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Амурский зоологический журнал, 2021, т. XIII, № 1
Amurian Zoological Journal, 2021, vol. XIII, no. 1
www.azjournal.ru
UDC 595.76:502.72 https://www.doi.org/10.33910/2686-9519-2021-13-l-12-35
http://zoobank.org/References/BCBB2C54-12EE-46BB-97E7-843E7317AClB
Notes on the fauna of beetles (Insecta, Coleoptera) adjacent to the territory of the Mordovia State Nature Reserve
A. B. Ruchin1®, L. V. Egorov1' 2, S. K. Alekseev3, G. B. Semishin1, M. N. Esin1
1 Joint Directorate of the Mordovia State Nature Reserve and National Park "Smolny" 30, Krasnaya Str., 430005, Saransk, Russia 2 State reserve "Prisursky", 9, Lesnoy av., 428034, Cheboksary, Russia 3 Ecological club "Stenus', 4, Staroobryadcheskii Lane, 248023, Kaluga, Russia
Abstract. The paper presents the results of the study of findings made in the adjacent of the Mordovia State Nature Reserve (Republic of Mordovia, Russia) in 2012-2019 (partly, from previous years). The geography of the collection includes 6 districts of the Nizhny Novgorod Region (Vyksa, Voznesenskoe, and Pervomaisk districts as well as Sarov and its surroundings) and the Republic of Mordovia (Temnikov, Elniki, and Tengushevo districts). Collection was carried out manually as well as by soil traps and crown traps with fermenting baits. In total, 260 species from 29 families of Coleopterans were found.
Authors
Alexaner B. Ruchin
E-mail: ruchin.alexander@gmail.com
SPIN: 1655-5762
Scopus Author ID: 6602618456
ResearcherlD: AAY-6928-2020
ORCID: 0000-0003-2653-3879
Leonid V. Egorov
E-mail: platyscelis@mail.ru
SPIN: 6331-1661
Scopus Author ID: 1592504730
ResearcherID: R-2054-2019
ORCID: 0000-0002-3680-8891
Sergei K. Alekseev
E-mail: stenus@yandex.ru
RSCI AuthorID: 141079
Scopus Author ID: 57207825754
Gennadiy B. Semishin E-mail: g.semishin@mail.ru Mikhail N. Esin E-mail: esinmishka@gmail.com
Copyright: © The Authors (2021). Published by Herzen State Pedagogical University of Russia. Open access under CC BY-NC License 4.0.
Keywords: beetles, Coleoptera, fauna, biodiversity, Mordovia State Nature Reserve, Republic of Mordovia, Nizhny Novgorod Region.
Материалы к фауне жесткокрылых (Insecta, Coleoptera) окрестностей Мордовского заповедника
А. Б. Ручинш, Л. В. Егоров1, 2, С. К. Алексеев3, Г. Б. Семишин1, М. Н. Есин1
1 Объединенная дирекция Мордовского государственного природного заповедника имени П. Г. Смидовича и национального парка «Смольный», ул. Красная, д. 30, 430005, Саранск, Россия
2 Государственный заповедник «Присурский», пос. Лесной, д. 9, 428034, Чебоксары, Россия
3 Экологический клуб <^епш», Старообрядческий пер., д. 4, 248023, Калуга, Россия
Сведения об авторах Ручин Александр Борисович E-mail: ruchin.alexander@gmail.com SPIN-код: 1655-5762 Scopus Author ID: 6602618456 ResearcherlD: AAY-6928-2020 ORCID: 0000-0003-2653-3879 Егоров Леонид Валентинович E-mail: platyscelis@mail.ru SPIN-код: 6331-1661 Scopus Author ID: 1592504730 ResearcherID: R-2054-2019 ORCID: 0000-0002-3680-8891 Алексеев Сергей Константинович E-mail: stenus@yandex.ru РИНЦ AuthorID: 141079 Scopus Author ID: 57207825754
Семишин Геннадий Борисович E-mail: g.semishin@mail.ru Есин Михаил Николаевич E-mail: esinmishka@gmail.com Права: © Авторы (2021). Опубликовано Российским государственным педагогическим университетом им. А. И. Герцена. Открытый доступ на условиях лицензии CC BY-NC 4.0.
Аннотация. Приводятся результаты изучения сборов 2012-2019 гг. (частично — предыдущих лет) на территориях, прилегающих к Мордовскому заповеднику (Республика Мордовия, Россия). География сборов включает 6 районов Нижегородской области (Выксунский, Вознесенский, Первомайский районы и г. Саров с окрестностями) и Республики Мордовия (Темниковский, Ельниковский, Теньгушевский районы). Сборы осуществлялись вручную, а также с помощью почвенных и кроновых ферментных ловушек. В общей сложности обнаружено 260 видов из 29 семейств жесткокрылых.
Ключевые слова: жесткокрылые, Со1еор1ега, фауна, биоразнообразие, Мордовский заповедник, Республика Мордовия, Нижегородская область.
Introduction
Biodiversity conservation is a very important issue, especially, in the face of challenges posed by envir onmental pollution, climate change, and urbanization (Oppenheimer 1989; Bengtsson et al. 2000; Dossou, Gle-houenou-Dossou 2007; Kovats, Akhtar 2008; Gebremedihin et al. 2018; Kopoteva, Kuptso-va 2019). It is generally accepted that the protection of life on Earth is not possible through individual efforts of dedicated teams — it is a responsibility of humanity at large and, at the same time, a condition of our survival on the planet. The most effective way to preserve biological diversity is through specially protected areas (PA) (Akpatou et al 2018; Xavier da Silva et al. 2018). Russia has several categories of protected areas with reserves and national parks subject to the strictest regulation. They are designed to preserve biodiversity in certain areas of land and sea (Grebennikov 2016; Markevich, Esin 2018; Bakirova, Zharkikh 2019; Kirilyuk et al. 2019; Uligova et al. 2019).
The Mordovia State Nature Reserve is sur-rouned by a diversity of landscapes and interesting biotopes. The coleopterofauna found in these areas has also been studied to a certain extent (Ruchin et al. 2013; 2018; 2019c; Ruchin, Egorov 2018e). Border landscapes are also habitats for a range of species, and, in some cases, they are home to certain ecological groups. For example, the forest area of the reserve includes few meadow (open) stations, while open biotopes are widespread at the edge of the reserve. The Mordovia State Nature Reserve is located in the Temnikov District of the Republic of Mordovia (Fig. 1) on the wooded right bank of the Moksha river. The Reserve covers an area of 321,62 km2. This is one of the first nature reserves in Russia. According to paheoecological reconstructions, broad-leaved forests have dominated suitable habitats in the reserve since 5,000 years ago. The Western border is along the Cher-naya, Satis and Moksha rivers. Forest-steppe near the southern border naturally delineates the border of the protected area (Khapugin, Ruchin 2019). The forest structure of the reserve includes coniferous-broadleaf forests
with forest-steppe at its edges. Forest communities cover 89,3% of the total territory. In general, the herb layer of the Mordovia State Nature Reserve has plants typical for taiga. However, it also includes species of flora common for broad-leaved forests. Pine (Pinus sylvestris L.) is the main forest-forming species in the reserve. It forms pure or mixed plant communities in the Southern, Central and Western parts of the Mordovia State Nature Reserve. Birch (Betula péndula Roth) occupies the second largest forest area in the reserve. These are mainly secondary post-fire and post-felling communities that have replaced pine forests. There are especially many young birch trees in forest areas heavily damaged by fire in 2010 (Ruchin et al. 2019a). Lime forest is found mainly in the Northern part of the Mordovia State Nature Reserve. These are secondary plant communities that have emerged on the site of pine and lime-spruce forests. Spruce (from Picea abies L.) and alder trees (from Alnus glutinosa (L.) Gaertn.) are located mainly in floodplains of rivers and streams (Pushta, Vyaz-Pushta, Vorsklyai, Arga, etc.) and occupy small areas. Vegetation communities of small-leaved species (birch, aspen, alder) are formed in post-fire areas (Khapugin et al. 2016). The reserve borders the Nizhny Novgorod Region in the North (Pervomaisk, Diveevo, and Voznesen-skoe districts, and Sarov). The rest of the border passes through the Republic of Mordovia (Temnikov and Yelniki districts).
