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Амурский зоологический журнал, 2022, т. XIV, № 4
Amurian Zoological Journal, 2022, vol. XIV, no. 4
www.azjournal.ru
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https://www.doi.org/10.33910/2686-9519-2022-14-4-562-569 http://zoobank.org/References/71A78E4C-1D32-45C6-AAD4-87F9F5929703
UDC 595.773.4
Notes on the genus Psacadina (Diptera, Sciomyzidae)
N. E. Vikhrev
Zoological Museum of Moscow University, 2 Bolshaya Nikitskaya Str., 125009, Moscow, Russia
Author
Nikita E. Vikhrev
E-mail: nikita6510@yandex.ru
SPIN: 1266-1140
Scopus Author ID: 32467511100
Copyright: © The Author (2022). Published by Herzen State Pedagogical University of Russia. Open access under CC BY-NC License 4.0.
Abstract. A review of the Palaearctic genus Psacadina Enderlein, 1939 is given. Diagnostic characters of Psacadina species offered in previous publications by Rozkosny (1987), Vala (1989) and Rivosecchi (1992) are discussed. The author came to the conclusion that using the shape of surstylus as the main diagnostic character leads to difficulties, while the form of sternite 5 makes it possible to make an unambiguous diagnosis. The diagnostic value of the wing pattern is confirmed. A new identification key for both sexes of Psacadina is proposed. Two new synonyms are proposed: Psacadina zernyi Mayer, 1953 = P. verbekei Rozkosny, 1975, syn. nov. and P. vittigera Schiner, 1864 = P. disjecta Enderlein, 1939, syn. nov. Distribution of Psacadina species in the eastern part of Palaearctic is significantly clarified.
Keywords: Psacadina, taxonomy, distribution, Sciomyzidae, Diptera
Заметки роду Psacadina (Diptera, Sciomyzidae)
Н. Е. Вихрев
Зоологический музей МГУ им. М. В. Ломоносова, ул. Большая Никитская, д. 2, 125009,
г. Москва, Россия
Сведения об авторе Вихрев Никита Евгеньевич E-mail: nikita6510@yandex.ru SPIN-код: 1266-1140 Scopus Author ID: 32467511100
Права: © Автор (2022). Опубликовано Российским государственным педагогическим университетом им. А. И. Герцена. Открытый доступ на условиях лицензии СС БУ-ЫС 4.0.
Аннотация. Предложен обзор палеарктического рода Psacadina Enderlein, 1939. Дано критическое обсуждение диагностических признаков Psacadina, использованных в предыдущих работах: Rozkosny (1987), Vala (1989) и Rivosecchi (1992). Автор пришел к выводу, что использование формы сурстилей как основного диагностического признака приводит к трудностям, тогда как форма стернита 5 позволяет поставить однозначный диагноз. Предложен новый определительный ключ видов Psacadina, который позволяет надежно отличать как самцов, так и самок. Предложены новые синонимы: Psacadina zernyi Mayer, 1953 = P. verbekei Rozkosny, 1975, syn. nov. и P. vittigera Schiner, 1864 = P. disjecta Enderlein, 1939, syn. nov. Уточнено распространение видов в восточной части Палеарктики.
Ключевые слова: Psacadina, систематика, распространение, Sciomyzidae, Diptera
Introduction
Psacadina Enderlein, 1939 is a genus of Tetanocerini endemic to the Palaearctic, in which five species have been regarded as valid since 1975. It is characterized as follows: two pairs of orbital setae; pedicel short, more wide than long (Fig. 1); arista with long black hairs; anepisternum with one strong seta at posterior margin and covered with several setulae; subalar setae absent; anepimeron with setu-lae; scutellum with two pairs of setae; wing with dark pattern as in Figs. 2-3; hind coxa with setulae on inner posterior margin; male f3 ventrally with two rows of spinulose setae.
Previously, Psacadina was considered as a subgenus of Pherbina. These genera share most characters including the same wing pattern, but Pherbina has pedicel longer than wide; subalar setae present and anepimeron apart from setulae with 1-2 strong seta(e).
