CC BY
8Mi
УДК 595.72
© Амурский зоологический журнал. I (1), 2009. 30-36.
© Amurian zoological journal. I (1), 2009. 30-36.
NEW SPECIES AND NEW RECORDS OF THE RHYNCHITID BEETLES (COLEOPTERA, RHYNCHITIDAE) FROM ASIA A.A. Legalov
[А.А. Легалов, Новые виды и новые находки жуков-ринхитид (Coleoptera, Rhynchitidae) из Азии]
Siberian Zoological Museum, Institute of Systematics and Ecology of Animals, Russian Academy of Sciences, Siberian Branch, Frunze street 11, Novosibirsk, 630091, Russia. E-mail: legalov@ngs.ru
Сибирский зоологический музей, Институт систематики и экологии животных СО РАН, ул. Фрунзе 11, Новосибирск, 630091, Россия. E-mail: legalov@ngs.ru
Summary. New species of rhynchitid beetles: Pseudomesauletes (Rubrauletes) thompsoni Legalov, sp.n. (China: Yunnan), Eusproda fengshuensis Legalov, sp.n. (China: Fujian), Temnocerus (Temnocerus) daliangensis Legalov, sp.n. (China: Sichuan), Auletomorphinus dundai Legalov, sp.n. (Thailand), A. sutensis Legalov, sp.n. (Malaysia: Sarawak), Byctiscus qingensis Legalov, sp.n. (China: Shaanxi), Aspidobyctiscus (Aspidobyctiscus) marshalli Legalov, sp.n. (Nepal, India), A. (A.) nanpingensis Legalov, sp.n. (China: Sichuan) are described. The status of Byctiscus princeps regalis (Roelofs, 1874), stat.n. is restored from synonym of Byctiscus princeps (Salsky, 1872) to subspecies. The studied material on rare species (Nelasiorhynchites ussuriensis (Legalov, 2002), Epirhynchites (Tshernyshevinius) auratus (Scopoli, 1763), Svetlanaebyctiscus vitis (Ter-Minassian, 1959), Byctiscophilus championi Voss, 1931, Byctiscus bilineatoides Legalov, 2007, B. macros Legalov, 2004, B. fukienensis Voss, 1948, B. fulminans Voss, 1930, B. populi (Linnaeus, 1758)) is listed.
Резюме. В статье описываются новые виды ринхитид: Pseudomesauletes (Rubrauletes) thompsoni Legalov, sp.n. (Китай, Юннань), Eusproda fengshuensis Legalov, sp.n. (Китай, Фудзянь), Temnocerus (Temnocerus) daliangensis Legalov, sp.n. (Китай, Сычуань), Auletomorphinus dundai Legalov, sp.n. (Таиланд), A. sutensis Legalov, sp.n. (Малайзия, Саравак), Byctiscus qingensis Legalov, sp.n. (Китай, Шанси), Aspidobyctiscus (Aspidobyctiscus) marshalli Legalov, sp.n. (Непал, Индия), A. (A.) nanpingensis Legalov, sp.n. (Китай, Сычуань). Byctiscus princeps regalis (Roelofs, 1874), stat.n. восстановлен из синонимов Byctiscus princeps (Salsky, 1872) в подвиды. Приводится материал для редких видов: Nelasiorhynchites ussuriensis (Legalov, 2002), Epirhynchites (Tshernyshevinius) auratus (Scopoli, 1763), Svetlanaebyctiscus vitis (Ter-Minassian, 1959), Byctiscophilus championi Voss, 1931, Byctiscus bilineatoides Legalov, 2007, B. macros Legalov, 2004, B. fukienensis Voss, 1948, B. fulminans Voss, 1930, B. populi (Linnaeus, 1758).
INTRODUCTION
Leaf-rolling weevils of family Rhynchitidae are common in the Oriental and in Southeastern Palaearctic, with the diversity centre in the Southeast Asia. These beetles are common in forest biotopes; usually cutting plant parts, less frequently rolling leaves. The group is inadequately studied, with many species still undescribed. Present paper continues the author's research into Asian Rhynchitidae [Legalov, 2000 (2001), 2001, 2002, 2002 (2003), 2003, 2004a, 2004b, 2004c, 2006a, 2006b, 2006c, 2007, 2008; Legalov, Fremuth, 2002; Legalov, Liu, 2005; Legalov, Korotyaev, 2006].
MATERIAL AND METHODS
Types and specimens are stored in the following collections and museums: CKJU - P. Kresl Collection (Czech Republic: Janovice nad Uhlavou); ISNB - Institut Royal des Sciences Naturelles de Belgique (Belgium: Brussels); IZAS - Institute of Zoology, Academia Sinica, (China: Beijing); NMPC - National Museum of Natural History (Czech Republic: Prague); RDP - R. Dunda Collection (Czech Republic: Prague); SMTD = Staatliches Museum fur Tierkunde (Germany: Dresden); ZMHB - Museum fur Naturkunde der Humboldt-Universitat (Germany: Berlin); ZMN - Zoological Museum, Institute of Animal Systematics and Ecology (Russia : Novosibirsk).