The diversity of Coleoptera (Insecta, Coleoptera) of the Mordovia State Nature Reserve has been studied since its foundation (Redikortsev 1938). However, the last 15 years have showed an ever-increasing interest in research (Egorov, Ruchin 2012; 2014; Egorov et al. 2010; 2015; 2016; 2017; 2018; 2019; 2020b). The studies explore separate families (Feoktistov 2008; Semenov 2015; 2016; 2017; Ruchin et al. 2018; 2019; Kovalev, Egorov 2017; Mandelshtam, Egorov 2018; 2019) as well as some of the most interesting species (Egorov 2017; Egorov, Shapovalov 2017; Ruchin, Egorov 2018a; 2018b; 2018c; 2018d; 2018f; Tomaszewska et al. 2018; Kazantsev et
Fig. 1. Location map of the Mordovia State Nature Reserve
Рис. 1. Карта расположения Мордовского государственного природного заповедника
al. 2019; Ruchin et al. 2019a, 2019b), including those from the Red Books (Ruchin, Kur-maeva 2010; Ruchin, Egorov 2017; Ruchin, Khapugin 2019). To date, according to our data, 2,145 species from 88 Coleoptera families have been recorded from the Mordovia State Nature Reserve (Egorov et al. 2020a).
This article provides additional information on the Coleoptera fauna of the surrounding area of the Mordovia State Nature Reserve, including the territories of the Nizhny Novgorod Region and the Republic of Mordovia.
Material and methods
The material was collected in 2012-2019 using entomological methods of field research (Fasu-lati 1971) and crown traps with fermenting baits (Ruchin et al. 2020). The paper also publishes previously unreleased results from earlier years.
The paper provides an annotated list of discovered species (see below). For each of the species the list specifies the location of collection (previously unpublished data), dates of collection, biotopes (upon the availability of data), the number of collected specimens, names of collectors (AR — A. B. Ruchin, GS — G. B. Semi-
shin, ME — M. N. Esin, PC — P. V. Cherenkov, OA — O. N. Artaev, DK — D. K. Kurmaeva).
The Coleoptera system, volume, and nomenclature of most taxa are taken primarily from the Catalogue of Coleoptera of the Palearctic (Löbl, Smetana 2007; 2008; 2010; Löbl, Löbl 2015; 2016; 2017 ); the volume of families and nomenclature of Curculionoidea-from, from (Alonso-Zarazaga et al. 2017). Most species were defined by L. V. Egorov (not marked in the text) and S. K. Alekseev (most of the Carabidae, marked in the text). The sequence of taxa within families is given in alphabetical order. The studied material is partially stored in the collection of the reserve (Pushta). Abbreviations used in the text: RM — Republic of Mordovia, NNR — Nizhny Novgorod Region, district — distr, vill. — vilage, sett. — settlement, ex. — exemplar.
ANNOTATED LIST
Coleoptera
Adephaga
Carabidae
Acupalpus meridianus (Linnaeus, 1760) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 12 ex., det. S. K. Alekseev (OA); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex., det. S. K. Alekseev (ME).
Agonum fuliginosum (Panzer, 1809) Material: NNR: Sarov, 05-08.2013, 5 ex. (OA).
Agonum gracilipes (Duftschmid, 1812) Material: RM: Temnikov distr., Mikhailovka vill., 06-07.2012, coppice, 1 ex., det. S. K. Alekseev (OA). Elniki distr., Peredovoi sett., 05-09.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev (OA); Novye Pichingushi vill., 04-06.2014, birch planting, 11 ex., det. S. K. Alekseev (OA).
Agonum sexpunctatum (Linnaeus, 1758) Material: RM: Temnikov distr., Veselyi sett., 25.05.2013, mixed forest, 1 ex., det. S. K. Alekseev (AR). Elniki distr., Starye Shaly vill., 05-06.2014, pine forest, 2 ex., det. S. K. Alekseev (OA). NNR: Sarov, northeast, on wet sand in a ditch on the side of a highway running through the woods, 26.05.2018, 12.06.2018, 2 ex. (PC).
Agonum versutum Sturm, 1824 Material: RM: Temnikov distr., Andreevka vill., 11.05.2013, the edge of the forest, 1 ex., det. S. K. Alekseev (AR).
Amara aenea (De Geer, 1774) Material: RM: Elniki distr., Svobod-nyi sett., 04-06.2014, disturbed meadow, 18 ex., det. S. K. Alekseev; Peredovoi sett., 05-09.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev; Novye Pichingushi vill., 0406.2014, birch planting, 8 ex., det. S.K. Alekseev; Novye Shaly vill., 05-06.2014, young pine forest, 1 ex., det. S. K. Alekseev (OA); Cherlyai vill., 15.05-28.06.2019, deciduous forest, 4 ex. (ME); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex. (ME). NNR: Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 1 ex. (OA).
Amara aulica (Panzer, 1796) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 3 ex., det. S. K. Alekseev (OA).
Amara bifrons (Gyllenhal, 1810) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 1 ex., det. S. K. Alekseev (OA); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex. (ME).
Amara communis (Panzer, 1797) Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 11 ex., det. S. K. Alekseev (ME). Elniki distr., Peredovoi sett., 05-09.2014, broad-leaved forest, 3 ex., det. S. K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 7 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev (OA); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 57 ex. (ME); Cherlyai vill., 31.07-16.09.2019, deciduous forest, 1 ex. (ME). NNR: Pervo-maisk distr., 4 km S Nikolaevka vill., 30.0414.07.2014, 7 ex. (OA). Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow, 5 ex., det. S. K. Alekseev (ME).
Amara consularis (Duftschmid, 1812) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 5 ex., det. S. K. Alekseev (OA).
Amara curta Dejean, 1828 Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 2 ex., det. S. K. Alekseev (OA).
Amara eurynota (Panzer, 1796) Material: RM: Elniki distr., Novye Pichingushi vill., 04-06.2014, birch planting, 4 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev (OA); Cherlyai vill., 15.0528.06.2019, deciduous forest, 1 ex. (ME).
Amara familiaris (Duftschmid, 1812) Material: RM: Elniki distr., Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 1 ex., det. S. K. Alekseev (OA).
Amara fulva (O.F. Müller, 1776) Material: NNR: Vyksa distr., NE Vy-ksa, dirt (sandy) road to Rizadeevskoe lake, 13.08.2017, 1 ex. (PC). Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow,
1 ex., det. S. K. Alekseev (ME).
Amara ingenua (Duftschmid, 1812) Material: NNR: Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow,
2 ex., det. S. K. Alekseev (ME).
Amara ovata (Fabricius, 1792). Material: RM: Temnikov distr., Andreevka vill., 11.05.2013, the edge of the forest, 1 ex. (AR).
Amara majuscula (Chaudoir, 1850) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 1 ex., det. S. K. Alekseev (OA).
Amara ovata (Fabricius, 1792) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 12 ex., det. S. K. Alekseev (OA). NNR: Voznesen-skoe distr., Baranovka vill., 16.05-18.06.2019, meadow, 1 ex., det. S. K. Alekseev (ME).
Amara similata (Gyllenhal, 1810) Material: RM: Elniki distr., Novye Shaly vill., 05-06.2014, young pine forest, 2 ex., det. S. K. Alekseev (OA). NNR: Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 1 ex., det. S. K. Alekseev (OA).
Amara tibialis (Paykull, 1798) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 5 ex., det. S. K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 3 ex.; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 2 ex., det. S. K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 2 ex., det. S. K. Alekseev (OA).
Anchomenus dorsalis (Pontoppidan, 1763) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 15 ex., det. S. K. Alekseev (OA).
Anisodactylus binotatus (Fabricius, 1787) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 12 ex., det. S. K. Alekseev; Peredovoi sett., 05-09.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev (OA). Anisodactylus nemorivagus (Duftschmid, 1812)
Material: RM: Temnikov distr., Kitsaev-ka vill., 12.05.2013, mixed forest, 1 ex. (AR). NNR: Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 1 ex., det. S. K. Alek-seev (OA).
Anisodactylus signatus (Panzer, 1796) Material: RM: Temnikov distr., Kitsaevka vill., 12.05.2013, mixed forest, 1 ex., det. S. K. Alekseev (AR). Elniki distr., Novye Pichingushi vill., 04-06.2014, birch planting, 1 ex., det. S. K. Alekseev (OA).
Asaphidion flavipes (Linnaeus, 1760) Material: RM: Elniki distr., Malye Mordovs-kie Poshaty vill., 05-06.2014, broad-leaved forest, 4 ex., det. S. K. Alekseev (OA). NNR: Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 1 ex., det. S. K. Alekseev (OA). Badister bullatus (Schrank, 1798) (=bipustu-latus (Fabricius, 1792) nec (Fabricius, 1775)) Material: RM: Temnikov distr., Mikhailovka vill., 06-07.2012, coppice, 1 ex., det. S. K. Alekseev (OA). Elniki distr., Peredovoi sett., 05-09.2014, broad-leaved forest, 2 ex., det. S. K. Alekseev; Novye Shaly vill., 05-06.2014, young pine forest, 2 ex., det. S. K. Alekseev (OA).
