Каталог типовых экземпляров Colias Fabricius, 1807, хранящихся в естественно-историческом музее университета им. Гумбольдта в Берлине (Lepidoptera, Pieridae)

Корб С.К.

АРК_I © Амурский зоологический журнал. V¡(2), 2014.182-206 i Accepted: 7.04.2014

УДК 595.789 ¡ © Amurian zoological journal. Vl(2),2014.182-206 ¡ Published-. 30.06.2014

A CATALOGUE OF THE TYPE SPECIMENS OF COLIAS FABRICIUS, 1807 DEPOSITED IN THE MUSEUM FÜR NATURKUNDE AN DER HUMBOLDT-UNIVERSITÄT ZU BERLIN

(LEPIDOPTERA, PIERIDAE)

S.K. Korb

[Корб С.К. Каталог типовых экземпляров Colias Fabricius, 1807, хранящихся в естественно-историческом музее Университета им. Гумбольдта в Берлине (Lepidoptera, Pieridae)]

Russian Entomological Society, Nizhny Novgorod Branch, a/ya 97 Nizhny Novgorod 603009 Russia. E-mail: stanislavkorb@list.ru

Русское Энтомологическое Общество, Нижегородское отделение, а/я 97, Н.Новгород 603009 Россия. E-mail: stanislavkorb@list.ru

Key words: Colias, type specimens, catalogue, lectotype designations, neotype designations, Museum für Naturkunde an der Humboldt-Universität zu Berlin

Ключевые слова: Colias, типовые экземпляры, каталог, обозначения лектотипов, обозначения неотипов, естественно-исторический музей университета им. Гумбольдта в Берлине

Summary. A catalogue of the type specimens of Colias Fabricius, 1807 of the Museum für Naturkunde an der HumboldtUniversität zu Berlin is given, information about all type specimens of this genus and their labels is listed in alphabetical order (274 type specimens: 79 primary types (1 neotype, 7 holotypes, 32 syntypes, 39 lectotypes) and 195 secondary types (11 paratypes, 184 paralectotypes)). The lectotype and neotype specimens established in current paper are figured. Lectotypes of the following Holarctic taxa are designated: Colias melinos var. alpina Elwes, 1899, Colias baeckeri Kotzsch, 1930, Colias palaeno var. cretacea Aurivillius, 1888, Colias cocandica culminicola Kotzsch, 1936, Colias aurora var. decolorata Staudinger, 1898, Colias erschoffi Alphéraky, 1881, Colias felderi Grum-Grshimailo, 1891, Colias heldreichi Staudinger, 1862, Colias helichtha Lederer, 1853, Colias sifanica herculeana Bollow, 1930, Colias lada Grum-Grshimailo, 1891, Colias libanotica Lederer, 1858, Colias marcopolo Grum-Grshimailo, 1888, Colias palaeno var. orientalis Staudinger, 1892, Colias palaenopoktussani O.Bang-Haas, 1935, Colias romanovi Grumm-Grshimailo, 1885, Colias sagartia Lederer, 1869, Colias ladakensis seitzi Bollow, 1930, Colias sieversi Grum-Grshimailo, 1887, Colias sifanica Grum-Grshimailo, 1891, Colias aurora thia O.Bang-Haas, 1934, Colias aurorina var. transcaspica Christoph, 1887. Lectotypes of the following Neotropic taxa are designated: Colias lesbia var. andina Staudinger, 1894, Colias lesbia var. arena Staudinger, 1899. The neotype of Colias aurora var. kenteana Heyne, [1895] is designated. A replacement name is proposed to resolve the case of primary homonymy: Colias arida hisilomeanus Korb, nom. nov. proflavescens Rühl, 1893, necflavescens Gabrowski, 1892. Резюме. Предлагается каталог типовых экземпляров Colias Fabricius, 1807 Музея естественной истории университета им. Гумбольдта в Берлине, информация обо всех хранящихся в Музее типовых экземплярах представлена в алфавитном порядке (274 типовых экземпляра: 79 номенклатурных типов (1 неотип, 7 голотипов, 32 синтипа, 39 лектотипов) и 195 прочих типов (11 паратипов, 184 паралектотпа)). Лектотипы и неотипы, обозначенные в настоящем сообщении, изображены. Обозначаются лектотипы следующих голарктических таксонов: Colias melinos var. alpina Elwes, 1899, Colias baeckeri Kotzsch, 1930, Colias palaeno var. cretacea Aurivillius, 1888, Colias cocandica culminicola Kotzsch, 1936, Colias aurora var. decolorata Staudinger, 1898, Colias erschoffi Alphéraky, 1881, Colias felderi Grum-Grshimailo, 1891, Colias heldreichi Staudinger, 1862, Colias helichtha Lederer, 1853, Colias sifanica herculeana Bollow, 1930, Colias lada Grum-Grshimailo, 1891, Colias libanotica Lederer, 1858, Colias marcopolo Grum-Grshimailo, 1888, Colias palaeno var. orientalis Staudinger, 1892, Colias palaeno poktussani O.Bang-Haas, 1935, Colias romanovi Grumm-Grshimailo, 1885, Colias sagartia Lederer, 1869, Colias ladakensis seitzi Bollow, 1930, Colias sieversi Grum-Grshimailo, 1887, Colias sifanica Grum-Grshimailo, 1891, Colias aurora thia O.Bang-Haas, 1934, Colias aurorina var. transcaspica Christoph, 1887. Обозначены лектотипы следующих неотропических таксонов: Colias lesbia var. andina Staudinger, 1894, Colias lesbia var. arena Staudinger, 1899. Обозначается неотип Colias aurora var. kenteana Heyne, [1895]. Предлагается замещающее название: Colias arida hisilomeanus Korb, nom. nov. pro flavescens Rühl, 1893, nec flavescens Gabrowski, 1892.

INTRODUCTION

The Museum für Naturkunde an der Humboldt-Universität zu Berlin is a depository of the largest and most important reference collection of Lepidoptera in Europe [Nekrutenko, 2000], containing the most important type specimens of O.Staudinger, A. and O. Bang-Haas, R.Püngeler and others. The complete description of this collection has been made by Y.P. Nekrutenko [2000].

This publication is one of the results of my research visits to the Museum für Naturkunde an der Humboldt-Universität zu Berlin in 2010-2014: 2010, Oktober; 2011, December; 2012, April; 2013, January; 2014, February.

In the present publication the complete list of type specimens (including all type specimens categories: holotypes, paratypes, lectotypes, paralectotypes, neotypes

and syntypes) of Colias in the Museum für Naturkunde an der Humboldt-Universität zu Berlin is arranged in alphabetical order. The lectotypes are designated for several Colias taxa; all of these lectotype specimens are figured according to the Rec. 74B of the Code.

The Colias collection in the Museum für Naturkunde an der Humboldt-Universität zu Berlin now housed in the 3rd floor of the main Museum building at the address Invalidenstrasse 43, Berlin. It occupies 3 wooden cabinets (fig. 1A); the whole collection was rearranged and partly reorganized by J.Grieshuber in the beginning of the current century (two cabinets: middle and lower) (fig. 1B). But it was done to O.Staudinger and A. and O. Bang-Haas collections; some other drawers are still in the old conditions (specimens are placed very compact, using ancient system and names, etc.) (upper cabinet) (fig. 1C).

In this paper 24 lectotypes (22 for Holarctic taxa, 2 for Neotropical taxa) and 1 neotype are designated. Alltogether data about 274 type specimens are listed: 79 primary types (1 neotype, 7 holotypes, 32 syntypes, 39 lectotypes) and 195 secondary types (11 paratypes, 184 paralectotypes). The taxonomic status and position of every listed taxon is given, as well as the original combination and original description citation, and primary type designation data.