About 30 years ago the genus Psacadina was reviewed in three almost simultaneous publications: Rozkosny (1987), Vala (1989) and Rivosecchi (1992). However, proposals of these authors were in many respects con-tradictive. This circumstance lead to permanent difficulties in identification of Psacadina species which mostly based on the structure of male surstyli (= gonostyli of authors). In the present work I offer two new synonyms: Psacadina zernyi Mayer, 1953 = P verbekei Rozkosny, 1975, syn. nov. and P vittigera Schiner, 1864 = P disjecta Enderlein, 1939, syn. nov. With two synonyms newly proposed, the genus Psacadina is no longer problematic; three remaining valid species may be reliably distinguished in both sexes as proposed in the key below. Moreover, reliable identification of the rich material in the Zoological Museum of Moscow University (not indicated in text) and Saint Petersburg Zoological Institute (indicated as ZIN) significantly clarified the distribution of Psacadina in the eastern part of Palaearctic.
Material and methods
Localities are given as follows: country, region/state/province (in italics), and geographi-
cal coordinates in decimal-degree format. Full names of the regions of the Russian administrative subdivisions are an entangled result of political and historical events of no interest for zoology, so they are listed as a name (taken from the English version of Wikipedia) and the word "region" (abbreviated in the text as "Reg").
Illustrations are original unless otherwise credited. When referring to figures, I capitalize the first letter (Fig. or Figs.) for those appearing in this paper and use lowercase (fig. or figs.) for those published elsewhere to avoid confusion.
Material examined
Psacadina kaszabi Elberg, 1978 Figs. 9, 13
RUSSIA: Amur Reg.: Zeya env. (53.77°N 127.28°E), 12-14.08.1981, A. Shatalkin, 2$; Klimoutsy (40 km W of Svobodny (51.46°N 127.59°E), 31.08.1958, A. Zinovjev, 4? (ZIN); Buryatia Reg.: Tunka env., 51.7°N 102.6°E, 750 m, 7-11.06.2021, N. Vikhrev, 10$, 5?; Mondy env., 51.67°N 101.04°E, 1250 m, 17.06.2021, N. Vikhrev, 1?; Irkutsk Reg., Slyudyanka, 51.68°N 103.69°E, 480 m, 12-14.06.2021, N. Vikhrev, 5$; Khakas Reg., Shira Distr., Itkol L., 54.477°N 90.120°E, 11-20.04.2022, M. Esin, 1$, 1?; Primorsky Reg., Lotos L., 42.46°N 130.64°E, 1-3.07.2014, N. Vikhrev, 1$; Tuva Reg.: Uyuk R., 800 m, 52.072°N 94.047°E, 27.05.2018, N. Vikhrev, 11$, 7?; Kyzyl, poplar-birch forest, 51.7°N 94.7°E, 17-25.05.2018, N. Vikhrev, 1$; Zabaikalsky Reg., Ulyatui (51.17°N 116.25°E), 740 m, 25-26.05.2014, A. Medvedev, 1$, 1?; 3-6.06.2014, A. Medvedev, 1$. DISTRIBUTION. Not rare in Mongolia and south-eastern Siberia, uncommon in the Far East. To the west known till 90°E, to the north till 54°N.
Psacadina vittigera Schiner, 1864 Figs. 3-8, 14 Psacadina disjecta Enderlein, 1939, syn. nov. (see Discussion)
ESTONIA, Peedu (59.2°N 25.6°E), A. Stackelberg, 22-24.08.1951, 4$, 5? (ZIN).
Fig. 1. Psacadina zernyi, male Рис. 1. Psacadina zernyi, самец
KAZAKHSTAN, Pavlodar (= N Kazakhstan) Reg., Petropavlovsk, 54.84°N 69.10°E, 28.06.2015, O. Kosterin, 1$. RUSSIA: Kursk Reg., Oboyan env., Psyol R. (51.191°N 36.313°E), 26.05.2007, N. Vikhrev, A. Ozerov, 6$, 2?;
Moscow Reg., Ilyinskoe (55.755°N 37.233°E), 31.03.1922, B. Rodendorf, 1?; Kashira Distr., Ozherelye, 54.77°N 38.24°E, 2-4.05.2018, K. Tomkovich, 1$; Dmitrov Distr. (E of Kostino, 56.31°N 37.81°E), 30.08.2007, N. Vikhrev, 1?; Mordovia Reg.: 20 km W of Saransk, 54.137°N 44.906°E, 21.06.2020, N. Vikhrev, 1$; Pushta env., 54.71°N 43.22°E, 1-5.09.2020, N. Vikhrev, 1?; Saint Petersburg Reg., Luga Distr., Yaschera (59.15°N 29.91°E), 28-31.08.1963, A. Stackelberg, 6$, 2? (ZIN).