Badister collaris Motschulsky 1844 Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 1 ex., det. S. K. Alekseev (OA).
Badister lacertosus Sturm, 1815 Material: RM: Elniki distr., Starye Shaly vill., 05-06.2014, pine forest, 5 ex., det. S. K. Alekseev; Svobodnyi sett., 04-06.2014, disturbed meadow, 1 ex., det. S. K. Alekseev; Peredovoi sett., 05-09.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev; Novye Pichin-gushi vill., 04-06.2014, birch planting, 1 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 2 ex., det. S. K. Alekseev; Novye Shaly vill., 0506.2014, young pine forest, 2 ex. (OA); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 11 ex. (ME). NNR: Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 2 ex., det. S. K. Alekseev (OA).
Badister meridionalis Puel, 1925 Material: NNR: Sarov, 05-08.2013, 1 ex. (OA).
Bembidion articulatum (Panzer, 1796) Material: NNR: Sarov, northeast, on wet sand in a ditch on the side of a highway running through the woods, 9.06.2017, 1 ex. (PC).
Bembidion gilvipes Sturm, 1825 Material: RM: Elniki distr., Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex. (ME).
Bembidion lampros (Herbst, 1784) Material: RM: Elniki distr., Malye Mordov-skie Poshaty vill., 05-06.2014, broad-leaved forest, 41 ex., det. S. K. Alekseev (OA); Sta-rye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 2 ex. (ME).
Bembidion obliquum Sturm, 1825 Material: NNR: Vyksa distr., Vyksa, Ri-zadeevskoe lake, near a puddle, 29.07.2018, 1 ex. (PC).
Bembidion properans (Stephens, 1828) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 32 ex., det. S. K. Alekseev; Peredovoi sett., 05-09.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 9 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 5 ex., det. S. K. Alekseev (OA); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 10 ex. (ME).
Bembidion quadrimaculatum (Linnaeus, 1760) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 33 ex., det. S. K. Alekseev (OA).
Blemus discus (Fabricius, 1792) Material: NNR: Sarov, northeast, on wet sand in a ditch on the side of a highway running through the woods, 17.08.2018, 1 ex. (PC).
Broscus cephalotes (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Ala-masovo vill., on a dacha plot, in the ground, 6.06.2014, 1.05.2015, 2 ex. (PC).
Calathus ambiguus (Paykull, 1790) Material: RM: Elniki distr., Novye Pichingushi vill., 04-06.2014, birch planting, 89 ex., det. S. K. Alekseev (OA).
Calathus erratus (C.R. Sahlberg, 1827) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 25 ex., det. S. K. Alekseev; Malye Mordovs-kie Poshaty vill., 05-06.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev (OA). NNR: Sarov, 05-08.2013, 2 ex., det. S. K. Alekseev (OA). Voznesenskoe distr., Shaprikha vill., 16.05-18.06.2019, mixed forest, 5 ex., det. S. K. Alekseev (ME).
Calathus fuscipes (Goeze, 1777) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 6 ex., det. S. K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 139 ex., det. S. K. Alekseev (OA).
Calathus melanocephalus (Linnaeus, 1758) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 5 ex., det. S. K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 6 ex., det. S . K. Alekseev (OA); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex. (ME). NNR: Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow, 3 ex., det. S. K. Alekseev (ME).
Calathus micropterus (Duftschmid, 1812) Material: RM: Elniki distr., Starye Shaly vill., 05-06.2014, pine forest, 2 ex.; Novye Shaly vill., 05-06.2014, young pine forest, 6 ex., det. S. K. Alekseev; Peredovoi sett., 05-
09.2014, broad-leaved forest, 3 ex., det. S. K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 2 ex., det. S. K. Alekseev; Ma-lye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 7 ex., det. S. K. Alek-seev (OA). NNR: Sarov, 05-08.2013, 28 ex., det. S. K. Alekseev (OA).
Calosoma inquisitor (Linnaeus, 1758) Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest,
I ex., det. S. K. Alekseev (ME). Elniki distr., Cherlyai vill., 15.05-28.06.2019, deciduous forest, 1 ex. (ME).
Carabus arvensis Herbst, 1784 Material: RM: Elniki distr., Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 4 ex., det. S.K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 30 ex., det. S. K. Alek-seev (OA). Tengushevo distr., Klemeschshei vill., 14.05-21.06.2019, mixed forest, 51 ex., det. S. K. Alekseev (ME).
Carabus cancellatus Illiger, 1798 Material: RM: Elniki distr., Malye Mor-dovskie Poshaty vill., 05-06.2014, broad-leaved forest, 4 ex., det. S.K. Alekseev; Pere-dovoi sett., 05-09.2014, broad-leaved forest,
II ex., det. S. K. Alekseev (OA); Cherlyai vill., 15.05-28.06.2019, deciduous forest, 7 ex. (ME). NNR: Voznesenskoe distr., Alamasovo vill., 17.10.2015, 30.04.2016, 16.08.2016, 3 ex. (PC); Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 7 ex., det. S. K. Alek-seev (OA).
Carabus clathratus Linnaeus, 1760 Material: RM: Tengushevo distr., Khlebi-no vill., 14.05-20.06.2019, meadow, 1 ex., det. S. K. Alekseev (ME).
Carabus convexus Fabricius, 1775 Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 5 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 2 ex., det. S. K. Alekseev (OA).
Carabus coriaceus Linnaeus, 1758 Material: RM: Elniki distr., Malye Mordov-skie Poshaty vill., 05-06.2014, broad-leaved forest, 2 ex., det. S. K. Alekseev (OA); Cherlyai vill., 31.07-16.09.2019, deciduous forest, 1 ex.
(ME). NNR: Voznesenskoe distr., Alamaso-vo vill., mixed forest, 13.09.2015, 1 ex. (PC); Sarov, 05-08.2013, 3 ex., det. S. K. Alekseev (OA).
Carabus glabratus Paykull, 1790 Material: RM: Elniki distr., Malye Mordo-vskie Poshaty vill., 05-06.2014, broad-leaved forest, 11 ex., det. S. K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 3 ex., det. S. K. Alekseev; Peredovoi sett., 05-09.2014, broad-leaved forest, 4 ex., det. S. K. Alekseev (OA). Temnikov distr., Andreevka vill., 14.0521.06.2019, mixed forest, 2 ex. (ME). NNR: Sarov, 05-08.2013, 13 ex., det. S. K. Alekseev (OA). Voznesenskoe distr., Shaprikha vill., 16.05-18.06.2019, mixed forest, 1 ex., det. S. K. Alekseev (ME).
Carabus granulatus Linnaeus, 1758 Material: RM: Elniki distr., Starye Shaly vill., 05-06.2014, pine forest, 1 ex., det. S. K. Alek-seev; Peredovoi sett., 05-09.2014, broad-leaved forest, 7 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev (OA). NNR: Sarov, 05-08.2013, 35 ex., det. S. K. Alek-seev (OA). NNR: Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 21 ex., det. S. K. Alekseev (OA).
Carabus hortensis Linnaeus, 1758 Material: RM: Temnikov distr., Kitsaev-ka vill., 12.05.2013, mixed forest, 1 ex. (AR); Mikhailovka vill., 06-07.2012, coppice, 9 ex., det. S. K. Alekseev (OA); Andreevka vill., 31.07-16.09.2019, mixed forest, 1 ex. (ME). Elniki distr., Novye Shaly vill., 05-06.2014, young pine forest, 5 ex., det. S. K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 5 ex., det. S. K. Alekseev; Malye Mordovs-kie Poshaty vill., 05-06.2014, broad-leaved forest, 26 ex., det. S. K. Alekseev; Peredovoi sett., 05-09.2014, broad-leaved forest, 6 ex., det. S. K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 5 ex., det. S . K . Alekseev (OA). NNR: Voznesenskoe distr., Alamasovo vill., 17.08.2014, 1 ex. (PC); Sarov, 05-08.2013, 11 ex., det. S. K. Alekseev (OA); Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 1 ex., det. S. K. Alek-seev (OA).
Cicindela campestris Linnaeus, 1758 Material: RM: Temnikov distr., Kitsaevka vill., 12.05.2013, mixed forest, 2 ex. (AR). Ten-gushevo distr., Khlebino vill., 14.05-20.06.2019, meadow, 1 ex., det. S. K. Alekseev (ME). Elniki distr., Cherlyai vill., 15.05-28.06.2019, deciduous forest, 3 ex. (ME). NNR: Voznesenskoe distr., Alamasovo vill., 30.05.2015, 14.06.2015, 30.04.2017, 20.05.2017, 6 ex. (PC); Baranovka vill., 16.05-18.06.2019, meadow, 4 ex., det. S. K. Alekseev (ME).
