2011).

O. frigida (Sw.) Lynge s. lat. [incl. O. gonatodes (Ach.) Räsänen, O. inaequatula (Nyl.) Zahlbr.] — on plant debris, soil, mosses, bark of Betula divaricata, Pinus pumila, on wood; 1, 4-8, 10-15, 21, 23, 26, 28, 29, 31, 32, a2, a3, a7, a8, a12, a13, a24 [F]; H, FR, LECB (Neshataeva, Neshataev, 2018). Thalli contain lecanoric and gyrophoric acids.

O. mahluensis Räsänen — on bark of Alnus fruticosa, Betula divaricata, Pinus pumila; 8, 19, 28, 29 [R]; LECB. Thalli contain lecanoric and gyrophoric acids.

O. subplicans (Nyl.) Brodo subsp. subplicans — on rocks; 5, 6, 10 [R]; FR, LECB.

O. upsaliensis (L.) A. Massal. — on soil and plant debris; 1, 10, 11, 14 [R]; H, FR (Neshataeva, Neshataev, 2018). Thalli contain variolaric acid.

*Ophioparma ventosa (L.) Norman — on rocks; 1, 5, 10 [R]; FR, LECB. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al., 2010).

*Orphniospora moriopsis (A. Massal.) D. Hawksw. — on rocks; a8 [R1]; H. — Kam (Mikulin, 1987), Chu (Andreev et al., 1996), Mag (Kotlov, 1995), Yak (Andreev et al., 1996), Ala (Spribille et al., 2010).

Parmelia omphalodes (L.) Ach. subsp. omphalodes — on soil and rocks; 1, 4-6, 10-12, 14, a6-a8 [O]; H, FR (Neshataeva, Neshataev, 2018).

P. saxatilis (L.) Ach. — on soil, rocks, and gravel; 1, 2, 5, 10, 14, a2, a6, a7 [R]; H.

P. skultii Hale — on soil, rocks, and gravel; 1, 3, 5, 10, 14, a2, a6, a8 [R]; H, FR, LECB.

P. sulcata Taylor — on soil and rocks; 10, a2 [R]; LECB.

Parmeliopsis ambigua (Wulfen) Nyl. — on bark of Alnus fruticosa, Betula divaricata, B. ermanii, B. exilis, Pinus pumila, Potentilla fruticosa, Salix pulchra, Sorbus sambucifolia, Vaccinium uliginosum, on wood; 8, 9, 13, 15, 19, 20, 24-29, 32, a6, a9, a15, a24 [F]; H, LECB.

P. hyperopta (Ach.) Arnold — on bark of Alnus fruticosa, Betula divaricata, B. ermanii, B. exilis, Pinus pumila, Salix alaxensis, S. pulchra, Sorbus sambucifolia, Vaccinium uliginosum, on wood; 8, 9, 13, 15, 16, 18-20, 24-29, 32, a6, a9, a15, a24 [F]; H, LECB.

Peltigera aphthosa (L.) Willd. — on soil; a10-a12 [R]; H (Neshataeva, Neshataev, 2018).

P. canina (L.) Willd. — on soil; a17 [R1]; H.

P. collina (Ach.) Schrad. — on bark of Salix alaxensis; 18 [R1]; LECB.

P. didactyla (With.) J. R. Laundon — on soil and bark of Salix alaxensis; 10, 17, 18, 23, 31, a16 [R]; LECB (Neshataeva, Neshataev, 2018).

*P. elisabethae Gyeln. — on mosses; 22 [R1]; LECB. — Kam (Himelbrant et al, 2014), Yak (Poryadina, 2005), Ala (Geiser et al, 1998).

P. extenuata (Norrl. ex Vain.) Lojka — on soil; a12 [R1]; H.

P. latiloba Holt.-Hartw. — on soil; 23 [R1]; LECB.

P. leucophlebia (Nyl.) Gyeln. — on soil; 26, a10, a17 [R]; H, LECB.

P. malacea (Ach.) Funck — on soil; 21, 23, a9, a11 [R]; H, LECB (Neshataeva, Neshataev, 2018).

P. neopolydactyla (Gyeln.) Gyeln. — on soil; 17 [R1]; H.

P. polydactylon (Neck.) Hoffm. — on soil; a10 [R1]; LECB.

P. rufescens (Weiss) Humb. — on soil; 6, 10, 17, 31, a9 [R].

P. scabrosa Th. Fr. — on soil; 21, 23, 30, a3, a10 [R]; H (Neshataeva, Neshataev, 2018).

*Pertusaria bryontha (Ach.) Nyl. — on plant debris; 1, 11, a2 [R]; H, LECB (sub Taeniolella pertusariicola and Sphinctrina turbinata). — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

P. carneopallida (Nyl.) Anzi — on bark of Alnus fruticosa, Betula divaricata, B. ermanii, B. exilis, Pinus pumila, Sorbus sambucifolia; 9, 16, 19, 26-29, a24 [O]; LECB.