LIST OF SPECIMENS

afghana O.Bang-Haas, 1927

Colias hyale afghana Bang-Haas, 1927: 42; pl. 5, fig. 23.

Taxonomic status and position: junior subjective synonym of C. erate (Esper, [1805]) [Grieshuber et al., 2011: 36]. Very variable species with huge range of klinal variability, also known as a good migrant.

J alba Röber, 1909

[Colias euxanthe] ab. alba Röber, 1909: 90.

Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

Fig. 1. A - C: the Colias collection in the Museum für Naturkunde an der Humboldt-Universität zu Berlin: A - cabinets; B - drawer of reorganized part; C - drawer of non-reorganized part.

J alba Staudinger, 1871

[Colias myrmidone] ab. alba Staudinger, 1871: 6. Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

Syntypes 29: 1$: printed, white paper 'ex coll. | Staudinger'; printed, pink paper 'Origin.'; phand-written, yellow paper 'Wien'; printed, white paper '[Wien] | infrasubspecific taxon | name bearing specimen of Colias | myrmidone [ab.] alba | Staudinger, 1871 | det. Grieshuber, 2007'; 1$: printed, white paper 'ex coll. | Staudinger'; printed, pink paper 'Origin.'; handwritten, white paper 'Wien'; white circle; printed, white paper '[Wien] | infrasubspecific taxon | name bearing specimen of Colias | myrmidone [ab.] alba | Staudinger, 1871 | det. Grieshuber, 2007'.

J alba Röber, 1909 [Coliasphilodice] ab. alba Röber, 1909: 91. Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

alexandra Röber, 1907 C.[olias] wiskotti alexandra Stgr. i.l., Röber, 1907: 70 see rueckbeili Lukhtanov, 1994

J albina Röber, 1909 [Colias eurytheme] ab. albina Röber, 1909: Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

Taxonomic status and position: a good species. Lectotype 3: printed, white paper 'ex coll. | R.Püngeler'; printed, yellow paper 'GART | Exemplar

Taxonomic status and position: a good subspecies of C. nastes Boisduval, 1832.

alpherakii Staudinger, 1882 Colias Alpherakii Staudinger, Bang-Haas, 1882: 164 - 166. Lectotype designated by: Grieshuber, Worthy, 2006: 71 - 72; fig. 26.

Taxonomic status and position: a good species.

alpina Elwes, 1899

Colias melinos, var. alpina Elwes, 1899: 304, 319-320.

Taxonomic status and position: junior objective synonym of C. tyche (Böber, 1812) [Grieshuber et al., 2011: 43].

Lectotype < (col. pl. IX: 1-3), designated here according to the Art. 61.1 and 74.1 of the Code: 1<: printed, numbers handwritten, white paper 'ex coll. 1/6 | Staudinger'; printed, red paper 'Syntype | Colias melinos | [var.] alpina Etwes, 1899 | det. Grieshuber, 2007'; handwritten, white paper 'Melinos'; handwritten, white paper 'Lectotype <| alpina Elwes | S.Korb des. 2014'; green circle; printed, date handwritten, white paper 'S.E.Altai. | Tchuja Mts. | 6-8000 ft. | 5.7.98 | H.J.Elwes.'.

alta Staudinger, 1886

Col.[ias] Hyale L. var. Alta Staudinger, 1886: 200 -201. Lectotype designated by: Korb, 2012: 29.

Taxonomic status and position: a good species.

andina Staudinger, 1894

Colias Lesbia Fab. (var. Andina) Staudinger, 1894: 64.

Taxonomic status and position: a good subspecies of C. lesbia (Fabricius, 1775)

Lectotype 3 (col. pl. XIII: 70-72), designated here according to the Art. 61.1 and 74.1 of the Code: 13: printed, pink paper 'Origin.'; printed, red paper 'Colias lesbia andina | Staudinger, 1894 | Syntype | Grieshuber det. XII.2004'; handwritten, white paper 'var. Andina | Stgr.'; handwritten, white paper 'Lectotype 3 I andina Stgr. | S.Korb des. 2014'; handwritten, white paper 'Cocapata | Bolivia 3500 m. | 92 Garlepp'.

arena Staudinger, 1898

Colias Lesbia Fab. var. (?) arena Staudinger, 1898a: 23.

Taxonomic status and position: remains uncertain; possible a synonym of nominotypical subspecies.

Colias Wiskotti Stgr. var. (ab.) Aurantiaca Staudinger, 1892a: 230 - 232. Lectotype designated by: Grieshuber, Worthy, 2006: 57; fig. 7.

Taxonomic status and position: junior subjective

synonym of C. wiskotti separata Grum-Grshimailo, 1888 [Grieshuber, Worthy, 2006].

Colias wiskotti aurea Kotzsch, 1936: 45. Lectotype designated by: Grieshuber, Worthy, 2006: 64; fig. 16.

Taxonomic status and position: according to J.Grieshuber and B.Worthy [2006: 65], "the taxon is provisionally accepted as a subspecies of C. wis-kotti"; this opinion was not changed in the last Colias revision [Grieshuber et al., 2011: 54].

ten, white paper 'aurea 3? Cotyp 55 -'; handwritten, white paper '6/7 wiskotti'.

baeckeri Kotzsch, 1930.

Colias baeckeri Kotzsch, 1930: 236 - 237.

Taxonomic status and position: junior subjective synonym of C. wanda Grum-Grshimailo, 1907 [Grie-shuber, Churkin, 2003].

Lectotype 3 (col. pl. IX: 4-6), designated here according to the Art. 61.1 and 74.1 of the Code: 3: printed, red paper 'Co-Type | e Collection | Kotzsch'; printed, white paper '734.'; printed, white paper 'Pass Dingtsiangmiau | Richthofengebirge | Kansu 2800 m'; printed, red paper 'Syntype | Colias baeckeri Kotzsch, 1930 | det. Grieshuber, 2007'; handwritten, white paper 'Lectotype 3| baeckeri Kotzsch. | S.Korb des. 2014'.

J banghaasi Bollow, 1930

[Colias christophi] f. banghaasi Bollow, 1930: 108.

Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

Colias blameyi Jörgensen, 1916: 508. Lectotype designated by: Lamas in Verhulst, 2000: 222.

Taxonomic status and position: a good subspecies of C. weberbaueri Strand, 1912.

boothii Curtis, 1835

Colias Boothii Curtis, 1835: LXV - LXVI; pl. A, figs. 3-5.

Taxonomic status and position: probable, a subspecies of C. tyche (Böber, 1812) [Grieshuber et al., 2011: 63]. In the last revision of the genus Colias [Grieshuber et al., 2011: 62] the labelling of the syntypic specimen from the Museum listed incorrect: the red label listed as "Syntype...", but it is in fact "Paralectotype...".

Colias Myrmidone v? Caucasica Staudinger, 1871: 6. Lectotype designated by: Nekrutenko, 1990: 99.

Taxonomic status and position: a good species.

Lectotype 3: printed, numbers handwritten, white paper 'ex coll. 1/3 | Staudinger'; printed, white paper 'Coll. Led.'; printed, white paper 'void lecto-type designation; | no syntype of | Colias caucasica Staudinger, | 1871 | det. Grieshuber 2007'; handwritten, white paper 'Helenendorf'; printed, handwritten name and author, yellow paper 'LECTOTYPUS 3 | caucasica Stgr. | Y.Nekrutenko design.'; printed, name and author handwritten (Staudinger's hand), pink paper 'Origin. | Caucasica | Stgr.'.