Cicindela hybrida Linnaeus, 1758 Material: RM: Temnikov distr., Andreevka vill., 11.05.2013, the edge of the forest, 1 ex. (AR). NNR: Sarov, 2.06.2015, 10.07.2015,
18.04.2016, 4.06.2016, 4 ex.; Voznesenskoe distr., Alamasovo vill., 25.07.2015, 2.08.2016,
29.04.2017, 3 ex. (PC).
Cicindela sylvatica Linnaeus, 1758 Material: RM: Temnikov distr., Purdoshki vill., 15.08.2015, 1 ex. (AR). Elniki distr., Novye Shaly vill., 26.05.2013, pine forest, 1 ex. (AR). NNR: Voznesenskoe distr., Alamasovo vill.,
18.07.2015, 19.07.2015, 25.07.2015, 9.05.2016,
30.07.2016, 29.04.2017, 2.05.2018, 7 ex. (PC). Clivina fossor (Linnaeus, 1758)
Material: NNR: Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 1 ex., det. S. K. Alekseev (OA).
Cychrus caraboides (Linnaeus, 1758) Material: NNR: Sarov, 05-08.2013, 4 ex., det. S. K. Alekseev (OA). Cylindera arenaria viennensis (Schrank, 1781) Material: NNR: Vyksa, 17.08.2017,
18.08.2017, 31.07.2018, 4 ex. (PC).
Note. Psammophile. This is a very rare local species. It inhabits floodplain areas with extensive sandy banks and sand and mud banks (Gebert 2014), open sandy areas of anthropogenic origin (Spitzer et al. 2014). Cymindis angularis Gyllenhal, 1810 Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 5 ex., det. S. K. Alekseev (OA).
Dolichus halensis (Schaller, 1783) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 3 ex., det. S. K. Alekseev (OA).
Dromius schneideri Crotch, 1871 Material: NNR: Voznesenskoe distr., Ala-masovo vill., 14.05.2016, 1 ex. (PC).
Elaphrus cupreus Duftschmid, 1812 Material: RM: Tengushevo distr., Kle-meschshei vill., 14.05-21.06.2019, mixed forest, 1 ex., det. S. K. Alekseev (ME). NNR: Voznesenskoe distr., Alamasovo vill., 11.06.2017, 1 ex. (PC); Sarov, 05-08.2013,
3 ex., det. S. K. Alekseev (OA).
Elaphrus riparius (Linnaeus, 1758) Material: NNR: Sarov, northeast, on wet sand in a ditch on the side of a highway running through the woods, 9.06.2017, 12.06.2017, 20.06.2017, 3 ex. (PC).
Harpalus affinis (Schrank, 1781) Material: RM: Temnikov distr., Mikhailovka vill., 06-07.2012, coppice, 1 ex., det. S. K. Alek-seev (OA). Elniki distr., Novye Pichingushi vill., 04-06.2014, birch planting, 26 ex., det. S.K. Alekseev; Svobodnyi sett., 04-06.2014, disturbed meadow, 3 ex., det. S. K. Alekseev (OA). NNR: Sarov, 05-08.2013, 1 ex., det. S. K. Alekseev (OA). Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow,
4 ex., det. S. K. Alekseev (ME). Harpalus distinguendus (Duftschmid, 1812)
Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 11 ex., det. S. K. Alekseev; Svobodnyi sett., 04-06.2014, disturbed meadow, 4 ex., det. S. K. Alekseev (OA).
Harpalus griseus (Panzer, 1796) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 7 ex., det. S. K. Alekseev (OA).
Harpalus laevipes Zetterstedt, 1828 Material: RM: Elniki distr., Peredovoi sett., 05-09.2014, broad-leaved forest, 2 ex., det. S. K. Alekseev; Novye Pichingushi vill.,
04-06.2014, birch planting, 1 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill.,
05-06.2014, broad-leaved forest, 6 ex., det. S. K. Alekseev; Novye Shaly vill., 05-06.2014, young pine forest, 8 ex., det. S. K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 9 ex., det. S. K. Alekseev (OA). NNR: Sarov, 05-08.2013, 9 ex., det. S. K. Alekseev
(OA). Voznesenskoe distr., Shaprikha vill., 16.05-18.06.2019, mixed forest, 3 ex., det. S. K. Alekseev (ME).
Harpalus latus (Linnaeus, 1758) Material: RM: Temnikov distr., Mikhailovka vill., 06-07.2012, coppice, 1 ex., det. S. K. Alekseev (OA); Andreevka vill., 31.07-16.09.2019, mixed forest, 7 ex. (ME). Tengushevo distr., Klemeschshei vill., 14.05-21.06.2019, mixed forest, 5 ex., det. S. K. Alekseev (ME); Khlebi-no vill., 14.05-20.06.2019, meadow, 22 ex., det. S. K. Alekseev (ME). Elniki distr., Pere-dovoi sett., 05-09.2014, broad-leaved forest,
1 ex.; Novye Pichingushi vill., 04-06.2014, birch planting, 1 ex., det. S. K. Alekseev; Ma-lye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 4 ex., det. S. K. Alek-seev (OA); Cherlyai vill., 15.05-28.06.2019, deciduous forest, 5 ex. (ME); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 17 ex. (ME). NNR: Sarov, 05-08.2013, 2 ex., det. S. K. Alekseev (OA). Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 21 ex. (OA).
Harpalus picipennis (Duftschmid, 1812) Material: NNR: Voznesenskoe distr., Shaprikha vill., 16.05-18.06.2019, mixed forest,
2 ex., det. S. K. Alekseev (ME). Harpalusprogrediens Schauberger, 1922 Material: RM: Temnikov distr., Mikhailov-
ka vill., 06-07.2012, coppice, 1 ex., det. S. K. Alekseev (OA). Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 7 ex., det. S. K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 2 ex., det. S. K. Alekseev (OA). NNR: Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow, 6 ex., det. S. K. Alekseev (ME).
Harpalus pumilus Sturm, 1818 Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 3 ex., det. S. K. Alekseev (OA).
Harpalus rubripes (Duftschmid, 1812) Material: RM: Tengushevo distr., Khlebi-no vill., 14.05-20.06.2019, meadow, 2 ex., det. S. K. Alekseev (ME). Elniki distr., Svobod-nyi sett., 04-06.2014, disturbed meadow, 11 ex., det. S. K. Alekseev; Peredovoi sett., 05-
09.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev; Novye Shaly vill., 05-06.2014, young pine forest, 1 ex., det. S. K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 1 ex., det. S. K. Alekseev (OA); Starye Russk-ie Poshaty vill., 15.05-28.06.2019, meadow, 9 ex. (ME). NNR: Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 1 ex., det. S. K. Alekseev (OA). Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow, 4 ex., det. S. K. Alekseev (ME).
Harpalus rufipes (De Geer, 1774) Material: RM: Temnikov distr., Mikhailov-ka vill., 06-07.2012, coppice, 39 ex., det. S. K. Alekseev (OA). Elniki distr., Novye Pichingushi vill., 04-06.2014, birch planting, 378 ex., det. S. K. Alekseev; Peredovoi sett., 05-09.2014, broad-leaved forest, 23 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 4 ex., det. S. K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 1 ex., det. S. K. Alek-seev (OA); Cherlyai vill., 15.05-28.06.2019, deciduous forest, 2 ex. (ME); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 5 ex. (ME). NNR: Voznesenskoe distr., Alamasovo vill., on a dacha plot, in the ground, 7.09.2014, 16.08.2015, 2 ex. (PC); Baranovka vill., 16.0518.06.2019, meadow, 5 ex., det. S. K. Alekseev (ME); Shaprikha vill., 16.05-18.06.2019, mixed forest, 4 ex., det. S. K. Alekseev (ME). Sarov, 05-08.2013, 2 ex., det. S. K. Alekseev (OA).
Harpalus signaticornis (Duftschmid, 1812) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 1 ex., det. S. K. Alekseev (OA).
Harpalus smaragdinus (Duftschmid, 1812) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 1 ex., det. S. K. Alekseev (OA).