P. coriacea (Th. Fr.) Th. Fr. — on soil, plant debris, and wood of dwarf shrubs; 1, 5, 6, 10-12, 14, a2, a8 [O]; H, FR, LECB (Neshataeva, Neshataev, 2018).

*P. cribellata Branth — on rocks; 5, 10, a7 [R]; LECB. — Kam (Himelbrant et al, 2014).

*P. glomerata (Ach.) Schaer. — on rocks and soil over rocks; 5, 10, a2, a8 [R1]; LECB. Thalli contain norstictic acid (major), traces of cf. salazinic acid in specimens 5, 10, a2, and stictic acid

complex in specimen a8. — Kam (Nylander, 1887), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al., 1996), Ala (Thomson, 1997).

*Pertusaria multipuncta (Turner) Nyl. — on bark of Pinuspumila; 8 [R1]; LECB. Thallus contains fumarprotocetraric acid. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Ala (Thomson, Ahti, 1994).

*P. octomela (Norman) Erichsen — on plant debris; 5 [R1]; LECB. — Kam (Himelbrant et al, 2014), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Thomson, 1997).

P. oculata (Dicks.) Th. Fr. — on soil and plant debris; 4, 5, 7, 10, 21, 26 [R]; FR, LECB (Neshataeva, Neshataev, 2018).

!P. pertusa (Weigel) Tuck. var. pertusa — on rock; a8 [R1]; H. — Chu (Andreev et al, 1996), Yak (Poryadina, 2005). 1

+Phaeocalicium compressulum (Szatala) A. F. W. Schmidt — on bark of Alnus fruticosa; 9, 15, 16, 19, 27 [R]; LECB.

*+P. flabelliforme Tibell — on bark of Betula divaricata; 28, 29 [R]; LECB. — Kam (Neshataeva et al., 2005).

*Phaeophyscia constipata (Norrl. et Nyl.) Moberg — on soil over rocks, plant debris, and mosses; 1, 31 [R]; FR, LECB. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Thomson, Ahti, 1994).

*P. sciastra (Ach.) Moberg — on rocks, soil over rocks, and concrete of lighthouse; 2, 3, 31, a6 [R]; H, FR. — Kam (Kuznetsova, Himelbrant, 2006), Chu (Andreev et al, 1996), Mag (Zhelydeva, 2015), Yak (Poryadina, 2005), Ala (Thomson, Ahti, 1994).

Physcia caesia (Hoffm.) Fürnr. — on rocks and on concrete of lighthouse; 1-3, 31, a6 [R]; H.

P. dubia (Hoffm.) Lettau — on soil and rocks; 1, 2, 10, 31, a6, a7 [R]; H.

Physconia grumosa Kashiw. et Poelt — on soil over rocks, rocks, plant debris, and mosses; 1, 2, 10, a6 [R]; H, FR, LECB.

*P. muscigena (Ach.) Poelt — on soil and mosses; 1, 10, 14, 31, a6 [R]; H, FR. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Thomson, Ahti, 1994).

!Placidiopsis pseudocinerea Breuss — on soil over rocks; 2 [R1]; H. — Yak (Poryadina, 2005), Ala (Breuss, McCune, 1994). — New to the Russian Far East. The nearest known locality is Taimyr Peninsula in Krasnoyarsk Territory (Breuss, 1996). Morphologically similar to Catapyrenium cinereum which has narrower simple spores, whereas the spores of Placidiopsis pseudocinerea are one-septate. Differs from other species of the genus by absence of rhizines, presence of dark rhizohyphae instead, and ellipsoid spores 5.5-9.0 ^m wide (Breuss, 2010; McCune, 2017).

Placynthiella dasaea (Stirt.) Tensberg — on plant debris and mosses; 13, 22 [R]; LECB.

P. icmalea (Ach.) Coppins et P. James — on soil and plant debris; 1, 10 [R].

!P. oligotropha (J. R. Laundon) Coppins et P. James — on soil and plant debris; 10, 13, 14 [R]; H (sub Scytinium imbricatum). — Yak (Chesnokov et al, 2016), Ala (Spribille et al, 2010). New to the northern part of the Russian Far East; known from Sakhalin Region (Konoreva et al., 2020).

P. uliginosa (Schrad.) Coppins et P. James — on soil and plant debris; 7, 13, a3 [R]; LECB.

Polycauliona candelaria (L.) Frodén et al. — on soil over rocks, on rocks and driftwood; 1, 10, a2, a6, a7, a23 [R]; LECB.

*P. verruculifera (Vain.) Arup et al. — on seashore rocks; 2, 31 [R]; LECB. — Kam (Nylander, 1887), Ala (Talbot et al, 1997).