SPAIN, Almeria Prov., Tijola (37.35°N 2.43°W), 13.03.1964, L. Knutson, 1$, genitalia not dissected, identification label by J. Ver-beke, 1966: P disjecta Enderlein (ZIN).
TURKEY, Antalya Reg., Side, sand dune (36.76°N 31.43°E), 28.02.2008, N. Vikhrev, 1$; 25.02.2008, K. Tomkovich, 1?. DISTRIBUTION. European. Also recorded in Turkey and N Kazakhstan, uncommon in the eastern part of its range. To the north recorded till 60°N, to the south till 36°N.
Psacadina zernyi Mayer, 1953 Figs. 1, 2, 10-11
Psacadina verbekei Rozkosny, 1975, syn. nov. (see Discussion)
AZERBAIJAN, Nakhchivan (39.2°N 45.4°E), 28.05.1957, L. Zimina, 1$. BELARUS, Gomel Reg.: Mytva R., 51.854°N 29.223°E, 31.07.2019, N. Vikhrev, 1$; Mozyr env., 52.02°N 29.32°E, 21.05.2019, N. Vikhrev, 1$; 11-14.06.2019, N. Vikhrev, 1$, 1?; Vitebsk Reg., Ezerische env., 55.83°N 30.00°E, 16-18.05.2019, N. Vikhrev, 1$. ESTONIA, Peedu (59.2°N 25.6°E) , A. Stackelberg, 1.07.1951, 1?; 22.08.1951, 1$ (ZIN).
Figs. 2-3. Wing pattern of Psacadina: 2a, b: P. zernyi: a — wing with smaller rounded spots; b — wing with larger diamond-shaped spots; 3a, b: P. vittigera: a — wing with rectangular spots; b— wing with trapezoid spots
Рис. 2-3. Крыло Psacadina: 2a, b: P. zernyi: a — крыло с маленькими округлыми пятнами; b — крыло с большими ромбовидными пятнами; 3a, b: P. vittigera: a — крыло с прямоугольными пятнами; b — крыло с пятнами трапециевидной формы
KAZAKHSTAN: Aktobe Reg., Chesnokovo, 51.66°N 52.17°E, 26.08.2012, Tomkovich, 1$; Pavlodar (= N Kazakhstan) Reg., Ishim R., 53.35°N 67.05°E, 15.08.2015, O. Kosterin, 1$; KYRGYZSTAN: Choktal (Issyk-Kul L.), 42.58°N 76.75°E, 19-22.09.2013, N. Vikhrev, 2$; Issyk-Kul L., S of Rybachy (presently Ba-lykchi, 42.4°N 76.2°E), 16.08.1969, E. Nar-chuk, 4$,1? (ZIN).
RUSSIA: Altai Krai Reg., Zmeinogorsk Distr., Kolyvanskoe L. (51.36°N 82.18°E), 8.09.2007, O. Kosterin, 2$, 1?;
Amur Reg., Zeya env. (53.77°N 127.28°E), 12-14.08.1981, A. Shatalkin, 4$, 2?; Astrakhan Reg., Baskunchak L., freshwater pond, 48.193°N 46.813°E, 1-6.05.2010, K. Tomkovich, 1$, 2?; Bashkortostan Reg., Abzakovo env., 53.82°N 58.62°E, 500 m, 15-19.06.2020, N. Vikhrev, 1?; Buryatia Reg.: Kyren env., 51.7°N 102.1°E, 750 m, 16-19.06.2021, N. Vikhrev, 1$, 1?; Tunka env., 51.7°N 102.6°E, 750 m, 7-11.06.2021, N. Vikhrev, 1$, 4?; Irkutsk Reg., Slyudyanka, 51.68°N 103.69°E, 480 m, 12-14.06.2021, N. Vikhrev, 2$; Kaliningrad Reg., Baltiysk, 54.644°N 19.939°E, on Phragmites, 23.08.2013, K. Tomkovich, 1$;
Karachay-Cherkes Reg., B. Zelenchuk R., 44.22°N 41.72°E, 580 m, 6.07.2019, O. Kosterin, 1$;