Harpalus tardus (Panzer, 1796) Material: RM: Temnikov distr., Kitsaevka vill., 12.05.2013, mixed forest, 1 ex. (AR). Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 10 ex., det. S. K. Alekseev; Malye Mor-dovskie Poshaty vill., 05-06.2014, broad-leaved
forest, 4 ex., det. S. K. Alekseev (OA). NNR: Voznesenskoe distr., Baranovka vill., 16.0518.06.2019, meadow, 3 ex., det. S. K. Alekseev (ME); Shaprikha vill., 16.05-18.06.2019, mixed forest, 2 ex., det. S. K. Alekseev (ME). Harpalus xanthopus winkleri Schauberger, 1923 Material: RM: Tengushevo distr., Khlebino vill., 14.05-20.06.2019, meadow, 1 ex., det. S. K. Alekseev (ME). Elniki distr., Malye Mordo-vskie Poshaty vill., 05-06.2014, broad-leaved forest, 2 ex., det. S. K. Alekseev; Novye Shaly vill., 05-06.2014, young pine forest, 2 ex., det. S. K. Alekseev (OA); Cherlyai vill., 15.0528.06.2019, deciduous forest, 2 ex. (ME); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 30 ex. (ME). NNR: Sarov, 05-08.2013, 1 ex., det. S. K. Alekseev (O). Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 1 ex., det. S. K. Alekseev (OA). Lebia chlorocephala (J.J. Hoffmann, 1803) Material: RM: Elniki distr., Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex. (ME).
Lebia cruxminor (Linnaeus, 1758) Material: RM: Elniki distr., Svobodnyi sett.,
04-06.2014, disturbed meadow, 2 ex., det. S. K. Alekseev (OA); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex. (ME).
Leistus terminatus (Panzer, 1793) Material: NNR: Sarov, 05-08.2013, 1 ex., det. S. K. Alekseev (OA).
Licinus depressus (Paykull, 1790) Material: RM: Elniki distr., Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 3 ex. (ME).
Limodromus assimilis (Paykull, 1790) Material: RM: Elniki distr., Peredovoi sett.,
05-09.2014, broad-leaved forest, 5 ex., det. S. K. Alekseev (OA). NNR: Sarov, 05-08.2013, 1 ex., det. S. K. Alekseev (OA).
Loricera pilicornis (Fabricius, 1775) Material: RM: Elniki distr., Starye Russk-ie Poshaty vill., 15.05-28.06.2019, meadow, 2 ex., det. S. K. Alekseev (ME). NNR: Sarov, 05-08.2013, 1 ex., det. S. K. Alekseev (OA). Microlestes maurus (Sturm, 1827) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 26 ex.,
det. S. K. Alekseev; Novye Pichingushi vill.,
04-06.2014, birch planting, 1 ex., det. S. K. Alekseev (OA); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex. (ME).
Microlestes minutulus (Goeze, 1777) Material: RM: Elniki distr., Novye Pich-ingushi vill., 04-06.2014, birch planting, 5 ex., det. S. K. Alekseev; Starye Shaly vill., 0506.2014, pine forest, 1 ex., det. S. K. Alekseev (OA).
Notiophilus aquaticus (Linnaeus, 1758) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 1 ex., det. S. K. Alekseev (OA). NNR: Sarov, 0508.2013, 1 ex., det. S. K. Alekseev (OA). Notiophilus palustris (Duftschmid, 1812) Material: RM: Temnikov distr., Mikhailov-ka vill., 06-07.2012, coppice, 1 ex., det. S. K. Alekseev (OA). Elniki distr., Peredovoi sett.,
05-09.2014, broad-leaved forest, 2 ex., det. S. K. Alekseev; Novye Pichingushi vill., 0406.2014, birch planting, 1 ex., det. S. K. Alek-seev; Malye Mordovskie Poshaty vill., 0506.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 1 ex., det. S. K. Alekseev (OA).
Odacantha melanura (Linnaeus, 1767) Material: NNR: Sarov, 13.03.2016, 1 ex. (PC).
Ophonus azureus (Fabricius, 1775) Material: RM: Temnikov distr., Mikhailov-ka vill., 06-07.2012, coppice, 1 ex., det. S. K. Alekseev (OA). Elniki distr., Novye Pichingushi vill., 04-06.2014, birch planting, 13 ex., det. S. K. Alekseev (OA).
Ophonus rufibarbis (Fabricius, 1792) Material: RM: Elniki distr., Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 3 ex. (ME). NNR: Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow, 1 ex., det. S. K. Alekseev (ME).
Oxypselaphus obscurus (Herbst, 1784) Material: RM: Temnikov distr., Mikhailov-ka vill., 06-07.2012, coppice, 1 ex., det. S. K. Alekseev (OA). Elniki distr., Novye Shaly vill., 05-06.2014, young pine forest, 6 ex., det. S. K. Alekseev (OA); Cherlyai vill., 15.0528.06.2019, deciduous forest, 1 ex. (ME).
NNR: Sarov, 05-08.2013, 6 ex., det. S. K. Alek-seev (OA). Voznesenskoe distr., Shaprikha vill., 16.05-18.06.2019, mixed forest, 1 ex., det. S. K. Alekseev (ME).
Panagaeus bipustulatus (Fabricius, 1775) Material: RM: Elniki distr., Starye Shaly vill., 05-06.2014, pine forest, 1 ex., det. S. K. Alek-seev (OA); Starye Russkie Poshaty vill., 15.0528.06.2019, meadow, 1 ex. (ME). NNR: Sarov, 18.05.2017, 1 ex. (PC). Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow, 1 ex., det. S. K. Alekseev (ME).
Panagaeus cruxmajor (Linnaeus, 1758) Material: RM: Elniki distr., Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 2 ex. (ME).
Philorhizus sigma (P. Rossi, 1790) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 2 ex., det. S. K. Alekseev (OA); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex. (ME).
Poecilus cupreus (Linnaeus, 1758) Material: RM: Temnikov distr., Mikhailov-ka vill., 06-07.2012, coppice, 2 ex., det. S. K. Alekseev (OA). Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 3 ex., det. S. K. Alekseev (ME). Elniki distr., Peredovoi sett., 05-09.2014, broad-leaved forest, 9 ex., det. S.K. Alekseev; Novye Pich-ingushi vill., 04-06.2014, birch planting, 296 ex., det. S.K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 7 ex., det. S.K. Alekseev; Novye Shaly vill., 05-06.2014, young pine forest, 1 ex., det. S.K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 4 ex., det. S. K. Alekseev (OA); Cherlyai vill., 15.05-28.06.2019, deciduous forest, 3 ex. (ME); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 4 ex. (ME). NNR: Pervomaisk distr., 4 km S Nikolaevka vill.,
30.04-14.07.2014, 2 ex., det. S. K. Alekseev (OA). Voznesenskoe distr., Baranovka vill.,
16.05-18.06.2019, meadow, 2 ex., det. S. K. Alekseev (ME).
Poecilus lepidus (Leske, 1785) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 1 ex., det. S.
K. Alekseev (OA). NNR: Sarov, 05-08.2013, 2 ex., det. S. K. Alekseev (OA). Voznesenskoe distr., Shaprikha vill., 16.05-18.06.2019, mixed forest, 3 ex., det. S. K. Alekseev (ME). Poecilus punctulatus (Schaller, 1783) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 2 ex., det. S. K. Alekseev (OA).
Poecilus versicolor (Sturm, 1824) Material: RM: Temnikov distr., Veselyi sett., 25.05.2013, mixed forest, 1 ex.; Kitsaev-ka vill., 12.05.2013, mixed forest, 1 ex. (AR); Mikhailovka vill., 06-07.2012, coppice, 97 ex., det. S. K. Alekseev (OA). Tengushevo distr., Klemeschshei vill., 14.05-21.06.2019, mixed forest, 99 ex., det. S. K. Alekseev (ME). Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 4 ex., det. S.K. Alekseev; Peredovoi sett., 05-09.2014, broad-leaved forest, 23 ex., det. S.K. Alekseev; Novye Pichin-gushi vill.,
04-06.2014, birch planting, 144 ex., det. S.K. Alekseev; Malye Mordovskie Poshaty vill.,
05-06.2014, broad-leaved forest, 5 ex., det. S. K. Alekseev (OA); Cherlyai vill., 15.0528.06.2019, 31.07-16.09.2019, deciduous forest, 10 ex. (ME); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 45 ex. (ME). NNR: Sarov, 05-08.2013, 5 ex., det. S. K. Alekseev (OA). Pervomaisk distr., 4 km
5 Nikolaevka vill., 30.04-14.07.2014, 36 ex., det. S. K. Alekseev (OA). Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow,
6 ex., det. S. K. Alekseev (ME).
Pterostichus anthracinus (Illiger, 1798) Material: Elniki distr., Novye Pichingushi vill., 04-06.2014, birch planting, 2 ex., det. S. K. Alekseev; Novye Shaly vill., 05-06.2014, young pine forest, 5 ex., det. S. K. Alekseev (OA).
Pterostichus diligens (Sturm, 1824) Material: NNR: Sarov, 05-08.2013, 8 ex., det. S. K. Alekseev (OA).