IPolychidium muscicola (Sw.) Gray — on soil; a4 [R1]; LECB. — Chu (Andreev et al, 1996), Yak (Poryadina, 2005), Ala (Spribille et al., 2020).

Polysporina simplex (Taylor) Vezda — on gravel; 12 [R1]; LECB (sub Rhizocarpon intersitum).

Porpidia crustulata (Ach.) Hertel et Knoph — on rocks; 1, 17, a6 [R]; H, FR, LECB.

P. flavicunda (Ach.) Gowan — on rocks; 5 [R1]; FR.

P. macrocarpa (DC.) Hertel et A. J. Schwab — on rocks; 5, 23, 26 [R]; H.

P. melinodes (Körb.) Gowan et Ahti — on rocks; 1, a6 [R]; H.

IP. soredizodes (Lamy ex Nyl.) J. R. Laundon — on rocks; 17, a3 [R]; LECB. — Ala (Spribille et al, 2020). New to the Russian Far East. The nearest reported locality is Kodar Range in Trans-Baikal Territory (Makryi, 2002). Crustose lichen with thin grayish thallus and roundish soralia K + yellow and Pd+ orange due to presence of stictic acid (The lichens..., 2009).

IP. speirea (Ach.) Kremp. — on rocks; 2 [R1]; H. — Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

IP. thomsonii Gowan — on gravel; 11, 12 [R]; LECB. — Yak (Andreev et al, 1996), Ala (Spribille et al, 2020). New to the Russian Far East. The species is difficult to distinguish from P. macrocarpa, compared to which P. thomsonii has wider excipular hyphae, and its exciple is more contrasting in colour: paler inside and with black cortex (McCune, 2017).

*P. tuberculosa (Sm.) Hertel et Knoph — on rocks; 1, 23 [R]; H (sub Cecidonia xenophana). — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Chesnokov et al, 2017), Ala (Spribille et al, 2010).

*Protomicarea alpestris (Sommerf.) McCune — on soil; 23 [R1]; H. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Galanina, 2016), Ala (Thomson, Ahti, 1994).

Protopannaria pezizoides (Weber) P. M. Jorg. et S. Ekman — on soil and plant debris; 1, 5, 14, 17, 26, 31, a8, a16 [R]; H, LECB.

Protoparmelia badia (Hoffm.) Hafellner — on rocks; 1, 5, 31, a6-a8 [R]; H, FR.

*Pseudephebe minuscula (Nyl. ex Arnold) Brodo et D. Hawksw. — on rocks and gravel; 1, 5, 10, 12, a2 [R]; H, LECB. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Thomson, Ahti, 1994).

*P. pubescens (L.) M. Choisy — on rocks; 1, 5, 6, 10-12 [R]; H, FR, LECB. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

*Psora rubiformis (Ach.) Hook. — on soil over rocks; 5 [R1]; LECB. — Kam (Mikulin, 1988), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Poryadina, 2005), Ala (Spribille et al, 2020).

Psoroma hypnorum (Vahl.) Gray var. hypnorum — on soil and plant debris; 14, 21, 26, a12 [R] (Neshataeva, Neshataev, 2018).

*Pycnothelia papillaria Dufour — on soil; 11 [R1]. — Kam (Krasnaya..., 2018), Chu (Andreev et al, 1996), Ala (Dillman et al, 2012).

IPyrenopsis haemaleella (Nyl.) Blomb. et Forssell — on rocks; 26 [R1]; H. — Ala (Spribille et al, 2010). — New to the Russian Far East. The nearest known localities belong to Sverdlovsk Region (Paukov, Teptina, 2012). Differs from other species of the genus by squamulose-areolate thallus with finally open apothecioid ascomata (Jorgensen, 2007).

Ramalina almquistii Vain. — on rocks; 5, a8 [R]; H, FR.

IR. scoparia Vain. — on rocks and soil over rocks; 1, 2, 5, 10, 14, a2, a6-a8 [R]; H, FR, LECB. — Chu (Andreev et al, 1996), Ala (Dillman et al, 2012).

Ramboldia cinnabarina (Sommerf.) Kalb et al. — on bark of Betula divaricata and Pinus pumila; 8, 13, 29 [R]; LECB.

*Rhexophiale rhexoblephara (Nyl.) Hellb. — on mosses; a8 [R1]; H. — Kam (Nylander, 1887), Chu (Zhurbenko, 2009b), Yak (Zhurbenko, 2009b), Ala (Thomson, Ahti, 1994).

*Rhizocarpon badioatrum (Flörke ex Spreng.) Th. Fr. — on rocks; 5, 12, 31 [R]; H, LECB. — Kam (Himelbrant et al., 2014), Chu (Andreev et al., 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

*R. eupetraeoides (Nyl.) Blomb. — on rocks and gravel; 1, 5, 10, 12, a2, a7 [R]; H, LECB. — Kam (Nylander, 1887), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

!R. ferax H. Magn. — on gravel; 6 [R1]; LECB. — Chu (Andreev et al, 1996), Mag (Kotlov,

1995), Yak (Andreev et al, 1996), Ala (Derr, 2010).