RESULTS
Family Rhynchitidae Gistel, 1848 Supertribe Rhynchititae Gistel, 1848 Tribe Auletini Desbrochers des Loges, 1908 Subtribe Pseudomesauletina Legalov, 2003 Genus Pseudomesauletes Legalov, 2001 Subgenus Rubrauletes Legalov, 2003
Pseudomesauletes (Rubrauletes) thompsoni Legalov,
sp.n. (Col. pl. I - a, b; fig. 1 - a, b)
Material. Holotype, male (NMPC), "China, Yunnan prov., Weibaoshan mts.". Paratype: male (ZMN), female (RDP), idem.
Description. Body reddish brown. Head, rostrum, clava of antennae, scutellum, thorax, femora, 3rd and clausal segments of tarsi brown. Body with short adpressed pale setae.
Male. Rostrum long, 6.0-6.5 times longer than wide, 1.41-1.44 times longer than pronotum, weakly curved, widened to the apex, densely punctate. Antennae located in the middle of rostrum. Eyes not large, strongly convex. Forehead wide, strongly convex, finely punctate. Temples short and straight.
Antennae long, reaching beyond the front of prono-tum. Scapus and 1st segment of funicle oval. 2nd-4th segments long-oval, narrower. 2nd segment a little shorter than 1st; 3rd segment a little longer than 2nd; 4th segment shorter and wider than 3rd. 5th segment trapezoid, wider than 4th segment. 6th segment almost rounded. 7th seg-
ment transversal, wider than 6th segment. Clava wide, almost compact, pointed, a little shorter than funicle. 1st and 2nd segments transversal. 3rd segment stilliform, a little shorter than previous segments.
Pronotum almost campaniform, length/width = 0.941.0, weakly narrowed to basis and apex. Disk convex, small and densely punctate. Greatest width is in the middle. Scutellum trapezoid.
Elytra almost rectangular, elongated, 1.31-1.33 times longer than wide, their greatest width behind the middle. Humeri weakly smoothed. Striae reduced. Points large and deep. Intervals weakly convex. Apex of elytra with sex patches.
Thorax small and sparsely punctate. Metepisternum narrow.
Abdomen convex. 1st and 2nd ventrites wide; 2nd ventrite a little wider than 1st. 3rd and 4th ventrites narrower than 2nd. 5th ventrite narrow, narrower than 4th. Pygidium convex, punctate.
Legs long. Femora widened. Tibiae almost direct, weakly widened to apex. Protibiae narrow and long. Tarsi long. Protarsi hardly more flattened and longer than meso- and metatarsi. 1st tarsal segment long-triangular. 2nd segment wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 2.6-2.7 mm.
Female. Rostrum longer, 7.33 times longer than wide, 1.57 times longer than pronotum. Antennae narrower than it of male. Pronotum of equal length and width, with more weakly rounded sides. Elytra stronger widened to apex, 1.4 times longer than wide, without sex patches. Length of body: 3.2 mm.
Diagnosis. This new species is close to Pseudomesauletes (Rubrauletes) consimilis (Voss, 1930) but differs by red-brown tibiae, brown femora and by the armament of endophallus.
Etymology. New species is named in honour of R. Thompson.
Tribe Isotheini Scudder, 1893
Subtribe Deporaina Voss, 1929
Genus Eusproda Sawada, 1987
Eusproda fengshuensis Legalov, sp.n. (Col. pl. I - c)
Material. Holotype, female (NMPC), "China, N Fujian, Fengshui Guan, 27.9 N, 117.85 E, 1700 m, 1-4.VI.2004, J. Turna".
Description. Female. Body black, lustrous, with short semierect setae. First one-third of rostrum with dense erect long setae.
Rostrum long, 3.46 times longer than wide, 1.41 times longer than pronotum, weakly curved, widened to apex, almost smooth. Antennae located before the middle of rostrum. Eyes not large, strongly convex. Forehead wide, weakly convex, dense and small punctate. Temples straight lines, weakly elongated.
Antennae long, reaching first line of pronotum. Scapus and 1st segment of funicle elongated-oval, almost equal length. 2nd-4th segments elongated, narrow. 1st -3rd segments equal length. 4th segment shorter than 3rd segment. 5th segment trapezoid, shorter than 4th segment. 6th segment oval. 7th segment almost trapezoid, wide,
wider than 6th segment. Clava not compact, shorter than funicle. 1st and 2nd segments oval, equal length. 3rd segment tear-shaped, longer than 2nd segment.
