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15http://zoobank.org/urn:lsid:zoobank.org:pub:6AF64174-31C7-4F70-A83E-27512309CEAB
DOI: 10.24412/2226-0773-2023-12-5-468-491
EDN: JFGHVH
Taxonomy structure of Agapanthia (Epoptes) asphodeli (Latreille, 1804) (Coleoptera, Cerambycidae)
M.A. Lazarev
Free Economic Society of Russia, Department of Scientifics Conferences and
All-Russian Projects
Tverskaya str., 22a, Moscow 125009 Russia
e-mail: cerambycidae@bk.ru, humanityspace@gmail.com
Key words: Coleoptera, Cerambycidae, taxonomy, zoogeography, new status, new subspecies, new synonyms, name restored.
Abstract: The taxonomy of the species is revised; 8 subspecies are accepted: Agapanthia (Epoptes) a. asphodeli (Latreille, 1804) - type locality France, Bordeaux environs; A. (E.) a. lopadusae Rapuzzi & Sparacio, 2017, stat. n. - type locality Italy, Sicilian Channel, Lampedusa Island; A. (E.) a. renatae Steiner & Schmid, 2013, stat. n. - type locality Greece, Peloponnese, Arcadia, between Karitena and Kastro; A. (E.) a. cretica Bernhauer, 1978, stat. n. - type locality Crete, Mount Ida, Anogia; A. (E.) a. kastamonica ssp. n. - type locality Turkey, Kastamonu province, Yaralıgöz Dağı; A. (E.) a. nicosiensis Pic, 1927 is restored to its original rank - type locality Cyprus; A. (E.) a. zappii Sama, 1987, stat. n. - type locality Algeria (Batna), Telmet canyon (Aures Mountains), 1700 m; A. (E.) a. fadli Sama & Rapuzzi, 2006, stat. n. - type locality “14 km east of Burg al ‘Arab (west of Alexandria), 30°58’N 29°40’E”. One subspecies is described as new and the species rank of five names is downgraded to subspecies level. Two new synonyms are proposed: A. (E.) renatae Steiner & Schmid, 2013 = A. (E.) asphodeli balcanica Sláma, 2019, syn. n.; A. (E.) cretica Bernhauer, 1978 = A. (E.) probsti Holzschuh, 1984, syn. n. The holotype of A. reyi Mulsant & Godart, 1870 is depicted for the first time, so the synonymies by Sama (1979) A. (E.) asphodeli = A. (E.) reyi Mulsant & Godart, 1870 is proved.
Introduction
Agapanthia asphodeli (Latreille, 1804) is a rare poorly investigated species. Many local populations were described as separate species; sometimes (Lucas, 1848; Pic, 1927) A. asphodeli was mixed with A. boeberi (Fischer von Waldheim, 1806). Spanish populations of A. asphodeli were long time accepted as A. reyi Mulsant & Godart, 1870 after wrong identification of the holotype of the later. The area of the species was treated by many authors for the regions, where no specimens were available from now (Plavilstshikov, 1948 - Arax valley; 1965 - steppe area, Crimea; 1968 - steppe area of USSR eastwards Urals and Transcaucasia; Ogloblin, 1948 - south of forest-steppe and steppe area of European USSR; Danilevsky & Miroshnikov, 1985 - European USSR, Transcaucasia; Prisnyj & Vorobieva, 2005 - Belgorod. Only one male (from Teberda) is available now from Russia.
So, the actual adequate information is necessary to be published.
I accept the species with 8 subspecies:
1. A. (E.) a. asphodeli (Latreille, 1804)
Type locality. France, Bordeaux environs.
2. A. (E.) a. lopadusae Rapuzzi & Sparacio, 2017, stat. n.
Type locality. Italy, Sicilian Channel, Lampedusa Island.
3. A. (E.) a. renatae Steiner & Schmid, 2013, stat. n.
Type locality. Greece, Peloponnese, Arcadia, between Karitena and Kastro.
4. A. (E.) a. cretica Bernhauer, 1978, stat. n.
Type locality. Crete, Mount Ida, Anogia.
5. A. (E.) a. kastamonica ssp. n.
Type locality. Turkey, Kastamonu province, Yaralıgöz Dağı.
6. A. (E.) a. nicosiensis Pic, 1927, stat. rest.
Type locality. Cyprus.
7. A. (E.) a. zappii Sama, 1987, stat. n.
Type locality. Algeria (Batna), Telmet canyon (Aures Mountains), 1700 m.
8. A. (E.) a. fadli Sama & Rapuzzi, 2006, stat. n.
Type locality. “14 km east of Burg al ‘Arab (west of Alexandria), 30°58’N 29°40’E”.
Material and method
Material was collected manually. Specimens used in morphological studies were killed by ethyl acetate. All photographs were taken with Canon PowerShot G10 digital camera equipped with Cannon Zoom lens 5X IS 6.1-30.5 mm 1:2.8-4.5 and microscope AmScope SM745NTP. The illustrations were edited with Adobe Photoshop 7.0 and Helicon Focus 3.20.
Acronyms of collections:
MD - collection of M.L. Danilevsky (Moscow)
ML - collection of M.A. Lazarev (Moscow)
SM - collection of S.V. Murzin (Moscow)
MNHN - collection of Muséum National d'Histoire Naturelle (Paris)
ZMM - collection of Zoological Museum of Moscow University
Results
Agapanthia (Epoptes) asphodeli (Latreille, 1804)
Lamia asphodeli Latreille, 1804:282 - “environs de Bordeaux”.
Saperda asphodelis, Westwood, 1839: 365.
Agapanthia cynarae var. delagrangei Pic, 1894: 75 - “Akbès”.
Agapanthia delagrangei, Plavilstshikov, 1932: 199, part. (= A. verecunda Chevrolat, 1882).
Agapanthia asphodeli m. mimica, Breuning, 1961: 184.
Agapanthia asphodeli, Plavilstshikov, 1932: 194 - south of USSR, Caucasus; 1965: 416 - steppe area, Crimea, Caucasus; Ogloblin, 1948: 469 - south of forest-steppe and steppe area of European USSR; Villiers, 1978: 435 - “Europe méridionale et centrale, Caucase, Asie Mineure, Afrique du Nord”; Sama, 1979: 508, 511; 1992: 399 (= reyi Mulsant & Godart, 1870); 2003: 93 - Europe, North Africa, Asia Minor, Caucasus, Transcaucasia; Baragaño Galan et al., 1981: 161; Kasatkin & Arzanov, 1997: 67 - Anapa; Jeniš, 2001: 260; Prisnyj & Vorobieva, 2005: 40 - Belgorod.
Agapanthia (s. str.) asphodeli, Plavilstshikov, 1948: 167 - Arax valley; Caucasus; 1968: 143 - steppe area of USSR eastwards Urals, Caucasus and Transсaucasia; Danilevsky & Miroshnikov, 1985: 390 - European USSR, Caucasus, Transcaucasia.
Agapanthia (Agapanthiella) asphodeli, Pesarini & Sabbadini, 2004: 12; Özdikmen 2007: 347 - Europe (Portugal, Spain, France, Corsica, Italy, Sicily, Sardinia, Malta, Croatia, Bosnia & Herzegovina, Macedonia, Albania, Greece, Romania, Switzerland, ?Sweden, Ukraine, Crimea, ? Moldavia, European Russia), Kazakhstan, Caucasus, Transcaucasia, Central Asia, Turkey, ?Syria, ?Palestine, ?North Africa, (Turkey: Adana-Ankara-Antalya-Aydin-Bilecik- Çanakkale-Hatay-Isparta-Izmir-Yozgat Provices); 2008a: 77 - Turkey; 2008b: 394 - Turkey.
Agapanthia (Epoptes) asphodeli, Löbl & Smetana, 2010: 215 - Azerbaijan, Albania, Armenia, Bosnia-Herzegovina, Bulgaria, Croatia, France, Georgia, Greece, Italy, Malta, Macedonia, Moldavia, Portugal, Spain, Russia: South European Territory, Slovenia, Ukraine, Kazakhstan, Turkey; Özdikmen, 2013: 16 - “Turkey: (W half of Anatolia)”; Rapuzzi & Sparacio, 2017: 957 - “from Spain and Portugal to Southern Russia, West Kazakhstan, Balcan Peninsula and Asia Minor (locus typicus: Bordeaux, France)”; Kasatkin, 2020: 242, 243; Danilevsky, 2023: 592.