*R. geminatum Körb. — on rocks; 31 [R1]; H (sub Lambiella furvella). — Kam (Kuznetsova, Himelbrant, 2006), Chu (Andreev et al, 1996), Ala (Spribille et al, 2020).

R. geographicum (L.) DC. s. lat. — on rocks and gravel; 1, 5, 6, 10-12, a2, a6 [R]; H, FR, LECB (Neshataeva, Neshataev, 2018).

R. grande (Flörke ex Flot.) Arnold — on rocks; 1, 31 [R]; H.

*R. hochstetteri (Körb.) Vain. — on rocks; 26 [R1]; H (sub Porpidia macrocarpa). — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Yak (Andreev et al,

1996), Ala (Spribille et al, 2020).

R. infernulum (Nyl.) Lynge f. infernulum — on rocks; 10-12, 23, a3 [R]; H, LECB.

*R. intersitum Arnold — on gravel; 12 [R1]; LECB. — Kam (Himelbrant et al, 2014), Ala (Spribille et al, 2020).

*R. lavatum (Fr.) Hazsl. — on gravel; 12 [R1]; LECB. — Kam (Himelbrant et al, 2014), Yak (Poryadina, 2005), Ala (Spribille et al, 2020).

*R. lecanorinum Anders — on rocks; 12, 31 [R]; H, LECB. — Kam (Nylander, 1887), Ala (Spribille et al, 2020).

*R. macrosporum Räsänen — on rocks; 1, 31 [R]; H, LECB. — Kam (Himelbrant et al, 2014), Mag (Kotlov, 1995), Yak (Poryadina, 2005), Ala (Spribille et al, 2020).

*R. polycarpum (Hepp) Th. Fr. — on rocks; 2, 10, 31 [R]; H. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

*R. reductum Th. Fr. — on gravel; 12, a3 [R]; LECB. — Kam (Himelbrant et al, 2014), Mag (Kotlov, 1995), Yak (Poryadina, 2005), Ala (Spribille et al, 2020).

*R. rittokense (Hellb.) Th. Fr. — on rocks; 5, 10 [R]; LECB. — Kam (Himelbrant et al, 2014), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Thomson, Ahti, 1994).

!R. sublavatum Fryday — on rocks; 23 [R1]; H. — New to Beringia and Russia. Known from Europe and Asia. The species is somewhat similar to R. lavatum, but has grayish thallus, smaller sessile apothecia with thinner margin, and shorter and broader spores. Morphologically close R. reductum contains stictic acid, which is absent in R. sublavatum (The lichens... , 2009).

!R. viridiatrum (Wulfen) Körb. — lichenicolous on thallus of Aspicilia sp. on gravel; 11 [R1]; LECB. — Mag (Kotlov, 1995), Ala (Weber, Viereck, 1967).

Rimularia badioatra (Kremp.) Hertel et Rambold — on rocks; a8 [R1]; H.

!Rinodina calcigena (Th. Fr.) Lynge — on concrete of lighthouse; 3 [R1]; H. — Yak (Andreev et al, 1996), Ala (Thomson, 1997). — New to the Russian Far East. The species is recognized by heavily pigmented hymenium, spores Bischoffii-type with a wide pigmented band around septum, sometimes pruinose apothecia (slightly pruinose in our specimen) with finally dark margin (Plate II: 3), and inhabiting calcareous substrates (McCune, 2017).

IR. gennarii Bagl. — on seashore rocks and over thallus of Lecanora gangaleoides; 31 [R1]; H. — Yak (Poryadina, 2005), Ala (Spribille et al, 2020). — New to the northern part of Russian Far East, known from Kuril Islands. Recognized by spores Dirinaria-type and maritime habitat (Sheard et al., 2017).

R. septentrionalis Malme — on bark of Salix alaxensis; 18 [R1]; LECB.

R. turfacea (Wahlenb.) Körb. — on soil and plant debris, wood of dwarf shrubs; 1, 5, 6, 10, 11, 14, 21, 23, 26, 31, a12 [O]; LECB.

Ropalospora lugubris (Sommerf.) Poelt — on rocks; 5, a2 [R]; H, LECB.

I#Rosellinula haplospora (Th. Fr. et Almq. ex Th. Fr.) R. Sant. — on thallus of Aspicilia cinerea; 31 [R1]; H. — New to Beringia and Russian Far East. The nearest reported locality is in Trans-Baikal Territory (Zhurbenko, Yakovchenko, 2014). Lichenicolous fungus associated with Aspicilia spp., characterized by large black perithecioid ascomata, immersed in host thallus and flattened at the top, and polyspored asci with simple brown spores (Hafellner, 1985, 2004).