Pterostichus gracilis (Dejean, 1828) Material: RM: Elniki distr., Starye Shaly vill., 05-06.2014, pine forest, 1 ex., det. S. K. Alekseev (OA); Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex. (ME). NNR: Sarov, 05-08.2013, 1 ex., det. S. K. Alekseev
(OA). Voznesenskoe distr., Baranovka vill., 16.05-18.06.2019, meadow, 1 ex., det. S. K. Alekseev (ME).
Pterostichus minor (Gyllenhal, 1827) Material: RM: Elniki distr., Novye Shaly vill., 05-06.2014, young pine forest, 6 ex., det. S.K. Alekseev; Peredovoi sett., 05-09.2014, broad-leaved forest, 1 ex., det. S.K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 1 ex.; Starye Shaly vill., 05-06.2014, pine forest, 1 ex., det. S. K. Alekseev (OA). Tengushevo distr., Khlebino vill., 14.0520.06.2019, meadow, 1 ex., det. S. K. Alekseev (ME). NNR: Sarov, 05-08.2013, 11 ex., det. S. K. Alekseev (OA).
Pterostichus melanarius (Illiger, 1798) Material: RM: Temnikov distr., Mikhailovka vill., 06-07.2012, coppice, 17 ex., det. S. K. Alek-seev (OA). Elniki distr., Peredovoi sett., 0509.2014, broad-leaved forest, 36 ex., det. S.K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 13 ex., det. S.K. Alekseev; Ma-lye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 30 ex., det. S. K. Alekseev (OA); Cherlyai vill., 15.05-28.06.2019, deciduous forest, 1 ex. (ME). NNR: Voznesenskoe distr., Alamasovo vill., on a dacha plot, in the ground, 23.08.2015, 12.09.2015, 2 ex. (PC). Sarov, 05-08.2013, 22 ex., det. S. K. Alekseev (OA). Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 46 ex., det. S. K. Alek-seev (OA).
Pterostichus niger (Schaller, 1783) Material: RM: Temnikov distr., Andreevka vill., 31.07-16.09.2019, mixed forest, 1 ex. (ME). Elniki distr., Starye Shaly vill., 05-06.2014, pine forest, 1 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 22 ex., det. S. K. Alekseev; Svo-bodnyi sett., 04-06.2014, disturbed meadow, 2 ex., det. S. K. Alekseev; Peredovoi sett., 0509.2014, broad-leaved forest, 23 ex., det. S. K. Alekseev; Novye Pichingushi vill., 04-06.2014, birch planting, 1 ex., det. S. K. Alekseev (OA); Cherlyai vill., 15.05-28.06.2019, deciduous forest, 1 ex. (ME). NNR: Sarov, 05-08.2013, 66 ex., det. S. K. Alekseev (OA). Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 61 ex., det. S. K. Alekseev (OA).
Pterostichus nigrita (Paykull, 1790) Material: RM: Elniki distr., Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 6 ex. (ME). NNR: Sarov, 05-08.2013, 9 ex., det. S. K. Alekseev (OA).
Pterostichus oblongopunctatus (Fabricius, 1787) Material: RM: Temnikov distr., Andreev-ka vill., 31.07-16.09.2019, mixed forest, 2 ex. (ME). Elniki distr., Starye Shaly vill., 0506.2014, pine forest, 85 ex., det. S. K. Alek-seev; Novye Shaly vill., 05-06.2014, young pine forest, 3 ex., det. S. K. Alekseev; Pere-dovoi sett., 05-09.2014, broad-leaved forest, 44 ex., det. S. K. Alekseev; Novye Pich-ingushi vill., 04-06.2014, birch planting, 19 ex., det. S. K. Alekseev; Malye Mordovskie Poshaty vill., 05-06.2014, broad-leaved forest, 86 ex., det. S. K. Alekseev (OA); Cherlyai vill., 15.05-28.06.2019, deciduous forest, 1 ex. (ME). Tengushevo distr., Klemeschshei vill., 14.05-21.06.2019, mixed forest, 6 ex., det. S. K. Alekseev (ME). Temnikov distr., Andreev-ka vill., 14.05-21.06.2019, mixed forest, 3 ex. (ME). NNR: Sarov, 05-08.2013, 105 ex., det. S. K. Alekseev (OA). Pervomaisk distr., 4 km S Nikolaevka vill., 30.04-14.07.2014, 131 ex., det. S. K. Alekseev (OA). Voznesenskoe distr., Shaprikha vill., 16.05-18.06.2019, mixed forest, 31 ex., det. S. K. Alekseev (ME).
Pterostichus quadrifoveolatus Letzner, 1852 Material: RM: Elniki distr., Novye Shaly vill., 05-06.2014, young pine forest, 5 ex., det. S.K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 6 ex., det. S. K. Alekseev (OA).
Pterostichus rhaeticus Heer, 1837 Material: NNR: Sarov, 05-08.2013, 19 ex., det. S. K. Alekseev (OA).
Pterostichus strenuus (Panzer, 1796) Material: Elniki distr., Peredovoi sett., 0509.2014, broad-leaved forest, 1 ex., det. S. K. Alekseev; Starye Shaly vill., 05-06.2014, pine forest, 5 ex., det. S. K. Alekseev (OA); Sta-rye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 7 ex. (ME). NNR: Pervomaisk distr.,
4 km S Nikolaevka vill., 30.04-14.07.2014,
5 ex., det. S. K. Alekseev (OA). Voznesen-skoe distr., Shaprikha vill., 16.05-18.06.2019, mixed forest, 3 ex., det. S. K. Alekseev (ME).
Pterostichus vernalis (Panzer, 1796) Material: RM: Elniki distr., Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 1 ex., det. S. K. Alekseev (ME). NNR: Sarov, 05-08.2013, 6 ex., det. S. K. Alekseev (OA). Sericoda quadripunctata (De Geer, 1774) Material: RM: Temnikov distr., Mikhailov-ka vill., 06-07.2012, coppice, 1 ex., det. S. K. Alekseev (OA).
Stenolophus teutonus (Schrank, 1781) Material: NNR: Sarov, northeast, on wet sand in a ditch on the side of a highway running through the woods, 3.06.2018, 1 ex. (PC).
Stomis pumicatus (Panzer, 1796) Material: RM: Elniki distr., Starye Russkie Poshaty vill., 15.05-28.06.2019, meadow, 4 ex. (ME). NNR: Sarov, 14.04.2016, 1 ex. (PC).
Synuchus vivalis (Illiger, 1798) Material: RM: Temnikov distr., Mikhailov-ka vill., 06-07.2012, coppice, 1 ex., det. S. K. Alekseev (OA). Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 1 ex., det. S. K. Alekseev (OA). NNR: Sarov, 05-08.2013, 16 ex., det. S. K. Alekseev (OA).
Trechus quadristriatus (Schrank, 1781) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 1 ex., det. S. K. Alekseev (OA).
Trechus secalis (Paykull, 1790) Material: NNR: Sarov, 05-08.2013, 24 ex., det. S. K. Alekseev (OA).
Polyphaga Hydrophilidae
Hydrochara caraboides (Linnaeus, 1758) Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 1 ex. (ME).
Hydrophilus aterrimus Eschscholtz, 1822 Material: NNR: Sarov, 5.05.2017, 1 ex. (PC).
Histeridae
Gnathoncus buyssoni Auzat, 1917 Material: NNR: Voznesenskoe distr., Shaprikha vill., 19-24.06.2019, 1 ex. (AR).
Silphidae
Dendroxena quadrimaculata (Scopoli, 1771) Material: NNR: Sarov, 26.05.2017, 1 ex. (PC).
Nicrophorus humator (Gleditsch, 1767) Material: NNR: Sarov, 05-08.2013, 1 ex. (OA).
Nicrophorus vespillo (Linnaeus, 1758) Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 1 ex. (ME).
Nicrophorus vespilloides Herbst, 1783 Material: NNR: Alamasovo vill., 17.06.2017, mixed forest, 1 ex. (PC). Oiceoptoma thoracicum (Linnaeus, 1758) Material: NNR: Alamasovo vill., 20.05.2017, 1 ex. (PC).
Silpha carinata Herbst, 1783 Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 5 ex. (ME).
Silpha obscura Linnaeus, 1758 Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 1 ex. (ME).
Staphylinidae
Lordithon pulchellus (Mannerheim, 1830) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 2.10.2016, 1 ex. (PC).
Nudobius lentus (Gravenhorst, 1806) Material: NNR: Sarov, 10.04.2016, 1 ex. (PC). Ontholestes murinus (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 3.05.2015, 1 ex. (PC); Sarov, 25.07.2013, 1 ex. (AR).