Paralectotypes 13, 19: 13: printed, numbers handwritten, white paper 'ex coll. 3/3 | Staudinger'; printed, white paper 'Coll. Led.'; printed, white paper 'void lectotype designation; | no syntype of | Colias caucasica Staudinger, | 1871 | det. Grieshuber 2007'; printed, handwritten name and author, yellow paper 'PARA-LECTOTYPUS 3 | caucasica Stgr. | Y.Nekrutenko design.'; printed, name and author handwritten (Stauding-er's hand), pink paper 'Origin. | Caucasica | Stgr.'; 19: printed, numbers handwritten, white paper 'ex coll. 2/3 | Staudinger'; printed, white paper 'Coll. Led.'; printed, white paper 'void lectotype designation; | no syntype of | Colias caucasica Staudinger, | 1871 | det. Grieshuber 2007'; printed, handwritten name and author, yellow paper 'PARALECTOTYPUS 9 | caucasica Stgr. | Y.Nekrutenko design.'; printed, name and author handwritten (Staudinger's hand), pink paper 'Origin. | Caucasica | Stgr.'; handwritten, white paper 'Caucas'. chlorocoma Christoph, 1888

Colias chlorocoma Christoph, 1888: 308. Lectotype designated by: Nekrutenko, 1990: 102.

Taxonomic status and position: a good species.

Paralectotypes 33: 13: printed, numbers handwritten, white paper 'ex coll. 1/3 | Staudinger'; handwritten, green paper 'Kasikoporan | Chr.'; printed, red paper 'Paralectotype | Colias chlorocoma | Christoph, 1888 | det. Grieshuber, 2007'; 13: printed, numbers handwritten, white paper 'ex coll. 2/3 | Staudinger';

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cretacea Aurivillius, 1888

C.[olias] Palaeno L. Var. cretacea Schilde, Aurivillius, 1888: 6. The authorship and publication date given to Aurivillius as he raised infrasubspecific status of ab. cretacea Schilde, 1884 to the subspecific rank.

Taxonomic status and position: junior objective synonym of C. palaeno lapponica Staudinger, 1861.

culminicola Kotzsch, 1936

Colias cocandica culminicola Kotzsch, 1936: 44 - 45.

Taxonomic status and position: a subspecies of C. cocandica Erschoff, 1874 [Grieshuber et al., 2011]; should be revised using molecular and statistical data.

Lectotype $ (col. pl. IX: 10-12), designated here according to the Art. 61.1 and 74.1 of the Code: 1$: printed, yellow paper 'GART | Exemplar und Eti- | ketten dokumentiert | specimen and label | data docu-

decolorata Staudinger, 1898

Col.[ias] Aurora Esp. und var. (ab.) Decolorata Staudinger, 1898b: 322.

Taxonomic status and position: junior subjective synonym of C. heos heos (Herbst, 1792).

Lectotype S (col. pl. IX: 13-15), designated here according to the Art. 61.1 and 74.1 of the Code: 1S: printed, numbers handwritten, white paper 'ex coll. 2/9 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Kentei] | Syntype | Colias aurora [var.] | decolorata Staudinger, | 1898 | det. Grieshuber, 2007'; handwritten, green paper 'Kentei | 89 Dörr.'; handwritten, white paper 'Lectotype S | decolorata Stgr. | S.Korb des. 2014'.

diva Grum-Grshimailo, 1891 Colias Diva Grum-Grshimailo, 1891: 449 - 450. Lectotype designated by: Grieshuber, Churkin, 2003: 245 - 246. Taxonomic status and position: a good species. Paralectotypes 13, 49: 13: handwritten, white paper 'Aurora var.?'; handwritten, brown paper 'Thibet | Dschachar mont | 90 Gr. Gr.'; printed, red paper '[loc. Dzhakhar, Mudzhik] | Paralectotype | Colias diva | Grum-Grshimailo, 1891 | Horae Soc. ent. ross. | 25: 229-450, designated | by J.Grieshuber & | S.Churkin, 2003'; 19: printed, numbers handwritten, white paper 'ex coll. 2/5 | Staudinger'; handwritten, white paper 'Diva | mil <sic!> Silberfl.'; handwritten, white paper 'p. Myg^HKb <r.[iver] Mudzhik> | 9.VII 90'; handwritten, white paper 'Amdo'; printed, red paper '[loc. Dzhakhar, Mudzhik] | Paralectotype | Colias diva | Grum-Grshimailo, 1891 | Horae Soc. ent. ross. | 25: 229-450, designated | by J.Grieshuber & | S.Churkin, 2003'; 19: printed, numbers handwritten, white paper 'ex coll. 3/5 | Staudinger'; handwritten, white paper '9.VII.90'; handwritten, white paper 'Myg^HKb <Mudzhik>'; printed, red paper '[loc. Dzhakhar, Mudzhik] | Paralectotype | Colias diva | Grum-Grshimailo, 1891 | Horae Soc. ent. ross. | 25: 229-450, designated | by J.Grieshuber & | S.Churkin, 2003'; brown circle; 19: printed, numbers handwritten, white paper 'ex coll. 4/5 | Staudinger'; printed, red paper '[loc. Dzhakhar, Mudzhik] | Paralectotype | Colias diva | Grum-Grshimailo, 1891 | Horae Soc. ent. ross. | 25: 229-450, designated | by J.Grieshuber & | S.Churkin, 2003'; brown circle; 19: printed, numbers handwritten, white paper 'ex coll. 5/5 | Staudinger'; printed, red paper '[loc. Dzhakhar, Mudzhik] | Para-lectotype | Colias diva | Grum-Grshimailo, 1891 | Horae Soc. ent. ross. | 25: 229-450, designated | by J.Grieshuber & | S.Churkin, 2003'; brown circle. eos Herrich-Schäffer, 1848 Colias Eos Herrich-Schäffer, 1848: pl. 81, figs. 395, 396; pl. 82, figs. 397, 398. Lectotype designated by: Grieshuber et al., 2011: 94.

Taxonomic status and position: junior subjective synonym of C. thisoa Menetries, 1832.

ermak Groum-Grshimaïlo, 1890

Colias Myrmidone var. Ermak Groum-Grshimaïlo, 1890: 252, 301 (footnotes). Lectotype designated by: Grieshuber, 2006a: 93 - 94; pl. 16, fig. 3.

Taxonomic status and position: junior subjective synonym of C. myrmidone myrmidone (Esper, [1781]) [Grieshuber, 2006a].

Colias Erschoffi Alphéraky, 1881: 362 - 365; pl. 14, fig. 1, 2.

Note. Different specimens from the type series were named as "lectotype" [Tshikolovets, 2005: pl. 36, fig. 13] or even "paratype" [Verhulst, 1991: fig. 5]. S.Alphéraky did not provide any holotype designation, so paratypes are impossible in his type series; no valid lectotype designations have been published too, so "lectotype" is also impossible. Valid lectotype designation is given below.

Taxonomic status and position: a good species.

Lectotype S (col. pl. IX: 16-18), designated here according to the Art. 61.1 and 74.1 of the Code: 1S: printed, numbers handwritten, white paper 'ex coll. 1/2 | Staudinger'; printed, red paper 'Syntype | Colias erschoffii | Alphéraky, 1881 | det. Grieshuber, 2007'; handwritten, white paper 'Lectotype S | erschoffi Alph. | S.Korb des. | 2014'; handwritten, brown paper 'Tian Schan | Alph.'; handwritten, white paper '15 V 1879. | Tian-chan'.

felderi Grum-Grshimailo, 1891

Colias Felderi Grum-Grshimailo, 1891: 448 - 449.