Pronotum almost campaniform, equal length and width, with almost direct sides. Disk convex, small and densely punctate. Greatest width before middle. Scutellum trapezoid.
Elytra back-ovoid, 1.22 times longer than wide. Greatest width behind middle. Humeri weakly smoothed. Striae clear and deep. 9th striae merge with 10th striae near apex of elytra. Points large and dense. Intervals weakly convex.
Thorax punctate. Metepisternum almost wide.
Abdomen convex. 1st and 2nd ventrites wide. 3rd and 4th ventrites narrower. 3rd ventrite narrower than 2nd ventrite. 4th ventrite narrower than 3rd ventrite. 5th ventrite much narrower than 4th ventrite. Propygidium and pygidium convex, densely punctate.
Legs long. Femora widened. Tibiae almost direct, weakly widened to apex. Tarsi long. 1st segment elongated. 2nd segment triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 4.3 mm.
Diagnosis. This new species is close to Eusproda proxima (Faust, 1882) but differs by a larger body, stronger narrowed sides of pronotum, longer rostrum and more narrow elytra.
Etymology. The name is formed from the location "Fengshui" - "fengshuensis".
Tribe Rhynchitini Gistel, 1848
Subtribe Lasiorhynchitina Legalov, 2003
Genus Nelasiorhynchites Legalov, 2003
Nelasiorhynchites ussuriensis (Legalov, 2002)
Material. female (NMPC), "China, Shaanxi prov., Taibaishan nat. park, 1350 m, 10.VI.1999, M. Murzin"; male (ZIN), "S Korea, GB Prov., Kimcheong City, Chupung ryeong, resting place, 17.V.2000, B. Korotyaev"; female (ZMN), "Russia, Primorie terr., 15 km SW of Slavjanka, Quercus-forest, 16.VI.1993, S. BelNfatbyl sTijis species is recorded in China for the first time.
Subtribe Temnocerina Legalov, 2003
Genus Temnocerus Thunberg, 1815
Subgenus Temnocerus s. str.
Temnocerus (Temnocerus) daliangensis Legalov, sp.n. (Col. pl. I - d)
Material. Holotype, female (NMPC), "China, S Sichuan, Daliang Shan Mts., Zhaojue vill. env., pass Xichang - Meigu vill., 12-14.VI.1998, M. Tryzna"
Description. Body black, with goldish lustre, with sparsely, short, semierect setae. Antennae and tarsi dark brown.
Female. Rostrum short, 7.0 times longer than wide, 2.0 times longer than pronotum, weakly curved towards apex, lustrous. Topmost third weakly punctate. Other part of rostrum densely long punctate. Antennae located almost before the middle of rostrum. Forehead wide,
weakly convex, densely punctate. Eyes large, convex. Temples long, transversely wrinkled.
Antennae long, reaching the first line of pronotum. Scapus and 1st segment of funicle oval. Scapus shorter than the 1st segment. 2-4th segments narrow. 2nd segment shorter than 1st segment. 3rd segment equal to 2nd segment. 4th segment shorter than 3rd segment. 5th segment oval, shorter than 4th segment. 6th segment wider than 5th segment. 7th segment almost trapezoid. Clava shorter than funicle, thicker, not compact. 1st segment longer than 2nd segment. 3rd segment tear-shaped, longer than 1st segment.
Pronotum weakly elongated, 1.52 times wider than long. Sides almost straight. Disk little flattened, large and densely punctate. Intervals lacking lustre. Scutellum almost square, small punctate.
Elytra elongated, 1.52 times longer than wide. Greatest width behind the middle. Humeri weakly smoothed. Striae weak with dense fine points. Scutellar stria advanced. Intervals narrow, weakly convex, smooth.
Thorax sparsely punctate. Abdomen convex, finely and sparsely rugoso-punctate. 1st and 2nd ventrites wide. 3-4th ventrites narrow. 5 th ventrite narrower. 2-4th ven-trites weakly flattened on middle. Pygidium convex, finely and sparsely punctate.
Legs long. Femora weakly widened. Tibiae thin. Protibiae elongated and narrow, almost straight. Meso-and metatibiae thicker and shorter. Tarsi long and narrow, little shorter than tibiae. 1st segment long, longer than 2nd segment. 2nd segment triangular. 3rd segment bilobed. 5th segment elongated. Claws with long teeth.
Length of body: 3.0 mm.
Diagnosis. This new species is very close to Tem-nocerus (Temnocerus) dundai Legalov, 2006 but differs by the narrower and more sparsely punctate pronotum, and shorter rostrum.
Etymology. The name is formed from the location "Daliang" - "daliangensis".