Kemerovo Reg., 20 km SWW of Kemerovo, 55.25°N 85.77°E, 9.07.2021, D. Efimov, 1$; Krasnodar Reg., Adler env., 43.439°N 39.954°E, 23.04.2008, A. Ozerov, D. Gavryu-shin, 3& 1$;
Kursk Reg., Oboyan env., Psyol R. (51.191°N 36.313°E), 26.05.2007, N. Vikhrev, A. Ozerov, 1& 2$;
Mordovia Reg., Smolny NP, 54.734°N 45.397°E, K. Tomkovich, 2.08.2022, 1$, 1$; Moscow Reg.: Dmitrov Distr. (E of Kostino, 56.31°N 37.81°E), 9-11.05.2007, N. Vikhrev, 3& 2$; 2-9.07.2007, N. Vikhrev, 2$; Severovo, YPT, 55.04°N 37.50°E, 6-8.06.2021, K. Tomkovich, 2^, 1$; Volokolamsk env., Lama R., 55.97°N 36.03°E, 11.05.2021, N. Vikhrev, 3$;
Novosibirsk Reg. (Academgorodok env.), Irtysh R. oxbows, 54.86°N 83.04°E, 1-2.05.2016, O. Kosterin, 1$;
Omsk Reg., Cherlak env. (54.15°N 74.80°E), 10.05.2009, O. Kosterin, 1$, 2$; Orenburg Reg., Aituarskaya step (51.12°N 57.67°E), 11.05.2005, K. Tomkovich, 1$;
Figs. 4-6. Psacadina vittigera: drawings of male postabdomen with surstyli (= gonostyli) in ventral view: 4 — from Rozkosny (1987, fig. 321); 5 — drawing of surstyli from Vala (1989, fig. 96a) attached to Rozkosny's drawing of postabdomen; 6 — drawing of surstyli from Rivosecchi (1992, fig. 129c) attached to Rozkosny's drawing of postabdomen Рис. 4-6. Psacadina vittigera: рисунок постабдомена и сурстилей (= гоностилей), вид снизу: 4 — из Rozkosny (1987, fig. 321); 5 — рисунок сурстилей из Vala (1989, fig. 96a), прикрепленный к рисунку постабдомена из Rozkosny (1987); 6 — рисунок сурстилей из Rivosecchi (1992, fig. 129c), прикрепленный к рисунку постабдомена из Rozkosny (1987)
Saint Petersburg Reg.: Luga (58.7°N 29.9°E), 2-17.08.1952, A. Stackelberg, 9$, 3$ (ZIN); Luga Disr., Yaschera (59.15°N 29.91°E), 28-31.08.1963, A. Stackelberg, 6$, 2$ (ZIN); Primorsky Reg., 30 km SE of Ussuriysk (Kamenush-ka, 43.63°N 132.23°E), 31.05.1985, A. Ozerov, 1$; Stavropol Reg., Kislovodsk env., 43.97°N 42.79°E, 700 m, 14-17.04.2022, N. Vikhrev, 1$; Tver Reg., Rzhev env., 56.21°N 34.35°E, 18.08.2014, N. Vikhrev, 2$; Ulyanovsk Reg., Vyazovka env., 52.82°N 48.35°E, 2-8.05.2011, K. Tomkovich, 1$; Volgograd Reg., Kalach-na-Donu env., 48.65°N 43.60°E, 30.04.2013, N. Vikhrev, 1$; Yakutia-Sakha Reg., Yakutsk (62.0°N 129.7°E) env., 31.05.1985, A. Barkalov, 1$; Zabaikalsky Reg., Ulyatui (51.17°N 116.25°E), 740 m, 3-6.06.2014, A. Medvedev, 1$. TURKEY, Sakarya Reg., Karasu (41.1°N 30.7°E) env., 27.08.2009, N. Vikhrev, 1$. DISTRIBUTION. Palaearctic from W Europe to Far East. To the north recorded till 62°N, to the south till 39°N.