Agapanthia (Epoptes) asphodeli asphodeli Danilevsky, 2020: 301 - E: Albania, Bosnia-Herzegovina, Bulgaria, Croatia, France, Greece, Italy, Malta, Macedonia, ?Moldavia, Portugal, Romania, Spain, ?Russia: South European Territory, Slovenia, ?Ukraine, N: Morocco ?A: Azerbaijan, Armenia, Georgia, Kazakhstan, Turkey; Özdikmen, 2021: 1351 E: Albania, Bosnia-Herzegovina, Bulgaria, Croatia, France, Greece, Italy, Malta, Macedonia, ?Moldavia, Portugal, Romania, Spain, ?Russia: South European Territory, Slovenia, ?Ukraine, N: Morocco, ?A: Azerbaijan, Armenia, Georgia, Kazakhstan, Turkey.
Type locality. France, Bordeaux environs.
Description. Relatively big species with moderately narrow body, covered with more or less dense yellow pubescence; eyes small, with lower lobes shorter than length of genae or rather big, about two times longer, than genae; antennae usually thin, very long, in males about two times longer than body, surpassing elytral apex by 6th joint; in females - by 8th joint; most of basal parts of 3rd - 12th reddish or dark-brown; apical parts black; 3rd joint always without setae tufts, sometimes with a poor concentration of several long setae; prothorax transverse, angulate widened at middle; pronotum with dense and wide central yellow setae line, without recumbent pubescence along its sides, with very numerous long erect setae; scutellum semicircular with very dense recumbent yellow pubescence; elytra about 2.8-3.0 times longer than basal width, with moderately sharpened apices; poorly spotted elytral pubescence rathr sparse, so elytral shine can be distinct; erect setae are distributed from elytral base to the apex; elytral punctation very dense with interspaces less than each dot; grey humeral line can be distinct in certain subspecies; 3rd joint of anterior tarsi elongated; 1st and 2nd joints together of posterior legs longer than 3rd; ventral body side with very dense yellow pubescence; last abdominal tergites in males truncated, sternites - slightly emarginates; last abdominal tergites and sternites in females rounded; body length in males: 10.5-17.2 mm, body length in females: 13.8-23.0 mm.
Distribution. The species is distributed in Europe from Portugal along south Europe to Slovakia, Hungary, Greece, Bulgaria, and Turkey; it is also known from Ukraine and Moldavia, widely distributed in Anatolia and North Africa (Egypt, Morocco, Lybia, Algeria, Tunisia). The species was also recorded from Syria. All numerous records from Russia, Transcaucasia and Kazakhstan must be proved, as a single old male only is available from North Caucasus (Teberda), and no specimens are available from Transcaucasia and Kazakhstan. The records from Iran (Sakenin, 2011: 6 - “Ardabil province: Meshkinshahr”) are doubtful.
Biology. Larvae are connected with stems of Asphodelus; Thapsia and Ferula were also recorded; all records of Carduaceae are very doubtful; imagoes are active from April to June.
Many Agapanthia species close to A. asphodeli (Latreille, 1804) were listed by Rapuzzi & Sparacio (2017), but most of them are in fact good subspecies of A. asphodeli (Latreille, 1804).
Agapanthia (Epoptes) asphodeli asphodeli (Latreille, 1804)
Figs 1-3, 7
Lamia asphodeli Latreille, 1804:282 - “environs de Bordeaux”.
Saperda spencii Gyllenhal, 1817: 187 - “Gallia meridionali”.
Agapanthia asphodeli, Mulsant 1839: 174 - “midi de la France”; 1862: 355 - “espèce méridionale”; Küster, 1846: 66 - “Im südlichen Frankreich”; Chernay, 1854: 252 - Kharkov; Gemminger & Harold, 1873: 3175 - “Gallia mer., Hungaria”; Perris, 1877: 340 - “Hyères”; Ganglbauer, 1884: 541, part. - “Süd-Europa, Kleinasien”; Seidlitz, 1891: 849 - “Im südl. Eur.”; Reitter, 1898: 130 - “Südeuropa, Kleinasien”; 1913: 66 - “in Böhmen”; Winkler, 1929: 1213 - Bohemia, Transsylvania, Mediterranea; Bonnamour, 1935: 107 - “Dans l'Isère: Vienne, la Grande-Chartreuse”; Schaefer 1947 - France: Banyuls -sur-Mer; Breuning, 1961: 184, part. - “Eur. centr. et mer., As. mer. occ.”; Carrière, 1996a: 64 - “Hérault”; 1996b: 562 - “Aude, Hérault”; 1996c: 110, 111 - “Les Baumes, Causse de Neffiès; La Redoute-plage; Bois Mégé, Vissous”; 2001: 404 - “Hérault”; 2003: 257, 258 - “Causse de Neffiès (Hérault)”; 2005: 468 - “Causse de Fauzan”; 2007: 551 - “Pic de Vissous, Grange de Couderc, Pech Haut, Roque-Nègre, Cabrières (Montagne des Batailles)”; Verdugo-Páez, 1999: 19 - “San Fernando”, “Puerto Real”, “Sierra Grazalema”, “Alcalá de los Gazules”; Sudre et al., 1999: 171 - “Arboras”, “pic Saint-Loup”, “massif de la Gardiole”, “Murviel-lès-Montpellier”, “Aumelas”, “Saint-Georges-d'Orques”, “Cournonterral”, “Saint-Guilhem-le-Désert”; Marquet, 2001: 116 - “Parc naturel régional de la Brenne (Indre)”; Mifsud, 2002: 167 - “Malta”; Sheshurak & Sadovnicha, 2002: 242 - Chernihiv Region; Echevarría Mayo, 2003: 218 - “Ávila”; Valladares et al., 2003: 79 - “Province d'Almeria”; Brustel et al., 2003: 452 - “Villeneuve-les-Maguelonne (Hérault)”; Neculiseanu & Baban, 2005: 202 - “Moldova”; Diego Barquín & Martínez-Porres Cáceres, 2005: 145 - “Cantabria”; Peris-Felipo et al., 2008: 109 - “Parque Natural de La Tinença de Benifassà (Castellón)”; 2010: 362 - “Torrevieja (Alicante)”, “Parque Natural de Las Lagunas de La Mata-Torrevieja (Alicante, España)” - (N38º01’56.39’’ W000º42’47.80’’, N38º01’59.38’’ W000º42’44.91’’, N38º01’49.90’’ W000º42’00.13’’, N38º01’19.08’’ W000º39’51.80’’, N38º01’00.80’’ W000º40’24.10’’, N38º01’50.30’’ W000º42’40.20’’, N38º01’01.00’’ W000º40’22.40’’, N38º01’50.90’’ W000º42’34.40’’); Mouthiez & Péru, 2008: 110 - “Loiret”; Tiberghien, 2010: 63 - “Arguedas, Navarra; Zaragoza; Pyrénées-Atlantiques”; Bartenev & Terekhova, 2011: 139 - Poltava Region, Kharkov Region, Chernihiv Region; Valladares, 2013: 39 - “Sorbas, Marchalico Viñicas”, “Olula de Castro, Sierra de los Filabres”, “Agua Amarga”, “Nijar”; Cartier & Cartier, 2016: 232 - “Vienne: Quinçay, Vouillé”; Touroult & al., 2019: 105 - France & Corse; Bacal et al., 2020: 59 - Moldova: “Ivancea”.
Agapanthia insularis Gautier des Cottes, 1870: 263 - “Corse aux environs d'Ajaccio, du côté de l'embouchure dela Gavone; Sicile, environs de Palerme”; Bellier, 1869: 14 - “île de Corse, aux environs d'Ajaccio, embouchure de la Gravone. Sicile, environs de Palerme”; Gemminger & Harold, 1873: 3176 - “Ajaccio”
Agapanthia reyi Mulsant & Godart, 1870: 27 - “Patrie: l'Espagne”; Argod, 1891a: 82; 1891b: xxxviii; Pic, 1891a: 38; 1891b: 82; Fauvel, 1895: 116; Winkler, 1929: 1213 - “Gallia meridionalis”; Plavilstshikov, 1930: 18 - “Spanien”; Breuning, 1961: 183 - “Espagne”.