Subtribe Rhynchitina Gistel, 1848
Genus Auletomorphinus Legalov, 2007
Auletomorphinus dundai Legalov, sp.n. (Col. pl. I - e)
Material. Holotype, female (NMPC), "NW Thailand, Mae Hong Son, Ban Huai Po, 1600-2000 m, 8-18.V.1992, J. Horak". Paratype - female (RDP), "NE Thailand, Loei prov., Phu Kradung N. P., 16.52 N, 101.49 E, 1000 m, 16-17.V.1999, D. Hauck".
Description. Female. Body red-brown. Head, rostrum, antennae, pronotum, scutellum, elytra, apex of femora, tibiae and tarsi black. Humeri and edges of elytra green-violet. Body with sparse dark semierect setae. Meso- and metepisternum with stains from white adpressed setae.
Rostrum long, 6.9-7.5 times longer than wide, 1.461.61 times longer than pronotum, almost straight, without carina, weakly widened towards the apex, finely punctate. Antennae attached before the rostrum base. Eyes large, weakly convex. Forehead convex, wide, almost smooth. Vertex convex, almost smooth. Temples short, weakly transversal-wrinkled.
Antennae thin and long, reaching the middle of prono-tum. Scapus and 1st segment of funicle oval. 1st segment
longer than scapus. 2nd-5th segments narrow, elongated. 2nd segment shorter than 1st segment. 3rd segment longer than 2nd segment. 4th segment equal to 3rd segment. 5th segment shorter and wider than 4th segment. 6th segment oval. 7th segment almost rounded. Clava wide, sharp, not compact. 1st and 2nd segments trapezoid. 1st segment longer than 2nd segment. 3rd segment tear-shaped, shorter and narrower than 3rd segment.
Pronotum campaniform, 1.08-1.18. times wider than length. Sides rounded. Disk convex, dense roughly rugo-sely-punctate, with weak middle line. Greatest width before middle. Scutellum trapezoid, wide.
Elytra almost rectangular, 1.16-1.24 times longer than width. Greatest width in humeri behind middle. Humeri weakly convex. Intervals narrow, convex. Striae wide. Points in them large and deep.
Prothorax punctate. Mesothorax and mesepisternum densely punctate. Metathorax sparsely punctate. Metepis-ternum densely punctate. Abdomen convex, small punctate. 1st and 2nd ventrites wide. 3rd and 4th ventrites narrower. 5th ventrite very narrow. Pygidium convex, sparsely punctate.
Legs long. Femora widened. Tibiae almost straight, long, weakly widened to apex, with two spines at the apex. Tarsi long. 1st segment elongated. 2nd segment wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth.
Length of body: 4.3-5.0 mm.
Diagnosis. This new species differs from the others by the roughly rugosely-punctate pronotum, with its greatest width closer to the middle.
Etymology. This new species is named in honour of R. Dunda.
Auletomorphinus sutensis Legalov, sp.n. (Col. pl. I -f, g; fig. 1 - c, d)
Material. Holotype, male (NMPC), "Sarawak, Kapit dist., Rumah Ugap vill., Sut riv., 3-9.III.1994, J. Horak". Paratypes: male (ZMN), female (RDP), idem.
Description. Body red-brown. Head, rostrum, antennae, pronotum, scutellum, elytra, apex of femora, tibiae and tarsi black. Elytra with green-violet lustre. Body with sparse dark semierect setae. Meso- and metepisternum with stains from white adpressed setae.
Male. Rostrum long, 8.0-8.36 times longer than wide, 1.37-1.48 times longer than pronotum, weakly curved, weakly widened to the apex, finely punctate. Antennae attached before the rostrum base. Eyes large, weakly convex. Forehead convex, wide, almost smooth. Vertex convex, almost smooth. Temples short, weakly transversal-wrinkled.
Antennae thin and long, reaching pronotum middle. Scapus and 1st segment of funicle oval. 1st segment shorter than scapus. 2nd-5th segments elongated trape-zoid. 2nd segment shorter than 1st segment. 3rd segment longer than 2nd segment. 4th segment hardly shorter than 3rd segment. 5th segment shorter and wider than 4th segment. 6th segment trapezoid, wide and short. 7th segment almost rounded. Clava wide, not compact. 1st and 2nd segments almost trapezoid. 1st segment longer than 2nd segment. 3rd segment tear-shaped, hardly longer than 1st segment.
Fig. 1. Male genitalia: a-b - Pseuäomesauletes ihompsoni, c-d - Auletomorphinus sutensis, e - Aspidobyctiscus nanpingensis, f-g - A. marxhalli (ho)otype).
Pronotum campaniform, 1.07-1.09 times wider than length. Sides weakly rounded. Disk convex, weakly rugosity-punctate. Greatest width before middle. Scutel-lum trapezoid, wide.
Elytra almost rectangular, 1.08-1.21 times longer than wide. Greatest width in humeri behind middle. Humeri weakly convex. Intervals wide, convex. Striae wide. Points in them large and deep.