Rusavskia elegans (Link) S. Y. Kondr. et Kärnefelt — on rocks (especially seashore) and concrete of lighthouse; 2, 3, 5, 10, 31, a9 [R].

R. sorediata (Vain.) S. Y. Kondr. et Kärnefelt — on rocks (especially seashore); 2, 10, 31 [R]; H, FR, LECB.

Sagedia mastrucata (Wahlenb.) A. Nordin et al. — on rocks; 1 [R1]; LECB.

IS. zonata Ach. — on rocks; 1 [R1]; LECB. — Yak (Andreev et al, 1996). New to Beringia and Russian Far East. Widespread species with dark gray crustose K- thallus, concave apothecia with dark thalline margin, and ascospores less than 20 ^m long (Foucard, 2001; Nordin et al., 2007).

ISantessoniella arctophila (Th. Fr.) Henssen — on plant debris; a16 [R1]; H (Plate II: 4). — Chu (Kristinsson et al, 2010), Ala (Spribille et al, 2020).

ISarcogyne hypophaea (Nyl.) Arnold — on rocks; 31 [R1]; H (sub Rusavskia sorediata). — Chu (Andreev et al, 1996), Yak (Chesnokov et al, 2015).

+Sarea resinae (Fr.) Kuntze — on resin of Pinus pumila; 13, 29 [R]; LECB.

*Schaereria fuscocinerea (Nyl.) Clauzade et Roux — on rocks; 5 [R1]; H. — Kam (Kuznetsova, Himelbrant, 2006), Chu (Hertel, Andreev, 2003), Mag (Kotlov, 1995), Yak (Poryadina, 2005), Ala (Spribille et al, 2020).

I#Sclerococcum attendendum (Nyl.) Ertz et Diederich — on thallus of Lecidea epiphaea on dead mosses; 5 [R1]; LECB. — Chu (Zhurbenko, 2007).

*# S. pertusariicola (Tuck.) Ertz et Diederich — on thalli of Pertusaria cribellata on rocks; 5, a7 [R]; LECB (sub P. cribellata). — Kam (Himelbrant et al, 2014).

Scoliciosporum umbrinum (Ach.) Arnold — on rocks; 10-12 [R]; LECB.

*Scytinium imbricatum (P. M. Jorg.) Otalora et al. — on soil and plant debris; 5, 10, 14, 31, a16 [R]; H, LECB. — Kam (Nylander, 1887), Chu (Andreev et al, 1996), Ala (Spribille et al, 2020).

*S. tenuissimum (Dicks.) Otalora et al. — on soil over rocks; 2 [R1]; H. — Kam (Neshataeva et al, 2008), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

S. teretiusculum (Wallr.) Otalora et al. — on soil; 14 [R1]; H (sub S. imbricatum).

Solorina crocea (L.) Ach. — on soil; 17, 23, a10 [R] (Neshataeva, Neshataev, 2018).

I#Sphaerellothecium araneosum (Rehm ex Arnold) Zopf — on thalli of Ochrolechia frigida s. lat. on soil and plant debris; 5, 10, 11 [R]; H. — Chu (Zhurbenko, 2007), Yak (Zhurbenko, 2007), Ala (Spribille et al, 2020).

*#S. minutum Hafellner — on thalli of Sphaerophorus fragilis and S. globosus on soil and rocks; 1, 5, 6, 10 [R]; H, LECB. — Kam (Zhurbenko et al, 2012), Yak (Zhurbenko et al, 2005), Ala (Spribille et al, 2010).

Sphaerophorus fragilis (L.) Pers. — on soil and rocks; 1, 5, 6, 10, 12, a6 [R]; FR.

S. globosus (Huds.) Vain. — on soil and rocks; 1, 4-6, 10-12, 14, 15, 21, 23, a8 [O]; H (sub Sphaerellothecium minutum) (Neshataeva, Neshataev, 2018; Neshataeva et al., 2018).

*#Sphinctrina turbinata (Pers.; Fr.) De Not. — on thalli of Pertusaria coriacea on soil and plant debris; 14, a2 [R]; H, LECB. — Kam (Neshataeva et al., 2003), Ala (Thomson, 1997).

!Staurothele areolata (Ach.) Lettau — on rocks; 31 [Ri]; LECB. — Chu (Kristinsson et al., 2010), Ala (Thomson, 1997).

Stereocaulon alpinum Laurer — on soil; 7, 14, 20-23, 30, 32, a4, a17 [O]; H, LECB (Neshataev et al., 2017; Neshataeva, Neshataev, 2018).

*S. arcticum Lynge — on soil; 11 [Ri]; LECB. — Kam (Mikulin, 1988), Chu (Andreev et al., 1996), Ala (Talbot et al, 1997).