Agapanthia cynarae var. delagrangei Pic, 1894: 75 - “Akbès”.
Agapanthia (s. str.) reyi, Pic, 1910: 95 - “Espagne”; Aurivillius, 1923: 464 - “Spanien”.
Agapanthia (Agapanthiella) asphodeli, Özdikmen, 2006: 87 - Turkey: “Ankara: Kızılcahamam, Işık Mountain, Keçikaya hill, 1615 m”, “Ankara: Kızılcahamam, Işık Mountain”, “Ankara: Kızılcahamam, Soğuksu National Park, 1250 m”, “Ankara: Kızılcahamam, Aköz village, 1150 m”.
Agapanthia (s. str.) asphodeli, Pic, 1910: 96 - “Medit.”; Plavilstshikov, 1916: 110 - Poltava; Aurivillius, 1923: 459 “Mittelmeer-Gebiet, Böhmen, Siebenbürgen”; Mikšić & Korpič, 1985: 100 - “Pronaden je г Dalmaciji, Hercogovini i Makedoniji”; Önalp, 1989: 199 - France, Spain, Turkey (Çanakkale, Izmir, Bilecik, Ankara, Antalya, Adana, Hatay); Özdikmen & Hasbenlý A., 2004: 41 - Turkey: “Isparta: Isparta-Burdur road, exit of Isparta, 1025 m”, “Antalya: near Manavgat waterfall, 25 m”, “Antalya: Kemer, Olimpos mountain, 561 m”, “Isparta: Yalvaç, Sultan mountains, 1596 m”, “Isparta: Yalvaç, Sultan mountains, 1674 m”, “Yozgat: Çiðdemli, Gökiniþ village, 1233 m”; Bartenev, 2004: 41 (= spencii Gyllenhal, 1813; = insularis Gautier, 1870), part. - Southern Europe, Western Europe, southern European Russia, Caucasus, Transcaucasia, Middle East, Turkey, Syria, North Africa Region, Ukraine, Kirovograd Region; 2009: 386 -Ukraine.
Agapanthia asphodeli v. mimica Pic, 1927: 1 - “Pyr.-Or.”.
Agapanthia (Epoptes) asphodeli, Sama & Rapuzzi, 2011: 142 - Italy: Basilicata, Campania, Lazio, Piemonte, Puglia, Sardegna, Sicilia, Toscana; Steiner & Schmid, 2013: 3; Özdikmen & Tezcan, 2020: 470 - Turkey: İzmir province; Zamoroka, 2022: 64 - “Ukraine”; Trócoli et al., 2023: 245 - Spain: Moianès (Barcelona, Catalogne); Danilevsky, 2023: 592, part.
Agapanthia asphodeli asphodeli, Sláma, 2019: 209 - France: “F 06 St. Rafael, F 86 Riboux, F 66 Banyuls”; “Süd. Spanien, Andalusien”.
Type locality. France, Bordeaux environs.
Description. Light parts of antennal joints from reddish to brown, but usually rather pale; black parts of antennal joints can be longer or shorter, from about half of each joint to 1/8 inside each population; elytra without metallic luster; elytral pubescence relatively dense; humeral grey line always absent; body length in males: 15.8-17.2 mm; body length in females: 16.0-23.0 mm.
Material. 1 male, Italy, Roma, Castlfusane De Maggi 11.4.1949 - ML; 1 female, Portogul, Porto - MD; 1 male, Madrid, J. Ardois - ZMM; 1 male, Grecia, 16.5.1973, Schimmel - MD; 1 female, “Rhodes / (Turquie d`Asie) / L. Bleuse” - ZMM; 2 males, 4 females, Turkey, Mersin, Içel, 6.4.1935, …? - MD; 1 female, Turkey, Karalailyas, 9.4.1935 - MD;; 2 males, 1 female, Turkey, Bandirma env., near Erdek, 40.4196ºN 27.8272ºE, 80-130 m 21-30.4.2011 S. Murzin - ML, SM; 1 male, Syria, Jebel el Ansariye (northern part), 5.1982, J. Kratochvil - MD; 1 female, Syria, Idlip env., 5.1982, J. Kratochvil - MD; 1 male, “Akbes / Syrien / Em. Reitter” - MD; 1 male, 8 females, “Akbes / Syrien / Em. Reitter” - ZMM; 1 male, Caucasus, Teberda, 3.6.1924 - ZMM.
Holotype of Agapanthia reyi Mulsant & Godart, 1870 (Figs 2-3), male with six labels: 1) [red] “HOLOTYPE”, 2) “36”, 3) “Espagne”, 4) "Collect. / Godart”, 5) “MUSEUM PARIS / Coll. Godart / Coll A. ARGOD. 1931”, 6) “Reyi / Muls. & Godart” / Opusc. ent. 14 cahier / p. 27.” - MNHN.
Distribution. All species area (with the exception of North Africa, Crete, Cyprus, Lampedusa Island, Peloponnesus and Kastamonu province of Turkey) from West Europe to North Caucasus, including Portugal, Spain, France, Italy, Malta, Croatia, Bosnia et Herzegovina, Macedonia, Albania, Greece, Bulgaria, Romania, Moldova, Ukraine, North Caucasus (Teberda), about whole west half of Anatolia and Syria.
Fig. 1. Agapanthia (Epoptes) asphodeli asphodeli (Latreille, 1804): male, Caucasus, Teberda, 3.6.1924.
Figs 2-3. Agapanthia (Epoptes) asphodeli asphodeli (Latreille, 1804): 2 - Holotype of Agapanthia reyi Mulsant & Godart, 1870, male with six labels: 1) [red] “HOLOTYPE”, 2) “36”, 3) “Espagne”, 4) "Collect. / Godart”, 5) “MUSEUM PARIS / Coll. Godart / Coll A. ARGOD. 1931”, 6) “Reyi / Muls. & Godart” / Opusc. ent. 14 cahier / p. 27.”; 3 - labels of Agapanthia reyi Mulsant & Godart, 1870.
Agapanthia (Epoptes) asphodeli lopadusae
Rapuzzi & Sparacio, 2017, stat. n.
Agapanthia asphodeli, Pisciotta et.al., 2008: 408 - “Lampedusa, loc. Isola dei Conigli”.
Agapanthia (Epoptes) lopadusae Rapuzzi & Sparacio, 2017: 958 - “Italy, Sicilian Channel, Lampedusa Island”; Danilevsky, 2020: 303 - Italy.
Type locality. Italy, Sicilian Channel, Lampedusa Island.
Description. The taxon belongs to A. asphodeli because of very elongated third joint of anterior tarsi, reddish 3rd-12th antennal joints, absence of the setae tufts on 3rd antennal joints and regular elytral pubescence.
It can be distinguished from other subspecies by darker antennae which are usually brownish basally from the third segments to the apices. The closest known taxon to A. a. lopadusae is A. a. zappii from North Africa, but it has distinct grey humeral stripes. Body length: 15.2-19.0 mm.
Distribution. The taxon is known from Lampedusa Island.
Agapanthia (Epoptes) asphodeli renatae
Steiner & Schmid, 2013, stat. n.
Saperda spencii, Brullé, 1832: 261 - “Morée”.
Agapanthia asphodeli, Berger, 2005: 93 - “Grèce: Péloponnèse: Dhafniotisa (Ilia)”.
Agapanthia (Epoptes) renatae Steiner & Schmid, 2013: 1, 3 - “Griechenland, Peloponnes, Nomos Arkadias, zwischen Karitena und Kastro, N37°29'10" O22°00'30", 450-550 m Seehöhe”; Danilevsky, 2020: 303 - Greece.