Prothorax punctate. Mesothorax and mesepisternum densely punctate. Metathorax sparsely punctate. Metepis-ternum densely punctate. Abdomen convex, finely punctate. 1st and 2nd ventrites wide. 3rd and 4th ventrites narrower. 5th ventrite very narrow. Pygidium convex, sparsely punctate.
Legs long. Femora widened. Tibiae almost straight, long, wide, widened to apex. Tarsi long. 1st segment triangular. 2nd segment wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth.
Length of body: 4.9-5.4 mm.
Female. Rostrum longer, 9.14 times longer than wide, 1.83 times longer than pronotum. Pronotum 1.16 times longer than wide. Sides of pronotum weaker rounded. Elytra 1.14 times longer than wide. Length of body: 5.1 mm.
Diagnosis. This new species is close to Auletomor-phinus rubrofemoralis (Legalov, 2003) but differs by the elytra shorter and wider, pronotum sides more narrowed to apex and clava of the antennae narrower.
Etymology. The name is formed from the location "Sut" - "sutensis".
Genus Haplorhynchites Voss, 1924
Haplorhynchites hampsoni (Voss, 1938)
Material. Female (CKJU), "S India, Tamil Nadu st, Nilgin hills, alt. 2100 m, 10 km SW of Manjoor, near Car-ington estate, Thiasola res., forest, 76'35e, 11'12 n, 14-19.VI.1999, Kejval, Tryzna".
Genus Epirhynchites Voss, 1969
Subgenus Tshernyshevinius Legalov, 2003
Epirhynchites (Tshernyshevinius) auratus (Scopoli, 1763)
Material. Female (ZMN), "Tuva, Erzinskii Distr., 100 km SE of Balgazyn, Erzinskie gol'tzy, 2000 m, 25.VI.1999, A. Marikoda".
Notes. The species is new to Republic of Tuva.
Tribe Byctiscini Voss, 1923
Subtribe Svetlanaebyctiscina Legalov, 2003
Genus Svetlanaebyctiscus Legalov, 2001
Svetlanaebyctiscus vitis (Ter-Minassian, 1959)
Material. 10 ex. (ZMN), "Primorskii krai, Lazovskii rez., bukhta Petrova, on Vitis, 20-23.VII.2007, A. Lega-lov, E. Shevnin".
Genus Byctiscophilus Voss, 1930
Byctiscophilus championi Voss, 1931
Material. Female (CKJU), "Nepal, Lumie, Annapuma, Hima, 17-22.VI.1999, A. Kudma".
Subtribe Byctiscina Voss, 1923
Genus Byctiscus C.G. Thomson, 1859
Byctiscus princeps princeps (Solsky, 1872) (Col. pl. II - c)
Rhynchites princeps Solsky, 1872: 284
Notes. Nominate subspecies is distributed on a continental part of Asia (China, Japan, Korea and Russian Far East).
Byctiscus princeps regalis (Roelofs, 1874), stat.n.
(Col. pl. II - b)
Rhynchites regalis Roelofs, 1874: 142
Material. Lectotype, male (ISNB), "male", "type", "Coll. R. I. Sc. N. B., Japon: Rec. G. Lewis, Coll. Roelofs", "R. regalis. R, Japon, L.". Paralectotypes: male (ISNB), female (ISNB), "type", "Coll. R. I. Sc. N. B., Japon: Rec. G. Lewis, Coll. Roelofs". Specimens: Male (SMTD), "Japonia, Damel"; 3 males (RDP), 2 males (ZMN), 3 females (RDP), "Japan, Yamaguchi pref., To-kusa, 29.IV.1979, Y. Takeshita".
Diagnosis. This subspecies differs from nominate subspecies by the forehead, rostrum top, mesothorax, episternum and legs with green or golden lustre.
Byctiscus qingensis Legalov, sp.n. (Col. pl. II - a)
Material. Holotype, female (NMPC), "China, Shaanxi, Qing Ling Shan mts., road Baoji - Taibai vill., pass 35 km S of Baoji, 21-23.VI.1998, O. Safranek, M. Tryzna".
Description. Female. Body green, lustrous, smooth. Forehead, pronotum sides, 4 stains on elytra, partially mesothorax, meso- and metepisternum purple. Rostrum top, disk of pronotum, side of ventrites, procoxa with golden lustre. Sides of rostrum and legs with purple-violet lustre. Bottom of femora with blue-violet lustre.
Rostrum short, strongly curved, 3.0 times longer than wide, widened to apex, finely punctate. Antennae attached to the middle of rostrum. Forehead wide, pressed, punctate. Eyes not protruding from contour of head. Vertex convex, finely and densely punctate. Temples elongated.