*S. arenarium (Savicz) I. M. Lamb — on rocks; 5 [Ri]; H. — Kam (Dombrovskaya, 1996), Chu (Andreev et al, 1996), Mag (Kotlov, 1995), Ala (Spribille et al, 2020).

S. condensatum Hoffm. — on soil; 6, 23, 32 [R] (Neshataeva, Neshataev, 2018).

S. glareosum (Savicz) H. Magn. — on soil and plant debris; 5, 6, 10, 11, 23, 31, a2, a16 [R]; H, LECB (Neshataeva, Neshataev, 2018).

*S. grande (H. Magn.) H. Magn. — on soil; 17, a19 [R]; H, LECB. — Kam (Himelbrant et al, 2014), Yak (Poryadina, 2005), Ala (Spribille et al, 2020).

*S. lambii Dombr. — on soil over rocks; 5 [R1]; LECB. — Kam (Dombrovskaya, 1996), Yak (Chesnokov et al., 2016).

S. paschale (L.) Hoffm. — on soil; 1, 6, 7, 10, 11, 14, 15, 21, 23, 30, 32, a4 [O]; LECB (sub S. alpinum) (Neshataev et al, 2017; Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

S. subcoralloides (Nyl.) Nyl. — on soil over rocks; 5 [R1]; LECB.

S. symphycheilum I. M. Lamb — on rocks; 5, 10 [R]; H, LECB.

S. vesuvianum Pers. — on rocks; 5 [R1]; H.

*#Taeniolella pertusariicola D. Hawksw. et H. Mayrhofer — on thallus of Pertusaria bryontha on plant debris; 11 [R1]; H. — Kam (Zhurbenko et al, 2012), Yak (Zhurbenko, 2009b), Ala (Laursen, Seppelt, 2009). 1

Tephromela atra (Huds.) Hafellner — on rocks; 5, 31 [R]; FR.

!Tetramelas chloroleucus (Körb.) A. Nordin — on bark of Alnus fruticosa, Betula divaricata, B. ermanii, Pinus pumila, on wood of Alnus fruticosa; 6, 8, 19, 27-29, a9 [R]; H, LECB. — Ala (Spribille et al., 2020). New to the northern part of the Russian Far East; known from Khabarovsk Territory (Yakovchenko et al., 2013).

!T. concinnus (Th. Fr.) Giralt — on rocks; 2 [R1]; LECB. — Chu (Andreev et al, 1996), Yak (Andreev et al, 1996).

!T. geophilus (Flörke ex Sommerf.) Norman — on soil; 14 [R1]; LECB. — Chu (Andreev et al, 1996), Yak (Poryadina, 2005).

*T. insignis (Nägeli ex Hepp) Kalb — on plant debris and mosses; 10, 31, a6 [R]; H, FR. — Kam (Neshataeva et al, 2006), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

!T. pulverulentus (Anzi) A. Nordin et Tibell — lichenicolous on thallus of Physconia muscigena on mossy soil; 10 [R1]; H. — Chu (Zhurbenko, 2009b), Yak (Zhurbenko, 2009b), Ala (Dillman et al, 2012).

Thamnolia vermicularis (Sw.) Schaer. [F]; LECB (Neshataev et al, 2017; Neshataeva, Neshataev, 2018; Neshataeva et al, 2018).

var. vermicularis — on soil; 1, 4-7, 10, 11, 14, 15, 21-23, 30, a7, a8.

var. subuliformis (Ehrh.) Schaer. — on soil; 5-7, 10-12, 14, 20, 22, 23, a7.

IToensbergia geminipara (Th. Fr.) T. Sprib. et Resl — on soil; 7 [R1]; H. — Chu (Andreev et al., 1996), Yak (Andreev et al., 1996), Ala (Spribille et al, 2020).

IToninia squalida (Ach.) A. Massal. — on soil; a4 [R1]; LECB (sub Polychidium muscicola). — Chu (Andreev et al, 1996), Ala (Spribille et al, 2020).

Toniniopsis subincompta (Nyl.) Kistenich et al. — on bark of Alnus fruticosa and Salix alaxensis; 18, 19 [R]; LECB.

*Trapelia glebulosa (Sm.) J. R. Laundon — on gravel; 12 [R1]; LECB. — Kam (Nylander, 1887), Ala (Orange, 2018).

Trapeliopsis flexuosa (Fr.) Coppins et P. James — on bark of Pinuspumila and on driftwood; 13, a15 [R].

T. granulosa (Hoffm.) Lumbsch — on soil; 1, 6, 10, 13 [R].

Tremolecia atrata (Ach.) Hertel — on rocks and gravel; 1, 5, 6, 11, 12, a2, a7 [R]; H, FR, LECB.