Taxonomic status and position: a good species.

hisilomeanus Korb, nom. nov.

Colias arida hisilomeanus Korb, nom. nov. pro

C. arida flavescens Rühl, 1893, nec C. myrmidone flavescens Gabrowski, 1892.

Taxonomic status and position: a good subspecies of C. arida Alpheraky, 1889.

Etymology: "To the west of Thangorodrim lay Hisilome, the Land of Mist, for so it was named by the Noldor in their own tongue because of the clouds that Morgoth sent thither during their first encampment..." [Tolkien, 2013: 109].

Holotype (by monotypy) $: printed, red paper 'Holotype | Colias eogene flavescens | Rühl, 1893 | det. Grieshuber, 2007'; handwritten, white paper 'E 543.'; handwritten, white paper 'eogene - 31. | flavescens | Rühl'; handwritten, white paper 'Turkestan'. fontainei Reissinger, 1989

Colias alfacariensis fontainei Reissinger, 1989: 141 - 145; Taf. 39, Fig. 3; Taf. 41, Fign. 5 - 8; Taf. 42, Fign. 1 - 16; Taf. 43, Fign. 1 - 3, 5 - 11, 13, 14; Taf. 44, Fign. 1 - 14; Taf. 47, Fig. 3; Taf. 76, Fig. 9; Taf. 77, Fig. 9.

Taxonomic status and position: junior objective synonym of nominotypical C. alfacariensis Ribbe, 1905.

J guatemalena Röber, 1909

C.[olias] philodice guatemalena Röber, 1909: 91. Lectotype designated by: Lamas, 1993: 140.

Taxonomic status and position: listed as infrasu-bspecific taxon [Verhulst, 2000].

Colias Heldreichi Staudinger, 1862: 257 - 264.

Taxonomic status and position: a good subspecies of C. aurorina Herrich-Schäffer, 1850.

Lectotype $ (col. pl. X: 22-24), designated here

according to the Art. 61.1 and 74.1 of the Code: 1$: printed, handwritten numbers, white paper 'ex coll. 3/6 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias heldreichi | Staudinger, 1862 | det. Grieshuber, 2007'; handwritten, white paper 'Ve-luchi | Kr.'; yellow circle; handwritten, white paper 'Lectotype S | heldreichi Stgr. | S.Korb des. 2014'.

Paralectotypes 2S, 3$: 1S: printed, handwritten numbers, white paper 'ex coll. 1/6 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias heldreichi | Staudinger, 1862 | det. Grieshuber, 2007'; handwritten, yellow paper 'Velu-chi | Kr.'; handwritten, white paper '5/7'; 1S: printed, handwritten numbers, white paper 'ex coll. 2/6 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias heldreichi | Staudinger, 1862 | det. Grieshuber, 2007'; handwritten, white paper 'Veluchi | Kr.'; yellow circle; 1$: printed, handwritten numbers, white paper 'ex coll. 4/6 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias heldreichi | Staudinger, 1862 | det. Grieshuber, 2007'; handwritten, white paper 'Veluchi | Kr.'; yellow circle; handwritten, white paper '15/7'; 1$: printed, handwritten numbers, white paper 'ex coll. 5/6 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias heldreichi | Staudinger, 1862 | det. Grieshuber, 2007'; handwritten, white paper 'Veluchi | Kr.'; yellow circle; 1$: printed, handwritten numbers, white paper 'ex coll. 6/6 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias heldreichi | Staudinger, 1862 | det. Grieshuber, 2007'; handwritten, white paper 'Veluchi | Kr.'; yellow circle; green rectangle. helichtha Lederer, 1853

Colias Helichtha Lederer, 1853: 18, 33.

Taxonomic status and position: junior subjective synonym of C. erate (Esper, [1805]) [Grieshuber et al., 2011].

Lectotype S (col. pl. X: 25-27), designated here

Paralectotype $: printed, white paper 'ex coll. | Staudinger'; printed, white paper 'Coll. Led.'; printed, red paper 'Syntype | Colias helichtha | Lederer, 1853 | det. Grieshuber, 2007'.

herculeana Bollow, 1930.

C.[olias] sifanica herculeana Bollow, 1930: 109; pl. 7, row f.

Taxonomic status and position: junior subjective synonym of C. sifanica sifanica Grum-Grshimailo, 1891.

Lectotype S (col. pl. X: 28-30), designated here

herzi Staudinger, 1901

ColiasMelinos v. herzi Staudinger, 1901: 16. Lec-totype designated by: Grieshuber, 2006b: 79.

Taxonomic status and position: uncertain; the last revisional papers about C. tyche (Böber, 1812) [Grieshuber, 1998a, 1998b, 1998c] made by using only external features, not even one type specimen have been dissected, no genitalia figures have been published; also no DNA data have been sequenced and used. According to these facts I can conclude that all "revi-sional" data regarding this extremely variable species are useless; it is strongly needed to revise C. tyche by using both genital armatures of type specimens and DNA sequences of their fresh topotypes to resolve the real situation within its subspecific composition.

J holmbomi Bang-Haas, 1916

Colias sulitelma ab. holmbomi Bang-Haas, 1916: 194; Taf. 10, fig. 6.

Taxonomic status and position: an infrasubspe-cific taxon.

Syntypes 1S, 1$: 1S: printed, white paper 'ex coll.

hyrcanica Reissinger, 1989

Colias alfacariensis hyrcanica Reissinger, 1989: 131 - 137; Taf. 34, Fign. 1 - 8; Taf. 35, Fign. 1 - 9; Taf. 35a, Fign. 6 - 9; Taf. 36, Fign. 1 - 9; Taf. 39, fign. 1, 2; Taf. 40, Fign. 1, 2, 4; Taf. 47, Fign. 1, 2, 4; Taf. 76, Fig. 7; Taf. 77, Fig. 7.

Taxonomic status and position: according to the opinion of J.Grieshuber et al. [2011] it is a junior subjective synonym of C. alfacariensis alfacariensis Ribbe, 1905.

kenteana Heyne, [1895]

Colias aurora var. Kenteana Heyne, [1895]: 731.

Taxonomic status and position: junior subjective synonym of C. heos heos (Herbst, 1792).

Note. J.Grieshuber et al. [2011] shown that the original type specimens of kenteana were impossible to identify; only one specimen of uncertain status was found in O.Staudinger's collection. To resolve this situation (because "impossible to identify" = "lost") I designate the neotype of kenteana below.

J kostlani Strand, 1911

Colias electo L. ab. Kostlani Strand, 1911: 140.

Taxonomic status and position: an infrasubspecific taxon, thus unavailable.

lada Grum-Grshimailo, 1891

Colias Lada Grum-Grshimailo, 1891: 447 - 448.

Taxonomic status and position: a good species.

Lectotype S (col. pl. X: 31-33), designated here according to the Art. 61.1 and 74.1 of the Code: 1S: printed, numbers handwritten, white paper 'ex coll. 1/2 | Staudinger'; printed, red paper 'Syntype | Colias lada Grum- | Grshimailo, 1891, | Horae Soc. ent. | ross. 25: 447-448, | det. J. Grieshuber & | S. Churkin, 2004'; handwritten, white paper '45.'; handwritten, white paper 'Lectotype S | lada Gr.Gr. | S.Korb des. | 2014'; handwritten, brown paper 'Thibet | Sinin Alp | 90 Gr.Gr.'; handwritten, white paper '22.V.90'; handwritten, white paper 'Мыньдaньша <Myndansha>'.