Antennae short, not reaching pronotum. Scapus and 1st-5th segments of funicle oval. 1st segment longer than scapus. 2nd segment much shorter than 1st segment. 3rd segment hardly shorter than 2nd segment. 5th segment shorter and wider than 4th segment. 6th segment roundish. 7th segment transversal, wider than 6th segment. Clava long, little shorter than funicle, flattened, almost compact. 1st and 2nd segments almost equal length. 3rd segment tear-shaped, longer than 2nd segment.
Pronotum wide, 1.19 times wider than length, 1.15 times longer than rostrum. Sides rounded. Disk convex, finely punctate. Grooves weak, gently wrinkled. Scutel-lum wide, rectangular, finely punctate.
Elytra almost rectangular, 1.14 times longer than wide, weakly pressed before scutellum. Humeri weakly convex. Greatest width behind the middle. Intervals narrow, flat, finely punctate. Striae weak. Points in them small.
Thorax weakly rugosely-punctate. Abdomen convex, dense rugosely punctate. 1st ventrite with blades. 1st-3rd ventrites wide. 4th ventrite narrower. 5th ventrite very narrow. Pygidium convex, very small punctate.
Legs long. Femora weakly widened, rugosely punctate. Protibiae weakly curved, long, narrow, densely punctate. Meso- and metatibiae weakly biconcave, more strongly widened towards the apex. Tarsi long. 1st segment triangular. 2nd segment wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 6.6 mm.
Diagnosis. This new species is close to Byctiscus princeps (Solsky, 1872) but differs by a smaller and less densely punctate pronotum, with less rounded sides and forehead and metepisternum more sparsely punctate.
Etymology. The name is formed from the location "Qing" - "qingensis".
Byctiscus bilineatoides Legalov, 2007 (Col. pl. I - h)
Material. Female (RDP), China, Yunnan prov., Yu-longshan mts., 27.13 N, 100.16 E, 3200 m, 14.VII.1990, Vit Kuban; male (ZMN), female (RDP), China, Sichuan pr., Nanping (Jiuzhaigou), VI.1990, CWs. leg.
Byctiscus macros Legalov, 2004 (Col. pl. I - i)
Material. Female (RDP), "China, Shaanxi, Qinling mts., 6 km E of Xunyangba, 23.V.-13.VI.1998, I. H. Marshal"; female (IZAS), "Sichuan, Kangding, Wasigou, 21-22.VIII.1939, Zhou Yao, Zheng Fengliu, Hao Tianhe".
Byctiscus fukienensis Voss, 1948 (Col. pl. II - e)
Material. Male (ZMN), "China, Fujian, near Jianyang, 5.IX.1997".
Byctiscus fulminans Voss, 1930 (Col. pl. II - d)
Material. Male (ZMN), "China, Jiangxi, near Ningdu, 24.IV.1996".
Byctiscus populi (Linnaeus, 1758) (Col. pl. II - f)
Material. Female (RDP), "China, Sichuan pr., Nanping (Jiuzhaigou), VI.1990, CWs. leg".
Genus Aspidobyctiscus Schilsky, 1903
Subgenus Aspidobyctiscus s. str.
Aspidobyctiscus (Aspidobyctiscus) marshalli Legalov, sp.n. (Figs. 2h, 2i, 3f, 3g)
Material. Holotype, male (NMPC), "Nepal, Dha-walagiri, Wille - Ghorepani, 1600-2600 m, 10.VI.1986, CWs. leg". Paratypes: male (RDP), male (ZMN), female (RDP), female (ZMN), idem; female (CKJU), "Nepal b., Dhunche env., Rasuwa distr., 2500 m, 10-14.VI.1999, Kresl"; male (RDP), "N India, Uttar Pradesh, Joshimat, Pulna, 3 km of Ghangaria, 2300 m, 23.VII.1989, A. Riedel"; female (ZMHB), "India, Nilghedi Hills".
Description. Body bronze, naked.
Male. Rostrum long, weakly curved, 3.83-4.0 times longer than wide, widened towards the apex, finely punctate. Antennae located behind the rostrum middle. Forehead wide, pressed, densely punctate. Eyes not protruding from contour of head. Vertex convex, densely punctate. Temples elongated. Prementum almost semicircular.
Antennae short, not reaching pronotum. Scapus and 1st segment of funicle oval. 2nd segment elongated-oval, narrower and shorter than 1st segment. 3rd and 4th segments oval. 3rd segment shorter than 2nd segment. 4th segment shorter than 3rd segment. 5th segment roundish. 6th and 7th segments transversal. 6th segment wider than 5th segment. 7th segment wider than 6th segment. Clava long, shorter than funicle, flattened, wide. 1st and 2nd segments wide trapezoid, equal length. 3rd segment wide, tear-shaped, longer than 2nd segment.
Pronotum wide, 1.13-1.15 times wider than length, 1.07-1.18 times longer than wide. Sides weakly rounded. Disk strongly convex, densely wrinkled, with middle striae. Grooves weak. Scutellum wide, rectangular, finely punctate.