Identification key for Psacadina $$
1. Wing with rounded (Fig. 2a) or diamond-shaped (Fig. 2b) spots along costal margin; posterior crossvein S-shaped. (Body
length $?: 5.6-6.6 mm; Transpalae-arctic.) $: sternite 5 narrow, elongated, with curved posteriorly apices as in Fig. 11. (Surstyli ear-shaped as in Fig. 10.) ............................zernyi Mayer
— Wing with inverted trapezoid (Fig. 3b) or subquadrate (Fig. 3a) spots along costal margin; posterior crossvein not S-shaped, only with a fold at middle. $: sternite 5 not as above, surstyli not ear-shaped........2
2. Body length $?: 5.2-6.2 mm. European, also recorded in Turkey and N Kazakhstan, uncommon in the eastern part of its range. $: sternite 5 as in Fig. 12 (half moon shaped, with rounded projection on posterior margin and a notch on anterior margin)....................vittigera Schiner
— Body length $?: 3.8-4.8 mm. East Palae-arctic eastern of 90°E. Common in southeastern Siberia and Mongolia, uncommon in the Far East. $: sternite 5 as in Fig. 13 (with notch on posterior margin and long projection on anterior margin) ..........................kaszabi Elberg
Discussion
Most authors (Rozkosny 1987; Rivosecchi
1992) divided species of Psacadina into two
Figs. 7-10. Surstylus of Psacadina: 7, 8 — the same surstylus of P. vittigera from a different angle of view; 9 — P. kaszabi; 10 — P. zernyi
Рис. 7-10. Сурстили Psacadina: 7, 8 — один и тот же сурстиль P. vittigera при съемке с разных углов зрения; 9 — P. kaszabi; 10 — P. zernyi
groups based on the wing pattern: costal margin of wings with dark spots rounded or subquadrate. I also believe that the species of Psacadina form two distinct clades: P. zernyi and P. vittigera + P. kaszabi, the latter two species share the wing pattern and the general structure of genitalia. Unfortunately, the wing pattern is very variable (Figs. 2-3), often even the left and the right wings of the same specimen are significantly different. However, diagnostic use of the wing pattern as recommended in the key above seems reliable, since all specimens I had preliminary diagnosed as "rounded" or "subquadrate" were confirmed as P. zernyi or P. vittigera + P. kaszabi after examination of their genitalia.
Species level identification of Psacadina is usually based on examination of male genitalia: surstyli (mentioned as gonostyli in the majority of previous publications) and sternite 5.
The surstylus is a small sclerite of a complicated shape which is very differently visible
depending on the angle of view. As a result, authors offer very different drawings of it in their works. To illustrate these difficulties, in Figs. 4-6, I took Rozkosny's (1987, fig. 321) drawing of the male postabdomen of Psaca-dina vittigera, first with the original drawings of the surstyli by Rozkosny and then with the surstyli replaced with their drawings, again of P. vittigera, but taken from illustrations by Vala (1989, fig. 96a) and Rivosecchi (1992, fig. 129c).
How does surstyli of P. vittigera actually look like? I have separated surstyli of the three above considered species of Psacadina and photographed them; the results are shown in Figs. 7-10. I found out that the same surstylus of P. vittigera may either look as on Rozkosny's drawing (compare Figs. 7 and 4) or resemble Vala's or Rivosecchi's drawings (compare Figs. 8 with 5 and 6). The surstylus of P. kaszabi photographed from the same angle of view is similar to that of P. vittigera. The
Figs. 11-13. Sternite 5 of Psacadina, posterior margin above, anterior below: 11 — P zernyi, 12 — P. vittigera; 13 — P kaszabi
Рис. 11-13. Стернит 5 Psacadina, задний край обращен кверху, передний книзу: 11 — P. zernyi; 12 — P. vittigera; 13 — P. kaszabi
Fig. 14. Sternites 4 and 5 of Psacadina vittigera and P disjecta (from Verbeke 1964: figs. 19 and 20) Рис. 14. Стерниты 4 и 5 Psacadina vittigera и P. disjecta (из Verbeke 1964: figs. 19 and 20)
surstylus of P. zernyi looks different, it has an ear-like shape, but depending on the angle of view the "ear" looks either short and rounded or more elongated. Thus, the diagnostic value
of the shape of surstyli of Psacadina seems limited, or at least requires an agreement on unification of the angle of view.
Fortunately, it is possible not to use the shape of surstyli at all to distinguish Psaca-dina species. In contrast with the difficulties discussed above, the shape of sternite 5 makes it easy to make unambiguous conclusions almost regardless of the angle of view. P. zernyi has sternite 5 narrow, elongated, with curved posteriorly apices (Fig. 11). The ster-nite 5 of P. vittigera is crescent-shaped with a rounded projection on the posterior margin and a notch on the anterior margin (Fig. 12). The sternite 5 of P. kaszabi has a notch on the posterior margin and a long projection on the anterior margin (Fig. 13).