*Umbilicaria cylindrica (L.) Delise — on rocks; 5, 12, a7 [R]; H. — Kam (Mikulin, 1988), Chu (Andreev et al, 1996), Yak (Andreev et al, 1996), Ala (Spribille et al, 2020).

U. deusta (L.) Baumg. — on rocks; 31 [R1]; H.

U. hyperborea (Ach.) Hoffm. var. hyperborea — on rocks and gravel; 1, 5, 6, 10, 12, 31, a2, a6 [R]; LECB (sub Parmelia skultii).

U. proboscidea (L.) Schrad. — on rocks; 1, 5, 10, 12, a6-a8 [R]; H, FR.

U. torrefacta (Lightf.) Schrad. — on rocks; 5, 12, a6-a8 [R]; H.

U. vellea (L.) Hoffm. — on rocks; 5, 31 [R]; LECB.

Varicellaria rhodocarpa (Körb.) Th. Fr. — on plant debris and on bark of Pinus pumila; 5, 8, 29 [R]; LECB.

IVerrucaria cf. viridula (Schrad.) Ach. — on rocks; 17 [R1]; LECB. — Yak (Kristinsson et al, 2010), Chu (Kristinsson et al, 2010).

Vulpicida juniperinus (L.) J.-E. Mattson et M. J. Lai — on bark of Pinus pumila; 8, 13, 29 [R]; LECB.

V. pinastri (Scop.) J.-E. Mattson et M. J. Lai — on soil, bark of Alnus fruticosa, Betula divaricata, B. ermanii, B. exilis, Pinus pumila, Sorbus sambucifolia; 9, 13, 24, 26-29, a13 [O].

*Xanthoparmelia stenophylla (Ach.) Ahti et D. Hawksw. — on rocks; 2, 10 [R]; H, LECB. Thalli contain usnic and salazinic acids. — Kam (Himelbrant et al., 2014), Chu (Andreev et al., 1996), Mag (Kotlov, 1995), Yak (Andreev et al, 1996), Ala (Thomson, Ahti, 1994).

Xylographa parallela (Ach.) Fr. — on driftwood; a15 [R1]; LECB.

X. vermicularis T. Sprib. — on wood (incl. driftwood); 23, 32, a6, a15, a24 [R]; LECB. Thalli contain stictic and confriesiic acids.

The revealed diversity of lichens and allied fungi of Cape Goven counts 394 species: 373 lichens (including 5 lichenicolous species), 18 lichenicolous fungi and 3 non-li-chenized saprobic fungi related to lichens. Altogether 4 species are new to Russia (Miriquidica pulvinatula, Myriolecis andrewii, Ochrolechia alaskana, Rhizocarpon sub-lavatum), 1 — to Asiatic Russia (Collemopsidiumfoveolatum), 29 other species are new to the Russian Far East, 4 — to the northern part of the Far East. Additionally, 51 species are new to Kamchatka Territory, and 92 other species are new to Koryakia. Among the new species to Russia or Russian Far East, 11 are also reported for the first time for

Beringia. According to current results and earlier data (Himelbrant et al., 2018; Ne-shataeva, Neshataev, 2018), a total of 500 species of lichens and allied fungi are known from Koryakia now, mainly from Koryak State Reserve.

Compared to the neighbouring regions, the specificity of the lichen flora of Cape Goven is not high, but, taking into account the amount of new species to Beringia and Russian Far East, it is higher than in Parapolsky Dale (Himelbrant et al., 2018). Most of the species reported here for the first time for Koryakia are known from other compared regions, especially Alaska (151 species of 181 new to Koryakia), which has ecologically similar places, and is best studied. Eleven species only (3% of total diversity of the lichen flora of Cape Goven) represent new records for Beringia and thus reflect low specifity of lichen diversity of the investigated territory. It is noteable that comparably low specifity was discovered in the lichen flora of Parapolsky Dale — 8 species, 2.5% (Himelbrant et al., 2018). This demonstrates low specifity of Koryakian lichen flora in general.