Elytra almost rectangular, 1.08-1.12 times longer than wide, very weakly pressed for scutellum. Humeri weakly convex. Greatest width in humeri and on middle. Intervals wide, convex, small and densely punctate. Striae clear. Points in them partially merge. 9th striae merge with 10th striae before 2nd ventrite.
Prothorax weakly wrinkled, with long teeth directed forwards. Meso- and metathorax with episternum densely rugositely-punctate. Metepisternum wide. Abdomen convex, rugosely punctate, weakly flattened in the middle. 1st ventrite with blades. 1st-2nd ventrites wide. 2nd ventrite wider than 1st ventrite. 3rd ventrite narrower. 4th ventrite narrow. 5th ventrite very narrow. Pygidium convex, densely punctate.
Legs long. Femora weakly widened, small rugosely punctate. Protibiae almost direct, long, narrow, densely punctate. Meso- and metatibiae weakly biconcave, stronger widened towards the apex. Tarsi long. 1st segment triangular. 2nd segment wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 4.5-5.4 mm.
Female. Rostrum shorter, 3.14-3.43 times longer than wide. Pronotum narrower, 1.19-1.25 times wider than length, 1.0-1.18 times longer than rostrum. Elytra 1.081.19 longer than wide. Length of body: 4.7-5.4 mm.
Diagnosis. This new species is close to Aspidobyctis-cus (Aspidobyctiscus) yunnanicus (Voss, 1930) but differs by the more gentle sculpture of the elytra and by the armament of endophallus.
Etymology. This new species is named in honour of G.A.K. Marshall.
Aspidobyctiscus (Aspidobyctiscus) nanpingensis Legalov, sp.n. (Col. pl. II - g; fig. 1 - e)
Material. Holotype, male (NMPC), "China, Sichuan pr., Nanping (Jiuzhai gou), VI.1990, CWs. leg.". Para-types: male (RDP), male (ZMN), idem.
Description. Male. Body bronze, naked.
Rostrum long, weakly curved, 3.75-4.0 times longer than wide, widened to apex, small punctate. Antennae attached behind the rostrum middle. Forehead wide, pressed, densely punctate. Eyes not protruding from contour of head. Vertex convex, densely punctate, with weak striae. Temples elongated, transversely-wrinkled. Premen-tum lamellate.
Antennae short, not reaching pronotum. Scapus and 1st segment of funicle oval. Scapus shorter than 1st seg-
ment. 2nd - 4th segments narrower than 1st segment. 2nd segment much shorter than 1st segment. 3rd segment longer than 2nd segment. 4th segment shorter than 3rd segment. 5th segment trapezoid, shorter and wider than 4th segment. 6th segment trapezoid, more wider than 5th segment. 7th segment transversal, narrow. Clava long, shorter than funicle, flattened, wide. 1st and 2nd segments widely trapezoid. 2nd segment shorter than 1st segment. 3rd segment wide, tear-shaped, longer than 1st segment.
Pronotum wide, 1.06-1.16 times wider than length, 1.081.16 longer than rostrum. Sides rounded. Disk strongly convex, densely wrinkled, with middle striae. Grooves weak. Scutellum wide, rectangular, finely punctate.
Elytra almost rectangular, length/width = 1.0-1.1, weakly pressed for scutellum. Humeri weakly convex. Greatest width in humeri and in the middle. Intervals wide, convex, small and densely punctate. Striae narrow. Points in them partially merge. 9th striae merge with 10th striae before 2nd ventrite.
Prothorax weakly wrinkled, with long teeth directed forwards. Meso- and metathorax with episternum densely rugosely-punctate. Metepisternum wide. Abdomen convex, rugosely punctate, weakly flattened in the middle. 1st ventrite with blades. 1st-2nd ventrites wide. 2nd ventrite wider than 1st ventrite. 3rd ventrite narrower. 4th ventrite narrow. 5th ventrite very narrow. Pygidium convex, densely punctate.
Legs long. Femora weakly widened, small rugosely punctate. Protibiae almost straight, long, narrow, densely punctate. Meso- and metatibiae weakly biconcave, more strongly widened towards the apex. Tarsi long. 1st segment triangular. 2nd segment wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 5.6-6.2 mm.
Diagnosis. This new species is close to Aspidobyctis-cus (Aspidobyctiscus) lacunipennis (Jekel, 1860) but differs by a wider pronotum with stronger-rounded sides.
Etymology. The name is formed from the location "Nanping" - "nanpingensis".
ACKNOWLEDGEMENTS
I wish to thank D. Drugmand (Bruxelles), R. Dunda (Prague), D. Efimov (Kemerovo), J. Hajek (Prague), O. Jaeger (Dresden), K.-D. Klass (Dresden), A. Korshunov (Kemerovo), P. Kresl (Janovice nad Uhlavou), N. Liu (Beijing), P. Limbourg (Bruxelles) and J. Willers (Berlin), for help with the work.