Quedius dilatatus (Fabricius, 1787) Material: RM: Temnikov distr., Tarkhany vill., 16-25.07.2019, 25.07-1.08.2019, 3 ex. (AR). NNR: Pervomaisk distr., 9 km E Be-reschshino vill., 10-22.07.2019, 2 ex. (AR); Voznesenskoe distr., Shaprikha vill., 25.069.07.2019, 2 ex. (AR).
Scaphidium quadrimaculatum G.-A. Olivier, 1790 Material: NNR: Sarov, 19.05.2017, 1 ex. (PC).
Sepedophilus bipustulatus (Gravenhorst, 1802) Material: NNR: Sarov, dry birch, on tinder, 9.05.2019, 1 ex. (PC).
Lucanidae
Platycerus caraboides (Linnaeus, 1758) Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 3 ex. (ME). Elniki distr., Cherlyai vill., 15.0528.06.2019, deciduous forest, 1 ex. (ME).
Geotrupidae
Anoplotrupes stercorosus (L.G. Scriba, 1791) Material: NNR: Sarov, 05-08.2013, 2 ex. (OA).
Scarabaeidae
Aphodius fimetarius (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 30.04.2017, 1 ex. (PC).
Cetonia aurata (Linnaeus, 1758) Material: RM: Temnikov distr., Tarkhany vill., 16-25.07.2019, 25.07-1.08.2019, 2 ex. (AR). NNR: Voznesenskoe distr., Svobodnyi sett., 2-19.06.2019, 5 ex. (AR), Shaprikha vill., 2-19.06.2019, 25.06-9.07.2019, 7 ex. (AR). Chaetopteroplia segetum (Herbst, 1783) Material: NNR: Sarov, 2.06.2015, 1 ex. (PC). Gnorimus variabilis (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Shaprikha vill., 25.06.-9.07.2019, 1 ex. (AR).
Hoplia parvula Krynicki, 1832 Material: NNR: Voznesenskoe distr., Ala-masovo vill., 13.06.2016, 1 ex. (PC). Melolontha hippocastani Fabricius, 1801 Material: NNR: Voznesenskoe distr., Ala-masovo vill., on a dacha plot, in the ground, 30.04.2015, 29.04.2016, 2 ex. (PC).
Oryctes nasicornis (Linnaeus, 1758) Material: NNR: Sarov, 25.06.2016, 9.07.2016, 2 ex. (PC).
Oxythyrea funesta (Poda von Neuhaus, 1761) Material: NNR: Sarov, 5.07.2014, 1 ex.; Voznesenskoe distr., Alamasovo vill., 2.07.2016, 24.06.2017, 2 ex. (PC).
Phyllopertha horticola (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 24.06.2017, 1 ex. (PC).
Protaetia fieberi (Kraatz, 1880) Material: RM: Temnikov distr., Tarkhany vill., 16-25.07.2019, 3 ex. (AR). NNR: Voznesenskoe distr., Svobodnyi sett., 2-19.06.2019, 3 ex. (AR), Shaprikha vill., 2-19.06.2019, 25.069.07.2019, 19-24.06.2019, 24.06.-2.07.2019, 12 ex. (AR).
Protaetia marmorata (Fabricus, 1792) Material: RM: Temnikov distr., Tarkhany vill., 16-25.07.2019, 9 ex. (AR). NNR: Voznesen-skoe distr., Svobodnyi sett., 2-19.06.2019, 11 ex. (AR), Shaprikha vill., 2-19.06.2019, 25.06-9.07.2019, 19-24.06.2019, 33 ex. (AR); Pervomaisk distr., 9 km E Bereschshino vill., 10-22.07.2019, 1 ex. (AR), Kavkazskoe les-nichestvo sett., 10-22.07.2019, 3 ex. (AR); Sa-rov, 24.05.2017, 1 ex. (PC).
Protaetia metallica (Herbst, 1782) Material: RM: Temnikov distr., Tarkhany vill., 16-25.07.2019, 1 ex. (AR). NNR: Voznesen-skoe distr., Svobodnyi sett., 2-19.06.2019, 1 ex. (AR), Shaprikha vill., 19-24.06.2019, 1 ex. (AR); Pervomaisk distr., 9 km E Bereschshino vill., 10-22.07.2019, 1 ex. (AR).
Note. In Catalogue of Palaearctic Coleoptera (Vol. 3) (Bezdèk 2016), the species is listed as Protaetia cuprea volhyniensis (Gory & Percheron, 1833).
Protaetia speciosissima (Scopoli, 1786) Material: NNR: Voznesenskoe distr., Shaprikha vill., 2-19.06.2019, 2 ex. (AR).
Trichius fasciatus (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 16.07.2017, 1 ex. (PC).
Valgus hemipterus (Linnaeus, 1758) Material: NNR: Sarov, 22.05.2015, 2.06.2016, 2.05.2017, 3 ex. (PC).
Byrrhidae
Byrrhus fasciatus (Forster, 1771) Material: RM: Elniki distr., Novye Shaly vill., 26.05.2013, pine forest, 1 ex. (AR).
Byrrhus pilula (Linnaeus, 1758) Material: NNR: Sarov, 25.07.2013, 1 ex. (AR).
Heteroceridae
Heterocerus marginatus (Fabricius, 1787) Material: NNR: Sarov, 3.06.2018, 1 ex. (PC).
Elateridae
Agriotes lineatus (Linnaeus, 1767) Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 1 ex. (ME).
Agriotes obscurus (Linnaeus, 1758) Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 4 ex. (ME).
Agrypnus murinus (Linnaeus, 1758) Material: RM: Temnikov distr., Russkoe Karaevo vill., 8-16.07.2019, 1 ex. (AR). NNR: Voznesenskoe distr., Shaprikha vill., 1924.06.2019, 1 ex. (AR); Sarov, 05-08.2013, 6 ex. (OA).
Ampedus cinnabarinus (Eschscholtz, 1829)
Material: NNR: Sarov, mixed forest, under the bark of a lying tree, 4.09.2015, 1 ex. (PC). Athous subfuscus (O. F. Müller, 1764) Material: NNR: Sarov, 05-08.2013, 1 ex. (OA); Sarov, 25.07.2013, 1 ex. (AR).
Cardiophorus ruficollis (Linnaeus, 1758) Material: NNR: Sarov, 05-08.2013, 1 ex. (OA).
Dalopius marginatus (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Ala-masovo vill., under the bark of a birch stump in a clearing, 24.04.2016, 1 ex. (PC).
Danosoma conspersum (Gyllenhal, 1808) Material: NNR: Sarov, 14.05.2017, 1 ex. (PC).
Drapetes mordelloides (Host, 1789) Material: NNR: Vyksa distr., 21.05.2019, 16.08.2017, 2 ex. (PC).
Melanotus castanipes (Paykull, 1800) Material: NNR: Voznesenskoe distr., Shaprikha vill., 2-19.06.2019, 1 ex. (AR). Prosternon tesselatum (Linnaeus, 1758) Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 3 ex. (ME). NNR: Sarov, 05-08.2013, 2 ex. (OA); Sarov, 25.07.2013, 1 ex. (AR).
Selatosomus aeneus (Linnaeus, 1758) Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 2 ex. (ME). NNR: Sarov, 05-08.2013, 1 ex. (OA).
Cantharidae
Cantharis livida Linnaeus, 1758 Material: NNR: Voznesenskoe distr., Svo-bodnyi sett., 2-19.06.2019, 2 ex. (AR).
Cantharis pellucida Fabricius, 1792 Material: NNR: Voznesenskoe distr., Shaprikha vill., 2-19.06.2019, 2 ex. (AR).
Ptinidae
Ptinus rufipes G.-A. Olivier, 1790 Material: RM: Elniki distr., Malye Mordo-vskie Poshaty vill., 05-06.2014, broad-leaved forest, 1 ex. (AR).
Cleridae
Thanasimus formicarius (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 10.05.2015, 1 ex. (PC).
Trichodes apiarius (Linnaeus, 1758) Material: NNR: Sarov, 12.07.2016, 1 ex.; Voznesenskoe distr., Alamasovo vill., 10.06.2018, 1 ex. (PC).
Melyridae
Apalochrusfemoralis Erichson, 1840 Material: RM: Elniki distr., Novye Shaly vill., 19.07.2015, 2 ex. (AR).
Cordylepherus viridis (Fabricius, 1787) Material: RM: Elniki distr., Malye Mordov-skie Poshaty vill., 19.07.2015, 1 ex. (AR).
Dolichosoma lineare (P. Rossi, 1794) Material: RM: Elniki distr., Novye Shaly vill., 19.07.2015, 2 ex. (AR).