Paralectotype 9: printed, numbers handwritten, white paper 'ex coll. 2/2 | Staudinger'; printed, red paper 'Syntype | Colias lada Grum- | Grshimailo, 1891, | Horae Soc. ent. | ross. 25: 447-448, | det. J. Grieshuber & | S. Churkin, 2004'; handwritten, white paper '30V90'; handwritten, handwritten, white paper 'Мынъдaньша <Myndansha>'; brown circle.

J languidus Bang-Haas, 1927

Colias eogene eogene Feld. ab. languidus BangHaas, 1927: 42, 112; t. 6, fig. 20.

Taxonomic status and position: an infrasubspecific taxon, thus unavailable.

Colias Palaeno L. var. Lapponica Staudinger, 1861:

3. Lectotype designated by: Grieshuber et al., 2011: 140.

Taxonomic status and position: junior subjective synonym of C. palaenopalaeno (Linnaeus, 1761).

libanotica Lederer, 1858

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Colias libanotica Lederer, 1858: 140 - 141; pl. 2, figs. 1, 2.

Taxonomic status and position: a good subspecies of C. aurorina Herrich-Schäffer, 1850.

Lectotype 9 (col. pl. XI: 34-36), designated here according to the Art. 61.1 and 74.1 of the Code: 19: printed, white paper, handwritten numbers 'ex coll. 1/1 | Staudinger'; printed, white paper 'Coll. Led.'; handwritten, white paper 'Libanon'; printed, pink paper 'Origin.'; handwritten, white paper 'Lectotype 9 | libanotica Led. | S.Korb des. 2014'; printed, red paper 'Syntype | Colias libanotica | Lederer, 1858 | det. Grieshuber, 2007'.

J maculata Röber, 1909

[Colias romanovi] f. maculata Röber, 1909: 66.

Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

maja Grum-Grshimailo, 1891

Colias Cocandica var. Maja Grum-Grshimailo, 1891: 447. Lectotype designated by: Grieshuber, Churkin, 2003: 257.

Taxonomic status and position: a good subspecies of C. cocandica Erschoff, 1874.

Colias Marco-Polo Grum-Grshimailo, 1888: 304 - 305.

Taxonomic status and position: a good species.

Lectotype 3 (col. pl. XI: 37-39), designated here according to the Art. 61.1 and 74.1 of the Code: 13: printed, numbers handwritten, white paper 'ex coll. 2/8 | Staudinger'; handwritten, white paper 'Lectotype 3 | marcopolo Gr.Gr. | S.Korb des. | 2014'; brown circle; handwritten, white paper 'Hindukusch or. | 87 Gr.Gr.'; printed, red paper 'Syntype | Colias marcopolo Grum- | Grshimailo, 1888 | det. Grieshuber, 2007'; handwritten, white paper 'EeHKb <Beik Pass> | 16.7.87'.

Paralectotypes 23, 59: 13: printed, numbers handwritten, white paper 'ex coll. 1/8 | Staudinger'; handwritten, brown paper 'Hindukusch or. | 87 Gr.Gr.'; printed, red paper 'Syntype | Colias marcopolo Grum- | Grshimailo, 1888 | det. Grieshuber, 2007'; handwritten, white paper 'Marco Polo | Gr.Gr.'; 9: printed, numbers handwritten, white paper 'ex coll. 3/8 | Staudinger'; brown circle; handwritten, white paper 'Hindukusch or. | 87 Gr.Gr.'; printed, red paper 'Syntype | Colias marcopolo Grum- | Grshimailo, 1888 | det. Grieshuber, 2007'; 13: printed, numbers handwritten, white paper 'ex coll. 4/8 | Staudinger'; brown circle; handwritten, white paper 'Hindukusch or. | 87 Gr.Gr.'; printed, red paper 'Syntype | Colias marcopolo Grum- | Grshimailo, 1888 | det. Grieshuber, 2007'; 19: printed, numbers handwritten, white paper 'ex coll. 5/8 | Staudinger'; brown circle; handwritten, white paper 'Hindukusch or. | 87 Gr.Gr.'; printed, red paper 'Syntype | Colias marcopolo Grum- | Grshimailo, 1888 | det. Grieshu-ber, 2007'; handwritten, white paper 'EeHKb <Beik Pass> | 14.7.87'; 19: printed, numbers handwritten, white paper 'ex coll. 6/8 | Staudinger'; brown circle; handwritten, white paper 'Hindukusch or. | 87 Gr.Gr.'; printed, red paper 'Syntype | Colias marcopolo Grum- | Grshimailo, 1888 | det. Grieshuber, 2007'; 19: printed, numbers handwritten, white paper 'ex coll. 7/8 | Staudinger'; brown circle; handwritten, white paper 'Hindukusch or. | 87 Gr.Gr.'; printed, red paper 'Syntype | Colias marcopolo Grum- | Grshimailo, 1888 | det. Grieshuber, 2007'; 19: printed, numbers handwritten, white paper 'ex coll. 8/8 | Staudinger'; brown circle; handwritten, white paper 'Hindukusch or. | 87 Gr.Gr.'; printed, red paper 'Syntype | Colias marcopolo Grum- | Grshimailo, 1888 | det. Grieshuber, 2007'.

maureri Staudinger, 1901

Colias Staudingeri v. Maureri Staudinger, 1901: 18. Lectotype designated by: Grieshuber et al., 2011: 150.

Taxonomic status and position: junior subjective synonym of C. staudingeri Alpheraky, 1881.

Lectotype 3: printed, pink paper 'Origin.'; print-

J meridensis Neuburger, 1905

Colias dimera Doubl. Hew. ab. meridensis Neuburger, 1905: 42.

Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

J niediecki Strand, 1909

C.[olias] croceus Fourcr. (edusa F.) (ab. niediecki) Strand, 1909: 78.

Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

ten, with printed "Strand det.", white paper 'Col. croceus | ab. Nied- | eckii m. | Strand det.'.

niveata O.Bang-Haas, 1927

Colias cocandica niveata Verity, O.Bang-Haas, 1927: 110.

Taxonomic status and position: junior objective synonym of C. nebulosa nebulosa Oberthür, 1894.

orientalis Staudinger, 1892

Colias Palaeno var. Orientalis Staudinger, 1892b: 311 - 312.

Taxonomic status and position: a good subspecies of C. palaeno (Linnaeus, 1761).

Lectotype S (col. pl. XI: 40-42), designated here according to the Art. 61.1 and 74.1 of the Code: 1S: printed, numbers handwritten, white paper 'ex coll 1/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Kentei] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'var. Orientalis | Stgr.'; handwritten, green paper 'Kentei | 89 Dörr.'; handwritten, white paper 'Lectotype S | orientalis Stgr | S.Korb des. 2014'.