REFERENCES
Legalov A.A. Svetlanaebyctiscus gen.n., eine neue Gattung der Tribus Byctiscini aus dem Fernen Osten (Coleoptera, Attelabidae) // Russian Entomological Journal. 2000 (2001). Vol. 9. № 4. P. 341-343.
Legalov A.A. Revision der holarktischen Auletini (Coleoptera, Attelabidae) // Russian Entomological Journal. 2001. Vol. 10. № 1. P. 33-66.
Legalov A.A. Species of the genus Lasiorhynchites (Coleoptera, Rhynchitidae) in the Far Eastern fauna // Entomological Review. 2002. Vol. 82. № 8. P. 10991101.
Legalov A.A., Fremuth J. Neue Arten der Familie Rhynchitidae (Coleoptera) aus der Türkei // Russian Entomological Journal. 2002. Vol. 11. № 2. P. 215-219.
Legalov A.A. A new species of the genus Temnocerus Thunberg, 1815 (Coleoptera, Rhynchitidae) from Japan // Russian Entomological Journal. 2002 (2003). Vol. 11. № 4. P. 409-410.
Legalov A.A. Taxonomy, classification and phylogeny of the leaf-rolling weevils (Coleoptera: Rhynchitidae, Atte-labidae) of the world fauna. Novosibirsk, 2003. CD-R. № 0320301200. 733+350 p. (641 Mb.) [in Russian].
Legalov A.A. A new species of the genus Involvulus (Coleoptera, Rhynchitidae) from the South of the Far East Russia // Zoological Herald. 2004a. Vol. 38. № 1. P. 8587. [in Russian].
Legalov A.A. New data of the leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) of the world fauna with description of 35 new taxons // Baltic Journal of Coleopterology. 2004b. Vol. 4. № 1. P. 63-88.
Legalov A.A. A new species of the genus Hap-lorhynchites Voss (Coleoptera, Rhynchitidae) from India // Entomological Review. 2004c. Vol. 84. №. 9. P. 994-997.
Legalov A.A., Liu N. New leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from China // Baltic Journal of Coleopterology. 2005. Vol. 5. № 2. P. 99-132.
Legalov A.A. Three new species of the leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) from Russia, China and Korea // Baltic Journal of Coleopterology. 2006a. Vol. 6. № 1. P. 15-22.
Legalov A.A. Two new species of the genus Deporaus Sam. (Coleoptera: Rhynchitidae) from the Russian Far East and China // Far Eastern Entomologist. 2006b. № 164. P. 1-6.
Legalov A.A. To the knowledge of the genus Temnocerus Thunberg, 1815 (Coleoptera: Rhynchitidae) // Far Eastern Entomologist. 2006c. № 165. P. 1-14.
Legalov A.A., Korotyaev B.A. A new species of the genus Temnocerus Thunb. (Coleoptera: Rhynchitidae) from Kazakhstan // Baltic Journal of Coleopterology. 2006. Vol. 6. № 2. P. 125-127.
Legalov A.A. Leaf-rolling weevils (Coleoptera: Rhynchitidae, Attelabidae) of the world fauna. Novosibirsk: Agro-Siberia, 2007. 523 pp.
Legalov A.A. New species of the tribe Byctiscini (Coleoptera, Curculionidae) from Yunnan // Baltic Journal of Coleopterology. 2008. Vol. 8. № 1. P. 49-54.
Roelofs W. Curculionides recueillis au Japon par M. G. Lewis // Annales de la Société Entomologique de Belgique. 1874. T. 17. P. 121-176.
Solsky S.M. 1872. Description d'un charencon nouveau de la Sibérie orientale // Horae Societatis Entomologicae Rossicae. 1871 (1872). T. 8. No. 1-4. P. 284-286.
Fig. 1. Rhynchitidae gen. spp.: a - Pseudomcsauletes (hompsoni {cf, holotype), b - P. ihompsoni (9, paratype), с -Eusprodafengshuensis (9, holotype), d - Temnocerus daliangensis (9, holotype), e - Aulelomotphinm dundai (9. holotype), f-A. sutensis (0\ holotype), g-A. sutensis (9, paratype), h - Byctiscus bilineatoides (9), i-B, macros (9).
Fig, 2. Byctíscinagcn, »pp.: a -Byctiscusqingensis (9, holotypc), b - B. princeps regalis (9), с - В.princeps princeps (o')5 d - B. fuíminans (ct), e - B. fukienensis (cf). f - B, popiití (9), g - Aspidabyetiscus nanpingensis (cf, holotype), h -/í. imrshaíii {cf, holotype), i -Л. marshaüi (9, paratype).