Agapanthia renatae, Rapuzzi & Sparacio, 2017: 957 - “Peloponnese (Greece)”
Agapanthia asphodeli balcanica Sláma, 2019: 208, syn. n. - “Graecia, Attiki, Villia”, “Graecia, Gerania; Greece, Meteora, 2 km. NE of Kalampaka - Kastraku”, “Graecia, prov. Pelloponese, Tripotama env. (Achaia), 40 km SE of Kalavryta, 37°52N, 21°53E, 675 m”, “Grece, Pelopones, Arkaia prov., Paradela env.”, “Kaesariani Attika”.
Agapanthia (Epoptes) asphodeli balcanica, Danilevsky, 2020: 301 - Greece.
Type locality. Greece, Peloponnese, Arcadia, between Karitena and Kastro.
Description. Elytra shining, look glabrous; black apical parts of antennomeres are wider than in the nominative subspecies (France); pronotum is wider (width-to-length ratio is 1.29-1.42); yellow lateral erect pubescence of pronotum is denser and longer; body usually wider and shorter. Body length: 12.0-20.0 mm.
Distribution. Greece, Peloponnese. Attica.
Remarks. Agapanthia (Epoptes) renatae Steiner & Schmid, 2013 and A. (E.) asphodeli balcanica Sláma, 2019 were both described from one area (Peloponnese), so, it must be regarded as one taxon: A. renatae Steiner & Schmid, 2013 = A. asphodeli balcanica Sláma, 2019.
Agapanthia (Epoptes) asphodeli cretica Bernhauer, 1978, stat. n.
Agapanthia cretica Bernhauer, 1978: 70 - “Kreta, Ida-Gebirge b. Anogia”, “Kreta, Chora Sfakion”; Rapuzzi & Sparacio, 2017: 957 - “Kriti (Greece)”.
Agapanthia probsti Holzschuh, 1984: 370, syn. n. - “Graecia, Kreta, Nomos Chanion, Omalos, 1000 m”, “Kreta, Lefka Ori, 3.7 km vor Omalos, 900 m”, “Kreta, SW Lefka Ori, Ag. Irini, 900 m”.
Agapanthia (Agapanthiella) cretica, Pesarini & Sabbadini, 2004: 127.
Agapanthia (Agapanthiella) probsti, Pesarini & Sabbadini, 2004: 127.
Agapanthia (Epoptes) cretica, Löbl & Smetana, 2010: 215 - Greece (Kriti); Steiner & Schmid, 2013: 2, 3; Danilevsky, 2020: 301 - Crete.
Agapanthia (Epoptes) probsti, Löbl & Smetana, 2010: 216 - Greece (Kriti); Steiner & Schmid, 2013: 2; Danilevsky, 2020: 303 - Crete.
Type locality. Crete, Mount Ida, Anogia.
Description. The taxon is distinctly differrent from closely related other subspecies of the group by dark elytra and its strong luster; eyes small; antennae relatively short, only about one third longer than body in males, or about just a little longer; black space of each antennal joint is relatively small, occupies a top apex of 3rd joint, about apical 1/3 of 3rd joint, and about halves of other joints; antennal tufts absent, just several single setae present near top of each joint; prothorax transverse, enlarged posteriorly, lateral and dorsal pronotal stripes very dense and orange, as well as marginal elytral stripes; elytra nearly glabrous, with indistinct setae spots, without humeral elytral stripes; in males about 3 times longer than wide, or about 2.6 times longer than wide in females; body length: 10.5-16.5 mm.
Material. “Ag. probsti”: 1 male, 1 female, Creta, Theviso, 6.1985, P. Schurmann - MD; “Ag. cretica”: 3 males, 1 female, Creta, Chora Sfakion, 4.1982 - MD; 3 males, 7 females, Creta, Chora Sfakion, 22.3.1991 M.Sláma - MD; 1 male, Creta, Biro, Canea, 12.3.1906 - MD.
Remarks. Agapanthia cretica Bernhauer, 1978 was described from Crete. A. probsti Holzschuh, 1984 was in fact undistinguished from A. cretica and also descriped from Crete - from about same type locality - so, Agapanthia cretica Bernhauer, 1978 (13.0-16.0 mm) = A. probsti Holzschuh, 1984 (10.5-13.8 mm).
Agapanthia (Epoptes) asphodeli kastamonica ssp. n.
Figs 4-6
Type locality. Turkey, Kastamonu province, Yaralıgöz Dağı.
Description. The taxon differs from European populations by extremely big eyes, lower eye lobe about 3 times longer than genae; antennae relatively short, about one forth longer than body in males, or by about 5 apical joints surpassing elytral apex; in females antennae surpassing elytral apex by 2 joints only; black space of each antennal joint is relatively small, occupies a top of 3rd joint, about apical 1/3 of 4th joint, and a little less than a half of other joints; antennal tufts absent, just several single setae present near joint apices; prothorax transverse, enlarged posteriorly; pronotal stripes light yellow; dorsal pronotal and lateral elytral stripes poorly developed; elytra very dark with rather dense punctation; nearly glabrous, with indistinct setae spots, without humeral elytral stripe; in males about 2.6 times longer than wide, or about 2.3 times longer than wide in females; body length: 17.0-18.0 mm.
Material. Holotype, male, Turkey, Kastamonu, Yaraligoz, 7.7.1997, N. Auvray - ML; 1 paratype, female with same label - ML.
Distribution. Turkey, Kastamonu province, Yaralıgöz Dağı [41°45'N, 34°10'E].
Etymology. The new taxon is named after its locality.
Figs 4-5. Agapanthia (Epoptes) asphodeli kastamonica ssp. n.:
4 - Holotype, male, Turkey, Kastamonu, Yaraligoz, 7.7.1997, N. Auvray; 5 - paratype, female with same label.
Figs 6-7. Heads, lateral views, males: 6 - Agapanthia (Epoptes) asphodeli kastamonica ssp. n., holotype; 7 - A. (E.) a. asphodeli (Latreille, 1804), Greece, 16.5.1973, Schimmel leg.
Agapanthia (Epoptes) asphodeli nicosiensis Pic, 1927, stat. rest.
Agapanthia dahli nicosiensis Pic, 1927: 1 - “Chypre”.
Agapanthia dahli m. nicosiensis, Breuning, 1961: 185.
Agapanthia nicosiensis, Rapuzzi & Sparacio, 2017: 957 - “Cyprus”.
Agapanthia (Agapanthiella) nicosiensis, Pesarini & Sabbadini, 2004: 127.
Agapanthia (Epoptes) nicosiensis, Löbl & Smetana, 2010: 216 - Cyprus; Ambrus, Grosser & Hrbek, 2014: 188 “Cyprus, 26 km SW Larnaka, 3 km N Skarinou, Dhypotamos Dam env.”, “Cyprus, Pafos, Agia Varvara”, “Cyprus, Akamas peninsula, 27 km N Paphos, 3 km W Neo Chorio, Camping site “Smigies” env.”, “Cyprus, Larnaca distr., Lageia env., 34°50'6.00"N 33°15'33.48"E”; Danilevsky, 2020: 303 - Cyprus.
Type locality. Cyprus.
Description. Eyes small; elytra with dense, spotted yellow pubescence and distinct grey humeral stripes; pale basal parts of 3rd-12th antennal joints light-red; setae tufts of 3rd antennal joints well developed; elytra in males about 2.8 times longer than wide; body length: 16.0-18.0 mm.
Distribution. Cyprus.
Material. 1 male, Cyprus, Nicosia - MD.
Agapanthia (Epoptes) asphodeli zappii Sama, 1987, stat. n.
Agapanthia cynarae, Lucas, 1847: 499 -pl. 42, figs 6, 6a, environs du cercle de Lacalle, lieux qui avoisinnent les lacs Tonga et Houbeira. ; environs d'Arzew.
Agapanthia asphodeli, Villiers, 1946: 118 - “Méditerranéenne”, “Maroc: Mogador, Tanger, Mèknes, Mélilla, Fèş, Sefrou, Forêts de la Mamora et des Zaers, Oujda, Talmest (2200 m), Tizi n’Tislet, Dj. Amar”, Algérie: Arzew, Oran, Saint-Charles, Téniet el Haad, Tunisie, Forêts de Yakouren, Alger, Constantine, La Calle, Lacs Tonga et Houbeira”, “Tunisie: El Feidja, Le Kef, Souk el Arba”; Heyrovský, 1937: 7 - “Libaah, Djezin, Chtaura, Saida”.