Paralectotypes 8 S, 9 9: 19: printed, numbers handwritten, white paper 'ex coll 2/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Kentei] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Kentei | 89 Dörr.'; green circle; 1S: printed, numbers handwritten, white paper 'ex coll 3/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Kentei] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Kentei | 89 Dörr.'; 19: printed, numbers handwritten, white paper 'ex coll 4/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Kentei] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Kentei | 89 Dörr.'; green circle; 1S: printed, numbers handwritten, white paper 'ex coll 5/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Kentei] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Kentei | 89 Dörr.'; green circle; 19: printed, numbers handwritten, white paper 'ex coll 6/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Kentei] | Syntype | Colias palaeno [var.]

orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Kentei | 89 Dörr.'; green circle; 1$: printed, numbers handwritten, white paper 'ex coll 7/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Kentei] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Kentei | 89 Dörr.'; green circle; 1$: printed, numbers handwritten, white paper 'ex coll 8/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Kentei] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Kentei | 89 Dörr.'; green circle; 1$: printed, numbers handwritten, white paper 'ex coll 9/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Ussuri] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, blue paper 'Ussuri | Dörr.'; 1$: printed, numbers handwritten, white paper 'ex coll 10/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Ussuri] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Ussuri | Dörr.'; blue circle; 1$: printed, numbers handwritten, white paper 'ex coll 11/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Ussuri] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Ussuri | Dörr.'; 1$: printed, numbers handwritten, white paper 'ex coll 12/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Ussuri] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Ussuri | Dörr.'; blue circle; 1$: printed, numbers handwritten, white paper 'ex coll 13/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Ussuri] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Ussuri | Dörr.'; blue circle; 1 $: printed, numbers handwritten, white paper 'ex coll 14/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Ussuri] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Ussuri | Dörr.'; 1$: printed, numbers handwritten, white paper 'ex coll 15/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Ussuri] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, white paper 'Ussuri'; blue circle; handwritten, white paper 'v. Orientalis'; 1$: printed, numbers handwritten, white paper 'ex coll 16/18 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper '[Bikin] | Syntype | Colias palaeno [var.] orientalis | Staudinger, 1892 | det. Grieshuber, 2007'; blue circle; handwritten, blue paper 'Bikin | 87 Dörr.'; green rectangle; 1$:

orthocalida Reissinger, 1974 Colias alfacariensis orthocalida Reissinger, 1974: 1 - 7; pl. 7, figs. 1 - 4; pl. 8, figs. 1 - 22.

Taxonomic status and position: junior subjective synonym of C. alfacariensis alfacariensis Ribbe, 1905.

J pallida Staudinger, 1861 [Colias erate] ab. Pallida Staudinger, 1861: 3. Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

Syntype 9: printed, numbers handwritten, white paper 'ex coll. 1/1 | Staudinger'; printed, pink paper 'Origin.'; handwritten, green paper 'Sarepta'; handwritten, white paper 'Pallida'; printed, white paper 'infrasubspecific taxon | name bearing specimen of Colias | erate [ab.] pallida Staudinger, | 1861 | det. Grieshuber, 2007'.

pamira Groum-Grshimailo, 1890 Colias Staudingeri Alph. var. Pamira Groum-Grshimailo, 1890: 334 - 335; pl. 4, fig. 2.

Taxonomic status and position: a good subspecies of C. staudingeri Alpheraky, 1881.

Syntype 3: printed, numbers handwritten, white paper 'ex coll 1/1 | Staudinger'; handwritten, brown paper 'Zalai Pamir | 85. Grumm.'; printed, red paper 'Syntype | Colias staudingeri [var.] pamira | Grum-Grshimailo, 1890 | det. Grieshuber, 2007'; handwritten, white paper '20.VII.84.'; handwritten, white pa-

per 'Kapa <Kara> | 04.'.

J parva Hoyningen-Huene, 1901

Colias palaeno ab. parva Hoyningen-Huene, 1901: 460.

Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

poktussani O.Bang-Haas, 1935

Colias palaeno poktussani O.Bang-Haas, 1935: 183.

Taxonomic status and position: a good subspecies of C. palaeno (Linnaeus, 1761).

richthofeni Bang-Haas, 1927

Colias richthofeni Bang-Haas, 1927: 42; Taf. 6, Fign. 1 - 3.

Taxonomic status and position: a good subspecies of C. nebulosa Oberthür, 1894.

romanovi Grumm-Grshimailo, 1885

C.[olias] romanovi Grumm-Grshimailo, 1885: 229.

Taxonomic status and position: a good species. J.Grieshuber et al. [2011: 190]: "seems to be very closely related to C. erschoffi... Much further investigation is needed before this problem can be resolved". I did these investigations: dissected males of both species [see: Korb, 2012: figs. 91, 97]. These taxa have very good differences in male genitalia (in romanovi aedoeagus is equal to valva in length, in erschoffi aedoeagus at least 30 % longer than valva; other small differences are also present).

Lectotype 3 (col. pl. XI: 46-48), designated here according to the Art. 61.1 and 74.1 of the Code: 13: printed, numbers handwritten, white paper 'ex coll. 4/7 | Staudinger'; printed, red paper 'Syntype | Colias romanovi Grum- | Grshimailo, 1885 | det. Grieshuber, 2007'; brown circle; handwritten, white paper 'Lectotype 3 | romanovi Gr.Gr. | S.Korb des. | 2014'; handwritten, white paper 'S.W.Alai | 85 Grum.'; handwritten, white paper '27.VI.84'; handwritten, white paper 'Eang^yaH <Baldshuan>'.

printed, numbers handwritten, white paper 'ex coll. 2/7 | Staudinger'; printed, red paper 'Syntype | Colias romanovi Grum- | Grshimailo, 1885 | det. Grieshuber, 2007'; handwritten, brown paper 'S.W.Alai | 85 Grum.'; 1$: printed, numbers handwritten, white paper 'ex coll. 5/7 | Staudinger'; printed, red paper 'Syntype | Colias romanovi Grum- | Grshimailo, 1885 | det. Grieshuber, 2007'; brown circle; handwritten, white paper 'S.W.Alai | 85 Grum.'; 1$: printed, numbers handwritten, white paper 'ex coll. 6/7 | Staudinger'; printed, red paper 'Syntype | Colias romanovi Grum- | Grshimailo, 1885 | det. Grieshuber, 2007'; brown circle; handwritten, white paper 'S.W.Alai | 85 Grum.'; 1$: printed, numbers handwritten, white paper 'ex coll. 7/7 | Staudinger'; printed, red paper 'Syntype | Colias romanovi Grum- | Grshimailo, 1885 | det. Grieshuber, 2007'; handwritten, brown paper 'Zalai | Grum.'; handwritten, white paper '1'; handwritten, white paper '20.VII.84.'; handwritten, white paper 'Кара <Kara> | 84'; 1$: printed, red paper 'Syntype | Colias romanovi Grum- | Grshimailo, 1885 | det. Grieshuber, 2007'; brown circle; handwritten, white paper 'S.W.Alai | 85 Grum.'.

roschana Grum-Grshimailo, 1893 Colias Alpherakii var. Roschana Grum-Grshimailo, 1893: 381.

Taxonomic status and position: junior subjective synonym of C. alpherakii Staudinger, 1882.

rueckbeili Lukhtanov, 1994 Colias wiskotti rueckbeili Lukhtanov, Lukhtanov, 1994: 74.

Taxonomic status and position: a good subspecies of C. wiskotti Staudinger, 1882.

Syntype <: printed, white paper, numbers handwritten 'ex coll. 1/1 | Staudinger'; handwritten, white paper 'var. Alexandra | Stgr. in l.'; printed, red paper 'Syntype | Colias wiskotti | rueckbeili Lukhtanov, 1994 | nec. "Alexandra" Röber, | 1907 (unavailable) | nec. "rückbeili" Bang- | Haas, 1927 (unavail.) | Grieshuber det. XII.2004'; handwritten, white paper 'v. Chrysop-tera Gr.Gr. | falsch | wohl= v. Draconis Gr.Gr.'; handwritten, brown paper 'Alex. Geb. | 99. Rckbl.'. sagartia Lederer, 1869 Colias Sagartia Lederer, 1869: 75 - 76; pl. 4, figs. 1, 2. Taxonomic status and position: a good species. Lectotype S (col. pl. XII: 49-51), designated here

according to the Art. 61.1 and 74.1 of the Code: 1<: printed, white paper, numbers handwritten 'ex coll. 2/5 | Staudinger'; printed, pink paper 'Origin.'; handwritten, white paper 'Hadschyabad'; printed, red paper '[2/5] | Syntype | Colias sagartia Lederer, 1869 | det. Grieshuber, 2007'; printed, white paper 'Coll. Led.'; handwritten, white paper 'Lectotype S | sagartia Led. | S.Korb des. 2014'.