Like in Parapolsky Dale, in Cape Goven most diverse are lichens in rocky habitats. Due to mountainous landscape of the study area, rocky outcrops are abundant, and the number of species (238 species or 60% of the lichen flora) in such habitats here is significantly higher than in rocky biotopes in Parapolsky Dale (147 species). Moreover, rocky habitats are diverse in Cape Goven, and lichen communities on the coast differ from the rest area: 205 species were found on inland rocks surrounded by tundra (of them 80 species were specific, i. e. found only in this habitat type), and 76 — on seashore rocks (with 31 species occurring only on the seashore rocks within the study area). The tundra is also significantly reacher here than in Parapolsky Dale: 223 species, or 57% of the lichen flora were found in tundra in the study area, versus 139 species in Parapolsky Dale. At the same time, due to relatively low cover of shrublands in Cape Goven, the total amount of lichens in such habitats is low — 71 species, 18% of the lichen flora (Pinus pumila shrubs — 48 species, Alnus fruticosa — 37 species, Betula divaricata — 21 species). Forests are almost absent in the study area, only Salix spp. "forests", usually not higher than 2-3 m, survive in some stream valleys. In such flood-plain willow stands 19 species (5% of the lichen flora) were found, including Nephroma bellum and Peltigera collina, which are more typically forest species. Bogs in the study area are not widespread and not very rich in lichens as well (19 species, 5%). In contrast, in Parapolsky Dale a significant part of lichen diversity is represented in shrub-lands, forests, and bogs (Himelbrant et al., 2018). Quite scarce and poor is driftwood on the shoreline of Cape Goven, probably it is not preserved for a long time on one place: previously it could be collected and used by people, and nowadays the existing logs seem to move much during storms. Anyway, 13 species were found on driftwood, of them only Lecanora orae-frigidae and Xylographa parallela were not recorded in other biotopes. Several species were found in the study area exclusively due to the presence of concrete construction of the lighthouse (13 species altogether were found there, including 7 specific). An average number of species per standard sample plot is 36.9+4.5 (which doesn't differ significantly from Parapolsky Dale): 6-15 species per

sample plot in floodplain willow stands, 11-31 in shrublands, 12-60 in tundras, 24-63 on seashore rocks, and 92-111 on inland rocky outcrops.

Unlike Parapolsky Dale, where epiphytic lichens are most diverse, in Cape Goven the richest substrates are soil and plant debris (181 species, 46% of the lichen flora). Saxicolous lichens are represented by 153 species (39% of the lichen flora), of which 147 were collected from rocks and gravel, and 9 — from concrete. Epiphytic and lignicolous lichens are few (63 species, 16% of the lichen flora, and 22 species, or 6%, respectively); the richest phorophytes are Alnus fruticosa (29 species) and Pinus pumila (31 species). Lichenicolous fungi and lichens count 23 species, muscicolous lichens — 17 species. Single species were also found on resin of Pinus pumila, bones, and barnacles.

Probably due to habitat mosaicism within the study area, the percent of rare species is very high, even higher than in Parapolsky Dale — altogether 347 species or 88% of the studied biota are "rare" [R]; of them 153 were recorded once [RJ. Only 37 species (9%) occur "occasionally" [O], and 10 species (3%) were found "frequently" [F]: Cetraria laevigata, Cladonia arbuscula, C. rangiferina, C. uncialis, Gowardia nigricans, Lecanora symmicta, Ochrolechia frigida, Parmeliopsis ambigua, P. hyperopta, Thamno-lia vermicularis. Altogether 3 species of the recorded in Cape Goven are red-listed in Kamchatka and Russia (Krasnaya..., 2008, 2018): Cetraria kamczatica, Cetrelia alaska-na, Lichenomphalia hudsoniana.

The intensity and methods of investigation, as well as the amount of collected material did not vary significantly during the field studies of 2016 in Parapolsky Dale (Himelbrant et al., 2019) and 2018 in Cape Goven. Nevertheless, the major difference in the number of revealed species was found: 315 in Parapolsky Dale and 394 in Cape Goven (79 species or 25% increase). The high portion of new species to Koryakia and Kamchatka Territory (181, 46% of recorded species) demonstrates the insufficiency of our knowledge about the regional lichen diversity and the noticeable variation in lichen biota of different parts of Koryakia.

Acknowledgments

The head and the staff of Kronotsky State Reserve, as well as inspectors of Ko-ryaksky State Reserve are thanked for help in organization and support of the expedition. We would like to express our special gratitude to inspector Anatoly Soro-kin for friendly and invaluable help during field studies. We are also very grateful to Marina P. Vyatkina (Kamchatka Branch of Pacific Institute of Geography FEB RAS, Petropavlovsk-Kamchatsky) for kind assistance. We thank Ivan Frolov (Botanical Garden of the Urals Branch of Russian Academy of Sciences, Yekaterinburg) for revision of Caloplaca s. lat. specimens, Liudmila A. Konoreva and Sergey V. Chesnokov for revision of Micarea specimens, Lidia S. Yakovchenko for confirming specimen of Candelariella placodizans, Toby Spribille (Edmonton, Canada) for help in finding information on species distribution in Alaska, and to Leena Myllys and Saara Velmala (University of Helsinki, Finland) for assistance in organizing our studies in H. We also

express our gratitude to the reviewers for valuable comments. The study was carried out within the framework of the institutional research project "Flora and systematics of algae, lichens, and bryophytes of Russia and phytogeographically important regions of the world" (no. 121021600184-6) of the Komarov Botanical Institute RAS and supported by Russian Foundation for Basic Research (expedition: grants 16-05-00736, 18-05-60093; laboratory research: grants 18-05-60093, 19-05-00805).

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