Agapanthia zappii Sama, 1987: 62 - “Algérie (Batna): col de Telmet (Aurès)”, “Maroc, Juadi Mellah”, Maroc, dint. Zaouia Temga”, “Maroc, Casablanca”, “Maroc, Salé”, “Maroc, Agadir”, “tra Taddert et Tizi-n’Ticka, 1650-2200 m”, “Libia (Tripolitania), Homs”; Chavanon, 1999: 171 - “Maroc Oriental: entre Oujda et Guenfouda”; Rapuzzi & Sparacio, 2017: 957 - “in North Africa from Lybia to Morocco”; Sama, Ringenbach & Rejzek, 2005: 448 - Libya.
Agapanthia (Agapanthiella) zappii, Pesarini & Sabbadini, 2004: 127.
Agapanthia (Epoptes) zappii, Chavanon et al., 2014: 296 “Aklim, 34°52’N – 02°31’W, 72 m”, “sud du col de Jerada, 34°17’N - 02°04’W, 1044 m”, “Taourirt, route d’Oujda, au pont sur l’oued Sifsif, 34°29’N - 02°53’W, 355 m”, “Saka, route de Nador, km 15, 34°41’N - 03°19’W, 580 m”; Löbl & Smetana, 2010: 216 - Algeria, Lybia, Morocco, Tunisia; Chavanon, 2018: 106 - Morocco, Oriental Region, Guercif Provinces: “Alentours du barrage Elhassi Labyad (au sud du col de Jerada) et toute la région située au nord d’une ligne Guercif-Touissit”; Danilevsky, 2020: 303 - Algeria, Lybia, Morocco, Tunisia; Kasatkin, 2020: 246; Trócoli, 2020: 29 - “Maroc: entre Oujda et Guenfouda”; Trócoli, 2023: 210 - Morocco, “Ifrane, Dayet Ahoua, 1467 m”.
Agapanthia (Epoptes) asphodeli, Cocquempot et al., 2016: 99 - Liban; Trócoli, 2020: 29 - “Le Nord, Centre et Ouest du Maroc, entre Oujda, Tanger, Casablanca et l’Atlas central, parfois plus au sud, dans la région côtière de Essaouira, Agadir”; Trócoli, 2023: 210 - Morocco, “Bouznika”, “Khemiset”, “Imilchil”.
Type locality. Algeria (Batna): Telmet canyon (Aures Mountains), 1700 m.
Description. According to Trócoli (2020), A. zappii Sama, 1987 is very close to A. asphodeli: “Ce sont deux espèces très proches, et zappi n’est peut-être qu’une variété d’asphodeli”. So, it must be accepted as a southern subspecies A. asphodeli zappii Sama, 1987, stat. n. A. a. asphodeli is also known from Morocco, but observed usually in the north part of the country. Still the areas of both taxa shown by Trócoli (2020) in his areal maps coincided totally.
A. asphodeli zappii differs from the nominative subspecies by elytral pubescence short, lighter, grey-yellow, with grey contrast humeral band; body length: 12.0-16.0 mm.
Distribution. North Africa (Morocco, Lybia, Algeria, Tunisia).
Material. 1 male, Morocco, Region Souss-Massa-Daraa, Prefecture Agadir-Ida ou Tanane, Commune Ida ou Tanane, 12 km E Aourir, 23.1.2002, Øistein Berg - MD; 1 male, paratype, Maroc., umg. Agadir, 12.85 Dr. Schurmann - MD; 1 female, paratype, Algeria, Batna, Col de Telmet, 1700 m, 11.6.1980, G. Sama & G. Magnani - MD.
Agapanthia (Epoptes) asphodeli fadli Sama & Rapuzzi, 2006, stat. n.
Agapanthia lateralis, Alfieri, 1916: 70 - “Egypte”; Petroff, 1919: 64 - “environs d’Alexandrie”.
Agapanthia fadli Sama & Rapuzzi, 2006: 186 - “14 km east of Burg al ‘Arab (west of Alexandria), 30°58’N 29°40’E”, “Egypt: “Mariut / Mars / Soc. Roy. Entom. Egypte / Faune d’Egypte”, “Hammam / (Dakahlich) / Egitto”; “Marzo / Faune d’Egypte”, “Maryout: Kinghi; Hammam, Abu Mina, Ikinghi Maryut, Damietta; El Borg; Burg el-Arab”; Rapuzzi & Sparacio, 2017: 957 - “North Egypt”.
Agapanthia (Epoptes) fadli, Löbl & Smetana, 2010: 215 - Egypt; Danilevsky, 2020: 303 - Egypt.
Type locality. Egypt, 14 km east of Burg al ‘Arab (west of Alexandria), 30°58’N 29°40’E.
Description. Body dark with strong bronze luster; lower eyes lobes short, about as long as genae; pronotum densely rugose; prothorax transverse, enlarged posteriorly; pronotal stripes light yellow; dorsal pronotal and lateral elytral stripes poorly developed; antennae long, about one half longer than body in males, or about 5 apical joints longer than body; 3rd antennal segment with a small setae tuft; black space of each antennal joint is relatively small, occupies a top apex of 3rd joint, about apical 1/3 of 3rd joint, and about halves of other joints; elytra densely setose, but setae spots indistinct, without humeral elytral stripe; elytra in males about 2.6 times longer than wide; body length: 11.0-21.0 mm.
Distribution. The taxon is known from Egypt only.
Acknowledgments. I am very grateful to M.L. Danilevsky (A.N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow), A.A. Gusakov (Zoological Museum of Moscow State University) and S.V. Murzin (Moscow) for supplying me with specimens for study. I am very grateful to Gérard Tavakilian and Christophe Rivier (Muséum National d'Histoire Naturelle, Paris) for supplying me with the photo of Agapanthia reyi Mulsant & Godart, 1870.
REFERENCES
Alfieri A. 1916. Séance du 3 Mai 1916. Catalogue des Cerambycides de l'Egypte. - Bulletin de la Société entomologique d'Egypte. 9 (2) [1916]: 63-76.
Ambrus R., Grosser W. & Hrbek J. 2014. Contribution to the knowledge of longhorn beetles from Cyprus (Coleoptera: Cerambycidae). - Humanity space. International almanac. 3 (2): 173-190.
Argod-Vallon A.A. 1891a. Extraits du Bulletin de la Société entomologique de France. Note sur Agapanthia Reyi Muls. et Godard. Séance du 25 février 1891. - L'Échange, Revue Linnéenne. 7 (81): 82.
Argod-Vallon A.A. 1891b. Séance du 25 février 1891. [Note sur Agapanthia Reyi Muls. et Godard]. - Bulletin de la Société entomologique de France, Paris. 60 (3): xxxviii-xxxix.
Aurivillius Ch. 1923. Cerambycidae: Lamiinae. II. Coleopterorum Catalogus pars 74 [Vol. 23] II: 323-704. W. Junk & S. Schenkling, Berlin.
Bacal S., Burduja D., Buşmachiu G., Cebotari C. & Merkl O. 2020: Longhorn beetles in the entomological collections of the Republic of Moldova (Coleoptera: Cerambycidae). - Folia Entomologica Hungarica. 81: 43- 72.
Baragaño Galan J.R., Notario Gómez A. & Sa Montero C. 1981.- Agapanthia asphodeli Lat. (col.: cerambycidae). Cría artificial y estudio cariológico. - Boletin de Sanidad Vegetal Plagas. 7 (1-2): 161-167.
Bartenev A.F. 2004. A review of the long-horned beetles species (Coleoptera: Cerambycidae) of the fauna of Ukraine. - Izvestiya Kharkovskogo Entomologicheskogo Obshchestva [The Kharkov Entomological Society Gazette]. 11 (1-2) (2003): 24-43. [in Russian]
Bartenev A.F. 2009. Longicorn-beetles of Left-Bank Ukraine and Crimea. Kharkov: Kharkov National University. 405 pp. [in Russian]
Bartenev A.F. & Terekhova V.V. 2011. An addition and remarks to the fauna of cerambycid beetles (Coleoptera, Cerambycidae) of Left-bank Ukraine and Crimea. - The Journal of V.N. Karazin Kharkiv National University. Series: biology. 13 (947): 133-146. [in Russian]
Bellier de La Chavignerie E. 1869. Lettre. - Petites Nouvelles Entomologiques Paris. 1 (4): 14.