SYNONYMY. P. disjecta Enderlein, 1939. This species was described from two males from Spain and Estonia. The type series was reexamined by Verbeke (1964) who found that the Estonian specimen belongs to P. zernyi and is not conspecific to the Spanish one. According to Verbeke, the Spanish syn-type belongs to a different species related to P. vittigera. Verbeke (1964) also identified and listed one more specimen from Morocco and several specimens from Spain (Almeria Prov.; Tijola) as P. disjecta; one male from the Spanish series is stored in ZIN (see material P. vittigera above). The only comprehensible difference between P. disjecta and widespread
P. vittigera is the shape of sternites 4 and 5 as shown in Fig. 14.
Since 1964 other authors have repeated this recommendation (see, for example, Rivo-secchi (1992: 213)), but I found it incorrect.
(1) Sternite 4 of P. vittigera does not look as proposed in Fig. 14, but is deeply emarginated at the posterior margin, exactly as proposed by Verbeke for P. disjecta.
(2) The notch on the anterior margin of sternite 5 may be visible or not depending on the angle of view.
(3) I examined the male from Spain, Almeria Prov., Tijola in ZIN. Its genitalia were not dissected, but there is an identification label by J. Verbeke, 1966: "P disjecta Enderlein". However, the postabdomen is slightly bent and the posterior margin of sternite 5 with a rounded projection is visible in the intact specimen.
Thus, at least one specimen identified by Verbeke as P. disjecta is P. vittigera and the proposed difference in the shape of sternites was not justified, so, P. vittigera Schiner, 1864 = P. disjecta Enderlein, 1939, syn. nov.
P. verbekei Rozkosny, 1975 in Knutson et al. 1975. During a long time Psacadina with rounded spots along costal margin was known under the name Psacadina punctata (Fabricius, 1794) (= Musca punctata Fabricius, 1794 = Pherbina punctata (Fabricius, 1794)). Rozkosny (Knutson et al. 1975) reexamined the type of Musca punctata Fabricius, 1794 and found out that it actually belonged to another species, so what was understood as Psacadina punctata was a nameless species. Therefore, he introduced a
new name Psacadina verbekei Rozkosny, 1975. Besides, Musca punctata Fabricius, 1794 is a junior primary homonym of Musca punctata Poda, 1761 (Thompson et Pont 1993); thus, the Fabricius's species is presently known under the valid available name Pherbellia schoenherri Fallen, 1826.
Earlier Mayer (1953) described Psacadina zernyi which "is closely related to P punctata and differs from the latter only due to slightly different form of surstylus".
However, I find Mayer's description groundless, because the shape of surstylus looks different depending on the angle of view. For example, the most realistic drawings with a comparison of the surstylus and ster-nite 5 of P. zernyi and P. verbekei were given by Vala (1989, figs. 96c, d, e, f) and Rivosecchi (1992, figs. 127, 128). I found out that the same surstylus and sternite 5 can be rotated
so that they look like P. zernyi or P. verbekei. Hence P zernyi and P verbekei are the same species. Therefore, Psacadina zernyi Mayer, 1953 becomes the oldest available, not preoccupied and hence valid name of this species: P zernyi Mayer, 1953 = P verbekei Rozkosny, 1975, syn. nov.
With the two synonyms newly proposed here, the genus Psacadina is no longer problematic, and the three remaining valid species may be reliably distinguished in both sexes as recommended in the key.
Acknowledgements
I thank Olga Ovchinnikova and Galina Su-leymanova (Saint Petersburg) for the opportunity to examine the important material in ZIN. I thank Oleg Kosterin (Novosibirsk) and Maria Yanbulat (Moscow) for useful discussion and correcting the text.
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For citation: Vikhrev, N. E. (2022) Notes on the genus Psacadina (Diptera, Sciomyzidae). Amurian Zoological Journal, vol. XIV, no. 4, pp. 562-569. https://www.doi.org/10.33910/2686-9519-2022-14-4-562-569
Received 28 September 2022; reviewed 3 November 2022; accepted 8 November 2022.
Для цитирования: Вихрев, Н. Е. (2022) Заметки роду Psacadina (Diptera, Sciomyzidae). Амурский зоологический журнал, т. XIV, № 4, с. 562-569. https://www.doi.org/10.33910/2686-9519-2022-14-4-562-569 Получена 28 сентября 2022; прошла рецензирование 3 ноября 2022; принята 8 ноября 2022.