Paralectotypes 4$: 1$: printed, white paper, numbers handwritten 'ex coll. 4/5 | Staudinger'; printed, pink paper 'Origin.'; handwritten, white paper 'Hadschyabad'; printed, red paper '[4/5] | Syntype | Colias sagartia Lederer, 1869 | det. Grieshuber, 2007'; 1$: printed, white paper, numbers handwritten 'ex coll. 5/5 | Staudinger'; printed, pink paper 'Origin.'; printed, white paper 'Coll. Led.'; handwritten, white paper 'Hadschyabad'; printed, red paper '[5/5] | Syntype | Colias sagartia Lederer, 1869 | det. Grieshuber, 2007'; 1 $: printed, white paper, numbers handwritten 'ex coll. 3/5 | Staudinger'; printed, pink paper 'Origin.'; printed, white paper 'Coll. Led.'; handwritten, white paper 'Hadschyabad'; printed, red paper '[3/5] | Syntype | Colias sagartia Lederer, 1869 | det. Grieshuber, 2007'; 1$: printed, white paper, numbers handwritten 'ex coll. 1/5 | Staudinger'; printed, pink paper 'Origin.'; printed, white paper 'Coll. Led.'; handwritten, white paper 'Hadschyabad'; printed, red paper '[1/5] | Syntype | Colias sagartia Lederer, 1869 | det. Grieshuber, 2007'.

sareptensis Alpheraky, 1875

Colias Hyale L. var. Sareptensis Alpheraky, 1875: 153 - 154 (raising status of taxon [Colias] Hyale ab. (an hibr?) Sareptensis Staudinger, 1871).

Taxonomic status and position: junior subjective synonym of C. erate (Esper, [1805]). The separation of taxa sareptensis Alpheraky and sareptensis Staudinger by J.Grieshuber et al. [2011] was based on the wrong assumption that S.Alpheraky described his own subspecies basing on own material; in fact, and it is clearly stated in the Russian text of S.Alpheraky's paper, he just raised status of O.Staudinger's 'aberration' clearly to 'variety'. According to this fact, lectotype designation of sareptensis Alpheraky, 1875 from S.Alpheraky's specimens [Grieshuber et al., 2006: 110; fig. 1] is invalid; it was fixed by correct lectotype designation from O.Staudinger's specimens [Korb, 2013: 250].

J saturata Röber, 1909

C.[olias]flaveola Bl. ab. saturata Röber, 1909: 93.

Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

schildei Staudinger, 1892

Colias Palaeno L. var. Schildei Staudinger, 1892b: 310.

Taxonomic status and position: junior subjective synonym of C. palaeno palaeno (Linnaeus, 1761).

Syntypes 33, 1$: 13: printed, white paper, numbers handwritten 'ex coll. 1/16 | Staudinger'; printed, pink paper 'Origin.'; handwritten, white paper 'ab. Schildei | Stgr.'; printed, white paper '[Kusamo 1/16] | infrasubspecific taxon | name bearing specimen of Colias | palaeno [var.] schildei | Staudinger, 1892 | det. Grieshuber, 2007'; handwritten, green paper 'Kusamo | Schilde'; handwritten, white paper 'Kuusamo.'; 1$: printed, white paper, numbers handwritten 'ex coll. 2/16 | Staudinger'; printed, pink paper 'Origin.'; printed, white paper '[Kusamo 2/16] | infrasubspecific tax-on | name bearing specimen of Colias | palaeno [var.] schildei | Staudinger, 1892 | det. Grieshuber, 2007'; green circle; handwritten, white paper 'Kuusamo.'; handwritten, yellow paper 'Zu Pelidne leitend'; 13: printed, white paper, numbers handwritten 'ex coll. 4/16 | Staudinger'; printed, pink paper 'Origin.'; printed, white paper '[Kusamo 4/16] | infrasubspecific tax-on | name bearing specimen of Colias | palaeno [var.] schildei | Staudinger, 1892 | det. Grieshuber, 2007'; green circle; handwritten, white paper 'Küsamo.'; 13: printed, white paper, numbers handwritten 'ex coll. 3/16 | Staudinger'; printed, pink paper 'Origin.'; printed, white paper '[Kusamo 3/16] | infrasubspecific tax-on | name bearing specimen of Colias | palaeno [var.] schildei | Staudinger, 1892 | det. Grieshuber, 2007'; green circle; handwritten, white paper 'Kuusamo.'. seitzi Bollow, 1930

C.[olias] ladakensis suisp. <sic!> seitzi Bollow, 1930: 119.

Taxonomic status and position: junior subjective synonym of C. ladakensis ladakensis Felder et Felder, 1865.

Paralectotype $: printed, white paper 'ex coll. | BANG-HAAS'; printed, red paper 'Type'; handwritten, white paper ' C. ladakensis | subsp. seitzi | Type Bollow'; printed, red paper 'Syntypus | Colias

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sieversi Grum-Grshimailo, 1887 Colias Sieversi Grum-Grshimailo, 1887: 388, 397 - 398. Taxonomic status and position: a good species. Lectotype S (col. pl. XII: 55-57), designated here according to the Art. 61.1 and 74.1 of the Code: 1<: numbers handwritten, white paper 'ex coll. 1/6 | Staudinger'; labelled: printed, red paper 'Syntype | Colias sieversi Grum-Grshimailo, 1887 | det. Grieshuber, 2007'; handwritten, brown paper 'Darwas | 85 Grumm.'; handwritten, white paper '013/2014K'; handwritten, white paper 'Lectotype S | sieversi Gr. Gr. | S.Korb des. /14'; handwritten, white paper '1.VII | 85'; handwritten, white paper 'Бездани <Bezdani>'.

Paralectotypes 3<, 2$: 1<: printed, numbers handwritten, white paper 'ex coll. 2/6 | Staudinger'; brown circle; printed, red paper 'Syntype | Colias sieversi Grum-Grshimailo, 1887 | det. Grieshuber, 2007'; handwritten, white paper 'Darwas | 85 Grumm.'; 1 S: printed, numbers handwritten, white paper 'ex coll. 3/6 | Staudinger'; handwritten, white paper '27/ VI.85'; brown circle; handwritten, white paper 'Jn Yrizeks.'; printed, red paper 'Syntype | Colias sieversi Grum-Grshimailo, 1887 | det. Grieshuber, 2007'; handwritten, white paper 'Darwas | 85 Grumm.'; 1$: printed, numbers handwritten, white paper 'ex coll. 4/6 | Staudinger'; brown circle; printed, red paper 'Syn-type | Colias sieversi Grum-Grshimailo, 1887 | det. Grieshuber, 2007'; handwritten, white paper 'Darwas | 85 Grumm.'; 1<: printed, numbers handwritten, white paper 'ex coll. 6/6 | Staudinger'; brown circle; printed, red paper 'Syntype | Colias sieversi Grum-Grshimailo, 1887 | det. Grieshuber, 2007'; handwritten, white paper 'Darwas | 85 Grumm.'; 1$: brown circle; printed, red paper 'Syntype | Colias sieversi Grum-Grshimailo, 1887 | det. Grieshuber, 2007'; handwritten, white paper 'Darwas | 85 Grumm.'.

sifanica Grum-Grshimailo, 1891 Colias Sifanica Grum-Grshimailo, 1891: 447. Taxonomic status and position: a good species. Lectotype S (col. pl. XII: 58-60), designated here according to the Art. 61.1 and 74.1 of the Code: 1S: printed, white paper 'ex coll. 1/2 | Staudinger'; printed, red paper 'Syntype | Colias sifanica Grum-| Grshimailo, 1891 | det. Grieshuber, 2007'; handwritten, white paper 'Lectotype S | sifanica Gr.Gr. | S.Korb des. 2014'; handwritten, brown paper 'Thibet | Sinin Alp | 90. Gr.Gr.'.

ica Gr.Gr. | S.Korb des. 2014'.