Bernhauer D. 1978. Eine neue Agapanthien-Art aus Kreta (Coleoptera, Cerambycidae). - Nachrichtenblatt der Bayerischen Entomologen. 27 (4): 69-71.
Berger P. 2005. Contribution à la connaissance de la faune de Grèce: Coleoptera, Cerambycidae. 3ème note. - Biocosme Mésogéen, Nice. 22 (2): 81-96.
Breuning S. 1961. 4. Lieferung, pp: 183-284. In: S.Breuning, 1958-1969. Catalogue des Lamiaires du Monde (Col. Céramb.). Tutzing bei München, Verlag des Museums G. Frey: 1069 pp.
Brullé A.G. 1832-1836. Expédition scientifique de Morée. Section des Sciences Physiques. Tome III. - 1.re Partie. Zoologie. Deuxième Section. - Des animaux articulés. F.G. Levrault, Paris. 3 (1/2): 1-400, 52 pls. pages 289-400 issued in 1833; plates in 1832-1836.
Brustel H., Berger P. & Cocquempot Ch. 2003. Catalogue des Vesperidae et des Cerambycidae de la faune de France (Coleoptera). - Annales de la Société Entomologique de France, Paris (N. S.). 38 (4) (2002): 443-461.
Bonnamour S. 1935.Société linnéenne de Lyon. Faune des Coléoptères de la région lyonnaise. Famille des Longicornes (Suite et fin). In: Annales de la Société linnéenne de Lyon et des Société botanique de Lyon. - Société d'anthropologie et de biologie de Lyon réunies. 78 (1934): 105-116.
Carrière J. 1996a. Approche étho-biologique de Phytoecia virgula (Charpentier, 1825) sur Echinops ritro L, en région héraultaise (Coleoptera, Cerambycidae). - Lambillionea. 96 (numéro spécial du centenaire): 63-68, 6 figs.
Carrière J. 1996b. A propos d'Agapanthiini: note biologique complémentaire. (Coleoptera, Cerambycidae). - Lambillionea. 96 (3) 2: 561-570, 14 figs.
Carrière J. 1996c. Oviposition et stade post-embryonnaire d'Agapanthiini : à propos d'Agapanthia asphodeli (Latreille, 1804) et d'Agapanthia dahli (Richter, 1821) en région héraultaise, généraliés, aspect iconographique (Coleoptera, Cerambycidae). - Lambillionea. 96 (1) 1: 109-121, 11 figs.
Carrière J. 2001. Périnatalité et variantes morphologiques des larvules néonates d'Agapanthiini; réflexions sur les plantes-hôtes (Coleoptera, Cerambycidae). - Lambillionea. 101 (1) 3: 404-412, 12 figs.
Carrière J. 2003. Chalcidiens oophages et Agapanthiini: Note étho-biologique complémentaire. (Hymenoptera, Chalcididae - Coleoptera, Cerambycidae). - Lambillionea. 103 (2): 253-262.
Carrière J. 2005. Biogéographie et dynamique des populations Héraultaises d'Agapanthia kirby (Gyllenhal, 1817): approche physiologique comparative de l'oviposition (Coleoptera, Cerambycidae). - Lambillionea. 105 (3) 2: 463-470.
Carrière J. 2007. Impact des variations climatiques sur les populations saxicoles de Agapanthia asphodeli (Latreille, 1804). (Coleoptera, Cerambycidae). - Lambillionea. 107 (4) 1: 551-556, 8 figs.
Cartier J.-C. & Cartier G. 2016. Contribution à l'étude de l'entomofaune de la Vienne. Les Cerambycidae de la forêt de Vouillé-Saint-Hilaire (Coleoptera Cerambycidae). - L'Entomologiste. 72 (4): 221-234.
Chavanon G. 1999. Deuxième contribution à la connaissance des Cerambycidae du Maroc Oriental. - L'Entomologiste. 55 (4): 167-174.
Chavanon G. 2018. Catalogue des Coléoptères de la région orientale du Maroc (Province de Guercif exceptée). - Travaux de l’Institut Scientifique, Série Zoologie. 57: 1-192.
Chavanon G., Labrique H., François A. & Belal G.S. 2014. Troisième contribution à la connaissance des Cerambycidae du Maroc oriental (Coleoptera). - L’Entomologiste. 70 (5): 293-298.
Chernay A. 1854. Systematic catalog of objects stored in the zoological office of the Imperial Kharkiv University up to 1848. Kharkiv: University Printing House: 178-249. [in Russian]
Cocquempot Ch., Nemer N., Brustel H. & Tanios C. 2016. Nouvelles données et nouveau catalogue des Coléoptères Cerambycidae du Liban (Coleoptera, Cerambycoidea). - Bulletin de la Société Entomologique de France. 121 (1): 91-104.
Danilevsky M.L. 2020. Catalogue of Palaearctic Coleoptera. volume 6/1. Chrysomeloidea I (Vesperidae, Disteniidae, Cerambycidae) Updated and Revised Second Edition. Mikhail L. Danilevsky editor. Koninklijke Brill, Leiden 2: i-xxviii + 1-712.
Danilevsky M.L. 2023. Longicorn beetles (Coleoptera, Cerambycoidea) of Russia and adjacent countries. Part 3. М.: IAE, 2023. 874 p.
Danilevsky M.L. & Miroshnikov A.I. 1985. Timber-Beetles of Caucasus (Coleoptera, Cerambycidae). Key. Krasnodar: 419 pp. [in Russian]
Diego Barquín J. & Martínez-Porres Cáceres R. 2005. Cerambícidos nuevos para Cantabria, Burgos y Palencia (España) (Coleoptera, Cerambycidae). - Lambillionea. 105 (1): 143-145.
Echevarría Mayo J.M. 2003. Algunas citas complementarias para “Fauna Ibérica (Coleoptera, Cerambycidae)”. - Boletín de la Sociedad Entomológica Aragonesa. 32: 218.
Fauvel A. 1895. Notes synonymiques (suile el fin). - Revue Russe d’Entomologie. 14 (4-5):93-127.
Heyrovský L. 1937. Přispěvek k poznání tesaříků jižni Syrie. Beitrag zur kenntnis der Cerambyciden von Süed-Syrien. - Časopis Československé společnosti entomologické. 34: 6-9.
Ganglbauer L. 1884. Bestimmungs-Tabellen der europäischen Coleopteren. VIII. Cerambycidae. (Schluss.) Mit Berücksichtigung der Formen Algiers und des paläarktischen Asiens, exclusive jener von Japan. - Verhandlungen der kaiserlich-königlichen zoologisch-botanischen Gesellschaft in Wien. 33 (1883): 437-586, 2 figs.
Gautier des Cottes C. 1870. Nouvelles entomologiques ou recueil synonymique de descriptions d'espèces et genres nouveaux; monographique, de moeurs et remarques sur des insectes coléoptères de la faune européenne et méditerranéenne. Suite. - Mittheilungen der Schweizerischen entomologischen Gesellschaft. 3 (5): 257-264.
Gemminger M. & Harold E. von 1873. Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. - Munich. 10: 2989-3232 & index.
Gyllenhal L. 1817: [new taxa]. In: Schoenherr C. J.: Synonymia Insectorum, oder Versuch einer Synonymie aller bisher bekannten Insecten; nach Fabricii Systema Eleutheratorum etc. geordnet. Erster Band. Eleutherata oder Käfer. Dritter Theil. Hispa---Molorchus. Upsala: Em. Brucelius, 506 pp. + Appendix ad C. J. Schönherr Synonymiam Insectorum. Tom 1. Pars 3. Sistens descriptiones novarum specierum, 11+ 266 pp., pls. 5,6.
Holzschuh C. 1984. Beschreibung von 24 neuen Bockkäfern aus Europa und Asien, vorwiegend aus dem Himalaya (Coleoptera, Cerambycidae). - Entomologica Basiliensa. 9: 340-372.