J standfussi Röber, 1909

C.[olias] pelidne Bdv. form standfussi Röber, 1909: 91.

Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

C.[olias] Staudingeri Alphéraky, 1881: 368 - 373; pl. 14, figs. 3 - 4. Lectotype designated by: Grieshu-ber et al., 2004: 38 - 39; pl. 2, fig. 1.

Taxonomic status and position: a good species.

J strandi Röber, 1924

Colias strandi Röber, 1924: 90.

Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

tamerlana Staudinger, 1897

Colias Tamerlana Staudinger, 1897: 152 - 153; pl. 5, figs. 1, 3. Lectotype designated by: Korb, 2012: 28; pl. 4, figs. 10, 11.

Taxonomic status and position: a good subspecies of C. cocandica Erschoff, 1874; it has no priority over the name mongola Alpheraky, 1897 because the last name was published two months earlier than the name tamerlana [Gorbunov, 2012].

Lectotype 3: printed, numbers handwritten, white paper 'ex coll. 2/12 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias tamerlana Staudinger, | 1897 | det. Grieshuber, 2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle; microvial with genitalia preparate in glycerin, with glued label 'Studied by S.Korb | ZMHB | 0096K/2011'; handwritten, white paper 'Lectotype | tamerlana Stgr. | S.Korb des.'.

2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle; 1$: printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias tamerlana Staudinger, | 1897 | det. Grieshuber, 2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle; 13: printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias tamerlana Staudinger, | 1897 | det. Grieshuber, 2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle; handwritten (two lines black ink, one line pencil), white paper 'v.? Tamerlana | Stgr. | Thian. or.'; 13: printed, numbers handwritten, white paper 'ex coll. 6/12 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias tamerlana Staudinger, | 1897 | det. Grieshuber, 2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle; 1$: printed, numbers handwritten, white paper 'ex coll. 10/12 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias tamerlana Staudinger, | 1897 | det. Grieshuber, 2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle; 1$: printed, numbers handwritten, white paper 'ex coll. 12/12 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias tamerlana Staudinger, | 1897 | det. Grieshuber, 2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle; 1$: printed, numbers handwritten, white paper 'ex coll. 11/12 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias tamerlana Staudinger, | 1897 | det. Grieshuber, 2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle; 13: printed, numbers handwritten, white paper 'ex coll. 9/12 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias tamerlana Staudinger, | 1897 | det. Grieshuber, 2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle; 13: printed, numbers handwritten, white paper 'ex coll. 7/12 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias tamerlana Staudinger, | 1897 | det. Grieshuber, 2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle; 1$: printed, numbers handwritten, white paper 'ex coll. 4/12 | Staudinger'; printed, pink paper 'Origin.'; printed, red paper 'Syntype | Colias tamerlana Staudinger, | 1897 | det. Grieshuber, 2007'; handwritten, white paper 'Thian. or. | 96 Hbhr.'; brown circle. tatarica O.Bang-Haas, 1915

Colias cocandica tatarica O.Bang-Haas, 1915: 98 - 99. Lectotype designated by: Grieshuber, Churkin, 2003: 260 - 261; pl. 15, fig. 7.

Taxonomic status and position: a good subspecies of C. cocandica Erschoff, 1874.

Paralectotypes 23, 1$: 13: printed, white paper 'ex coll. BANG-HAAS'; printed, yellow paper 'GART | Exemplar und Eti- | ketten dokumentiert | specimen and label | data documented | 22.11.R0 2001 .'; printed, pink paper 'Origin.'; printed, white paper 'Schahidulla

Colias aurora thia O.Bang-Haas, 1934: 109 - 110.

Taxonomic status and position: junior subjective synonym of C. diva vespera O.Bang-Haas, 1927 [Grieshuber, 1996a].

tkatschukovi O.Bang-Haas, 1936

Colias chlorocoma tkatschukovi O.Bang-Haas, 1936: 287. Lectotype designated by: Nekrutenko, 1990: 102.

Taxonomic status and position: junior subjective synonym of nominotypical C. chlorocoma Christoph, 1888.

transcaspica Christoph, 1887

Colias Aurorina var. Transcaspica Christoph, 1887: 162.

Taxonomic status and position: a good subspecies of C. aurorina Herrich-Schäffer, 1850.

Lectotype 3 (col. pl. XII: 64-66), designated here according to the Art. 61.1 and 74.1 of the Code: 13: printed, numbers handwritten, white paper 'ex coll. 3/4 | Staudinger'; printed, red paper 'Syntype | Colias aurorina | [var.] transcaspica | Christoph, 1887 | det. Grieshuber, 2007'; handwritten, white paper 'Lectotype | transcaspica Chr. | S.Korb des. 2014'; handwritten, white paper '14 6 82 | Нохуръ <Nokhur>'.

tsurpuana Grieshuber, 1996

Colias nina tsurpuana Grieshuber, 1996b: 557 -558; pl. 1, figs. 1 - 9.

Taxonomic status and position: junior subjective synonym of C. nina Fawcett, 1904 [Grieshuber et al., 2011].

vespera O.Bang-Haas, 1927

Colias aurora vespera O.Bang-Haas, 1927: 44; pl. 6, figs. 14, 15.

Taxonomic status and position: junior subjective synonym of C. diva Grum-Grshimailo, 1891.

abgebildet | t. 6 f. 15 | beschrieb. p. 44'.

viridis Bollow, 1930

C.[olias] tyche Form viridis Bollow, 1930: 108; pl. 7, row e.

Taxonomic status and position: junior subjective synonym of nominotypical C. montium Oberthür, 1886.

wanda Grum-Grshimailo, 1907

Colias eogene wanda Grum-Grshimailo, 1907: 13, 363, 424. Lectotype designated by: Grieshuber, Churkin, 2003: 250 - 251.

Taxonomic status and position: a good species.

J werneri Geest, 1905

Colias Chrysotheme Esp. ab. Werneri Geest, 1905: 379.

Taxonomic status and position: an infrasubspe-cific taxon, thus unavailable.

Syntypes, 23, 1$: 1$: printed, white paper 'ex coll. | BANG-HAAS'; printed, yellow paper 'GART

wiskotti Staudinger, 1882

Colias Wiskotti Staudinger, Bang-Haas, 1882: 166 -167; Taf. 2, Fig. 9 - 10. Lectotype designated by: Grie-shuber, Worthy, 2006: 48.

Taxonomic status and position: a good species.

ACKNOWLEDGMENTS

I am deeply grateful to the curator of the lepidop-terological collections of the Museum für Naturkunde an der Humboldt-Universität zu Berlin Dr. W.Mey for the opportunity to study the collections.

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