Jeniš I. 2001. Tesařici - Long-horned Beetles. Disteniidae, Oxypeltidae, Vesperidae, Anoplodermatidae & Cerambycidae I. Vesperidae & Cerambycidae Evropy/of Europe I. Zlin: Atelier Regulus: 333 pp.
Kasatkin D.G. 2020. Contribution to the knowledge of the genus Agapanthia Audinet-Serville, 1835 (Coleoptera: Cerambycidae: Lamiinae) from the Near East and Transcaucasia. - Caucasian Entomological Bulletin. 16 (2): 233-249, 79 figs.
Kasatkin D.G. & Arzanov Ju.G. 1997. “Der Bockkaffer (Cerambycidae). Material fur Fauna der Kaffer (Coleoptera) norden Kaukasus und untere Don.” [wrong translation of the Russian title of the article; must be: “Die Bockkäfer (Cerambycidae) (Teil 2). Die Materialen zur Käferfauna (Coleoptera) des Nordkaukasus und des unteren Don] Records of Kharkov Entomological Society. 5 (2): 63-70.
Küster H.C. 1846. Die Käfer Europa's. Nach der Natur beschrieben. Mit Beiträgen mehrerer Entomologen. Nürnberg, Bauer & Raspe. 7: n° 1-100, 2 pls.
Latreille P.-A. 1804. Histoire Naturelle, Générale et particulière des Crustacés et des Insectes. Imprimerie F. Dufart, Paris. 11: iv + 1-424, pls 91-93.
Lucas P.H. 1847. Histoire Naturelle des animaux articulés. Coléoptères. In Exploration scientifique de l'Algérie pendant les années 1840, 1841, 1842. Sciences Physiques et Zoologie II. (Deuxième partie. Insectes) (1846) 2: 1-590, 47 pls.
Marquet J. 2001. Contribution à l'inventaire des insectes du département de l'Indre. Liste des Coléoptères du PNR Brenne. - L'Entomologiste. 57 (3-4): 101-122.
Mifsud D. 2002. Longhorn beetles (Coleoptera, Cerambycidae) of the Maltese Islands (Central Mediterranean). - The Central Mediterranean Naturalist, Malta. 3 (4): 161-169.
Mikšić R. & Korpič M. 1985: Cerambycidae Jugoslavije, III Dil. Sarajevo: Akademja Nauka i Umjetnosti Bosne i Hercegovine. 148 pp.
Mouthiez J. & Péru L. 2008. Liste des Longicornes observés dans le département du Loiret (Coleoptera Cerambycidae). - L'Entomologiste. 64 (2): 109-111.
Mulsant É. 1839. Histoire Naturelle des Coléoptères de France. Longicornes. Paris, Maison: vii-xii + 1-304 pp., 3 pls.
Mulsant É. 1862. Histoire Naturelle des Coléoptères de France. Longicornes. - Annales de la Société Impériale d'Agriculture, d'Histoire Naturelle et des Arts Utiles de Lyon: 1-480.
Mulsant E. & Godart A., 1870. Description d'une espèce nouvelle de Longicorne. - Opuscules Entomologiques. 14: 27-28.
Neculiseanu Z. & Baban E. 2005. Fauna cerambicidelor (Coleoptera: Cerambycidae) din Republica Moldova. - Analele stiintifice ale USM. Seria “Stiinte chimico-biologice”: 199-202.
Ogloblin D.A. 1948. 73. Fam. Cerambycidae – Timber Beetles or Longhorned Beetles. In: A Key of the Insects of the European Part of the USSR. Moscow-Leningrad: 450-471 [in Russian]
Önalp B. 1989. Agapanthia Serville 1835 (Coleoptera, Cerambycidae: Lamiinae) Türleri Üzerinetaksonomik Araştirmalar. - Hacettepe Üniversitesi EgitimFakültesi Dergisi. 4: 197-229.
Özdikmen H. 2006. Contribution to the knowledge of Turkish longicorn beetles fauna (Coleoptera: Cerambycidae). - Munis Entomology & Zoology. 1 (1): 71-90.
Özdikmen H. 2007. The Longicorn Beetles of Turkey (Coleoptera: Cerambycidae). Part I - Black Sea Region. - Munis Entomology & Zoology. 2 (2): 179-422.
Özdikmen H. 2008a. The Longicorn Beetles of Turkey (Coleoptera: Cerambycidae) Part II - Marmara Region. Munis Entomology & Zoology 3 (1): 7-152
Özdikmen H. 2008b. The Longicorn Beetles of Turkey (Coleoptera: Cerambycidae) Part III - Aegean Region. - Munis Entomology & Zoology. 3 (1): 355-436.
Özdikmen H. 2013. Turkish Agapanthiini Mulsant, 1839 with identification keys (Coleoptera: Lamiinae). - Munis Entomology & Zoology. 8 (1): 9-40.
Özdikmen H. 2021. An annotated catalogue: Cerambycoidea (Cerambycidae and Vesperidae) of Turkey (Coleoptera). - Munis Entomology & Zoology. 16 (Suplement): 1273-1556.
Özdikmen H. & Hasbenlý A. 2004. Contribution to the Knowledge of Longhorned Beetles (Coleoptera, Cerambycidae) from Turkey, Subfamily Lamiinae. - Journal of the Entomological Research Society. 6 (2): 25-49.
Özdikmen H. & Tezcan S. 2020. An important contribution to the knowledge of Lamiinae Fauna of Turkey (Coleoptera: Cerambycidae). - Munis Entomology & Zoology. 15 (2): 463-476.
Perris É. 1877. Larves de Coléoptères - suite. - Annales de la Société Linnéenne de Lyon. (2) 23 (1876): 1-590, 14 pls. (Paris, Deyrolle, Naturaliste 590 pp, 14 pls).
Peris-Felipo F.J., Moreno-Marí J., Oltra-Moscardó M.T. & Jimenez-Peydró R. 2008. Cerambícidos (Coleoptera: Cerambycidae) capturados en el Parque Natural de La Tinença de Benifassà (Castellón, España). - Boletín de la Asociación Española de Entomología, Salamanca. 32 (1-2): 95-116.
Peris-Felipo F.J., Falcó-Garí J.V., Oltra-Moscardó M.T. & Jimenez-Peydró R. 2010. Contribución al conocimiento de los Cerambícidos (Coleoptera: Cerambycidae) del Parque Natural de Las Lagunas de La Mata-Torrevieja (Alicante, España). - Boletín de la Asociación Española de Entomología, Salamanca 33 (3-4) (2009): 355-366.
Pesarini C. & Sabbadini A. 2004. Osservazioni sulla sistematica della tribù Agapanthiini Mulsant, 1839 (Coleoptera Cerambycidae). - Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano. 145 (1): 117-132, 16 figs.
Petroff A. 1919. Quelques notes sur les Cerambycides des environs d’Alexandrie. - Bulletin de la Société Entomologique d’Egypte: 58-64.
Pic M. 1891a. Toujours des Longicornes. - L'Échange, Revue Linnéenne. 7 (77): 38.
Pic M. 1891b. Note sur Agapanthia reyi Muls. et Godard. - L'Échange, Revue Linnéenne. 7 (81): 82.
Pic M. 1894. Trois coléoptères syriens. - L'Échange, Revue Linnéenne. 10 (114): 75-76.
Pic M. 1910. Catalogue bibliographique et synonymique des longicornes d’Europe et régions avoisinantes: suite. Pp. 95-98 [pagination speciale]. - In: Matériaux pour servir à l'étude des longicornes. 7ème cahier, 2ème partie. Lyon: Imprimerie Jacquet Frères: 95-98 pp.
Pic M. 1927. Notes diverses, descriptions et diagnoses (Suite.). - L'Échange, Revue Linnéenne 43 (427): 1-2.
Pisciotta S., Sajeva M. & Sparacio I. 2008. New records of Coleoptera Cerambycidae for Lampedusa Island. (Pelagian Is., Sicily). - Naturalista siciliano. 32: 405-409.
Plavilstshikov N.N. 1916. Les longicornes du governement de Poltava (Coleoptera, Cerambycidae). - Revue Russe d'Entomologie. 16 (1-2): 106-111.
Plavilstshikov N.N. 1930. Die Agapanthia-Arten der palaearktischen Region. Bestimmungs-Tabellen der europäischen Coleopteren. 98. Heft. Troppau: Edmund Reitter’s Nachfolger Emmerich Reitter, 40 pp.
Plavilstshikov N.N. 1932. Timber-beetles - Timber Pests. Moscow, Leningrad: 200 pp. [in Russian]
Plavilstshikov N.N. 1948. A Key for Longicorn Beetles of Armenia. Erevan: 232 pp. [in Russian]
Plavilstshikov N.N. 1965. 75-th Fam. Cerambycidae - Timber Beetles, Longicornes. - In: A Key to Insects of the European Part of the USSR, v. 2, Coleoptera and Strepsiptera. Moscow-Leningrad, “Nauka”: 389-419.
Plavilstshikov N.N. 1968. Review of the genus Agapanthia Serv. (Coleoptera, Cerambycidae) of the USSR fauna. - Archives of Zoological Museum Moscow State University. 11: 113-168.
Prisnyj A.V. & Vorobieva O.V. 2005 Scientific collection funds of “Museum of Zoology” at the chair of zoology and ecology of Belgorod State University. Issue 1. Insects - Ectognatha. Beetles – Coleoptera. Belgorod: IPC “POLITERRA”: 63 pp. [in Russian]
Rapuzzi P. & Sparacio I. 2017. A new species of genus Agapanthia Audinet-Serville, 1835 (Coleoptera Cerambycidae) from Lampedusa Island (Sicily Channel, Italy). - Biodiversity Journal. 8 (4): 957-961.
Reitter E. 1898. Ueber die bekannten und einige neue palaearctische Agapanthia-Arten. (Coleoptera.). - Wiener Entomologische Zeitung. 17 (4-5): 130-135.
Reitter E. 1913. Fauna Germanica. Die Käfer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. IV. Band. [1912]. Stuttgart: K.G. Lutz’ Verlag: 236 pp., pl. 129-152.
Sama G. 1979. Una nuova specie di Agapanthia Serville di Macedonia (Coleoptera: Cerambycidae). - Bolletino del Museo Civico di Storia Naturale di Verona. 5 (1978): 505-513, 4 figs.
Sama G. 1987. Note préliminaire pour une faune des longicornes de l'Afrique du Nord. - Biocosme Mésogéen, Nice. 4 (1): 43-64, 24 figs.
Sama G. 2003. Atlas of the Cerambycidae of Europe and the Mediterranean Area. Volume 1: Northern, Western, Central and Eastern Europe. British Isles and Continental Europe from France (excl. Corsica) to Scandinavia and Urals. Vít Kabourek, Zlín, (2002): 1-173, 729 figs.
Sama G., Ringenbach J.-C. & Rejzek M. 2005. A preliminary survey of the Cerambycidae of Lybia (Coleoptera). - Bulletin de la Société Entomologique de France. 110 (4/5): 439-454.
Sama G. & Rapuzzi P. 2006. Preliminary report on a recent survey of the Egyptian Cerambycidae, with description of three new species (Insecta Coleoptera Cerambycidae). - Quaderni di Studi e Notizie di Storia Naturale della Romagna. 23: 179-194, 6 figs.
Sama G. & Rapuzzi P. 2011. Una nuova Checklist dei Cerambycidae d’Italia (Insecta Coleoptera Cerambycidae). - Quaderno di Studi e Notizie di Storia Naturale della Romagna. 32: 121-164.
Schaefer L. 1947. Captures de Coléoptères et observations diverses. (suite et fin). - Bulletin mensuel de la Société linnéenne de Lyon. 16 (3): 52-53.
Seidlitz G.C.M. 1891. Die Kaefer (Coleoptera) Siebenbürgens. - Fauna Transsylvanica. Hartung, Königsberg. 5-6: i-lvi + 1-914, 1pl.
Sheshurak P.N. & Sadovnicha L.V. 2002. Terms of summer of longhorn beetles in Chernihiv region (Coleoptera: Cerambycidae). - Kharkov Entomological Society Gazette. 9 (1-2) (2001): 241-244.
Sláma M. 2019. A description of two new subspecies of European Longhorn-Beetles (Coleoptera, Cerambycidae). - Humanity space. International almanac. 8 (2): 208-214.
Steiner S. & Schmid H. 2013. Eine neue Agapanthia-Art (Coleoptera: Cerambycidae: Lamiinae: Agapanthiini) aus Griechenland. - Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen. 65: 1-4.
Sudre J., Foucart A., Cocquempot Ch. 1999. Catalogue commenté et étude bibliographique des Coléoptères Cerambycidae du département de l'Hérault. - Bulletin mensuel de la Société linnéenne de Lyon. 68 (6): 133-192.
Tiberghien G. 2010. Observations sur les Coléoptères de la chaîne pyrénéenne et régions limitrophes. 12 ème note: Cerambycidae (Coleoptera Phytophagoidea), suite. - Bulletin de la Société Linnéenne de Bordeaux. (145) 38 (1): 59-66.
Touroult J., Cima V., Bouyon H., Hanot Ch., Horellou A. & Brustel H. 2019. Longicornes de France - Atlas préliminaire (Coleoptera: Cerambycidae & Vesperidae). - Supplément au bulletin d’ACOREP-France, Paris: 1-176.
Trócoli S. 2020. Actualización del catálogo de Longicornios de Marruecos Actualisation du catalogue des Longicornes du Maroc (Parte IV / Partie IV: Cerambycidae: Lamiinae). - Revue de l'Association Roussillonnaise d'Entomologie (R.A.R.E.). 29 (1): 26-65, 35 figs, cartes 83-133.
Trócoli S. 2023. Nuevas aportaciones al Catálogo de Longicornios de Marruecos Nouvelles contributions au catalogue des Longicornes du Maroc (Coleoptera, Cerambycidae). - Revue de l'Association Roussillonnaise d'Entomologie (R.A.R.E.). 32 (3): 204-212.
Trócoli S., Mercadé A., Oliete C. & Aibar R. 2023. Los Longicornios del Moianès (Barcelona, Catalunya) (Coleoptera, Cerambycidae, Vesperidae). - Revue de l'Association Roussillonnaise d'Entomologie (R.A.R.E.). 32 (4): 237-247.
Valladares L., Calmont B., Soldati F., Brustel H. 2013. Contribución al conocimiento de los coleópteros (Coleoptera) de la Provincia de Almería (Andalucía, sureste de España). 2ª nota. Boletín de la Sociedad Andaluza de Entomología. 22: 25-66.
Valladares L., Soldati F., Calmont B. & Valladares M. 2003. Contribution à la connaissance des Coléoptères de la province d'Almeria (sud-est de l'Espagne) - 1ère note. - Revue de l'Association Roussillonnaise d'Entomologie (R.A.R.E.). 12 (3): 77-86.
Verdugo-Páez A. Los coleópteros Cerambycidae de la provincia de Cádiz (España) (Insecta: Coleoptera). - Boletín Sociedad Entomológica Cordobesa (SOCECO). 8 (Suplemento): 1-28.
Villiers A. 1946. Coléoptères Cérambycides de l'Afrique du Nord. - Faune de l'Empire Français, ORSC Paris. 5: 1-152, 275 figs.
Villiers A. 1978. Faune des Coléoptères de France I. Cerambycidae. Paul Lechevalier, Paris. Encyclopédie Entomologique 42: i-xxviii + 611 pp, 1802 figs.
Westwood J.O. 1839. An introduction to the modern classification of insects; founded on the natural habits and corresponding organisation of the different families. Longicornes or Capricornes. London. 1: iii-xii + 1-465.
Winkler A. 1929. Catalogus Coleopterorum regionis palaearcticae. II. Pars 9-10. Wien: 1009-1264.
Zamoroka A.M. 2022. The longhorn beetles (Coleoptera, Cerambycidae) of Ukraine: Results of two centuries of research. - Biosystems Diversity. 30 (1): 46-73.
Received: 10.06.2023
Accepted: 06.09.2023