Nitidulidae
Cryptarcha strigata (Fabricius, 1787) Material: RM: Temnikov distr., Tar-khany vill., 16-25.07.2019, 1 ex. (AR). NNR: Voznesenskoe distr., Svobodnyi sett., 2-19.06.2019, 3 ex. (AR).
Cychramus luteus (Fabricius, 1787) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 24.07.2016, 1 ex. (PC).
Epuraea biguttata (Thunberg, 1784) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 1.05.2015, 1 ex. (PC); Sarov, 14.04.2016, 1 ex. (PC). (det. A. Kirejtshuk).
Glischrochilus grandis (Tournier, 1872) Material: RM: Temnikov distr., Tarkha-ny vill., 16-25.07.2019, 25.07-1.08.2019, 2 ex. (AR). Elniki distr., Starye Shaly vill., 0506.2014, pine forest, 1 ex.; Svobodnyi sett., 04-06.2014, disturbed meadow, 1 ex. (AR). NNR: Voznesenskoe distr., Shaprikha vill., 2-19.06.2019, 1 ex. (AR), Alamasovo vill., 10.05.2015, 1 ex. (PC).
Glischrochilus hortensis (Geoffroy, 1785) Material: RM: Elniki distr., Starye Shaly vill., 05-06.2014, pine forest, 1 ex. (AR). Glischrochilus quadripunctatus (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Alamasovo vill., 12.04.2015, 10.05.2015, 2 ex. (PC). Glischrochilus quadrisignatus (Say, 1835) Material: RM: Temnikov distr., Tarkhany vill., 16-25.07.2019, 25.07-1.08.2019, 3 ex. (AR).
Ipidia binotata Reitter, 1875 Material: NNR: Vyksa distr., Vyksa, 13.08.2019, 12.08.2017, 2 ex. (PC). Ipidia sexguttata (R.F. Sahlberg, 1834) Material: NNR: Vyksa distr., Vyksa, 21.05.2019, 22.05.2019, 2 ex. (PC).
Soronia grisea (Linnaeus, 1758) Material: RM: Temnikov distr., Tarkhany vill., 16-25.07.2019, 25.07-1.08.2019, 3 ex. (AR).
Bothrideridae
Bothrideres bipunctatus (Gmelin, 1790) Material: NNR: Sarov, 29.05.2015, 9.04.2015, 29.04.2016, 3 ex. (PC).
Endomychidae
Dapsa horvathi (Csiki, 1901) Material: RM: Elniki distr., Novye Shaly vill., 19.07.2015, 1 ex. (AR).
Mycetina cruciata (Schaller, 1783) Material: RM: Elniki distr., Novye Shaly vill., 05-06.2014, young pine forest, 1 ex. (AR).
Coccinellidae
Coccinella quinquepunctata Linnaeus, 1758 Material: NNR: Sarov, 25.07.2013, 1 ex. (AR).
Oenopia conglobata (Linnaeus, 1758) Material: RM: Temnikov distr., Tarkhany vill., 16-25.07.2019, 1 ex. (AR).
Melandryidae
Dircaea quadriguttata (Paykull, 1798) Material: NNR: Sarov, 2.06.2015, 19.06.2015, 2 ex. (PC); Voznesenskoe distr., Alamasovo vill., 16.07.2017, 2 ex. (PC).
Melandrya dubia (Schaller, 1783) Material: NNR: Sarov, 26.05.2017, 1 ex. (PC).
Meloidae
Cerocoma schaefferi (Linnaeus, 1758) Material: RM: Temnikov distr., Tarkhany vill., 7.07.2013, 1 ex. Elniki distr., Malye Mor-dovskie Poshaty vill., 31.05.2014, 1 ex.
Lytta vesicatoria (Linnaeus, 1758) Material: RM: Elniki distr., Starye Pichin-gushi vill., 20.06.2012, 1 ex. (GS).
Meloe proscarabaeus Linnaeus, 1758 Material: NNR: Sarov, 6.05.2017, 22.05.2017, 2 ex. (PC).
Meloe violaceus Marsham, 1802 Material: NNR: Sarov, 10.05.2016, 7.06.2017, 2 ex. (PC).
Oedemeridae
Chrysanthia viridissima (Linnaeus, 1758) Material: RM: Elniki distr., Novye Shaly vill., 19.07.2015, 2 ex.; Malye Mordovskie Poshaty vill., 22.07.2017, 1 ex. (AR). NNR: Sarov, 25.07.2013, 1 ex. (AR).
Oedemera femorata (Scopoli, 1763) Material: RM: Elniki distr., Novye Shaly vill., 19.07.2015, 1 ex.; Malye Mordovskie Poshaty vill., 23.07.2016, 22.07.2017, 2 ex.; Novoyam-skaya Sloboda vill., 22.07.2017, 3 ex. (AR).
Oedemera virescens (Linnaeus, 1767) Material: RM: Elniki distr., Novye Shaly vill., 26.05.2013, pine forest, 1 ex. (AR).
Tenebrionidae
Crypticus quisquilius (Linnaeus, 1760) Material: RM: Elniki distr., Svobodnyi sett., 04-06.2014, disturbed meadow, 14 ex. (AR). Tengushevo distr., Klemeschshei vill., 14.0521.06.2019, mixed forest, 3 ex. (ME); Khlebino vill., 14.05-20.06.2019, meadow, 2 ex. (ME).
Lagria hirta (Linnaeus, 1758) Material: RM: Elniki distr., Malye Mordovs-kie Poshaty vill., 19.07.2015, 23.07.2016, 2 ex.; Novye Shaly vill., 22.07.2017, 1 ex. (AR). Lagria laticollis Motschulsky, 1860 Material: Elniki distr., Malye Mordovskie Poshaty vill., 23.07.2016, 1 ex.
Opatrum sabulosum (Linnaeus, 1760) Material: RM: Elniki distr., Svobodnyi sett.,
04-06.2014, disturbed meadow, 212 ex. (AR). Upis ceramboides (Linnaeus, 1758)
Material: NNR: Voznesenskoe distr., Shaprikha vill., 16.05-18.06.2019, mixed forest, 2 ex. (ME).
Cerambycidae
Arhopalus rusticus (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 10.07.2016, 1 ex. (PC).
Aromia moschata (Linnaeus, 1758) Material: RM: Temnikov distr., Tarkhany vill., 16-25.07.2019, 1 ex. (AR). NNR: Voznesen-skoe distr., Shaprikha vill., 19-24.06.2019, 1 ex. (AR); Pervomaisk distr., 9 km E Bere-schshino vill., 10-22.07.2019, 2 ex. (AR); Per-vomaisk distr., Kavkazskoe lesnichestvo sett., 10-22.07.2019, 1 ex. (AR).
Chlorophorus herbstii (Brahm, 1790) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 12.06.2019, 1 ex. (PC).
Cortodera femorata (Fabricius, 1787) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 21.05.2016, 1 ex. (PC); Sarov,
05-08.2013, 2 ex. (OA).
Dinoptera collaris (Linnaeus, 1758) Material: NNR: Sarov, 23.06.2017, 1 ex. (PC).
Lamia textor (Linnaeus, 1758) Material: NNR: Sarov, mixed forest, 11.06.2016, 1 ex. (PC); Sarov, 25.07.2013, 1 ex. (AR).
Leptura quadrifasciata Linnaeus, 1758 Material: NNR: Voznesenskoe distr., Shaprikha vill., 2-19.06.2019, 3 ex. (AR); Per-vomaisk distr., 9 km E Bereschshino vill., 1022.07.2019, 2 ex. (AR); Voznesenskoe distr., Alamasovo vill., 27.06.2015, 4.07.2015, 3 ex. (PC).
Leptura thoracica Creutzer, 1799 Material: NNR: Voznesenskoe distr., Svobodnyi sett., 2-19.06.2019, 4 ex. (AR). NNR: Voznesenskoe distr., Shaprikha vill., 2-19.06.2019, 3 ex. (AR); Sarov, 20.06.2016, 1 ex. (PC).
Lepturobosca virens (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Alamasovo vill., 27.07.2017, 30.07.2017, 30.06.2018, 3 ex. (PC).
Mesosa myops (Dalman, 1817) Material: NNR: Sarov, 22.05.2015, 17.06.2016, 2 ex. (PC).
Monochamus sutor (Linnaeus, 1758) Material: RM: Temnikov distr., Russkoe Karaevo vill., 8-16.07.2019, 1 ex. (AR). NNR: Sarov, 29.06.2017, 1 ex. (PC).
Necydalis major Linnaeus, 1758 Material: NNR: Voznesenskoe distr., Shaprikha vill., 2-19.06.2019, 1 ex. (AR).
Oedecnema gebleri (Ganglbauer, 1889) Material: NNR: Sarov, 28.05.2019, 1 ex. (PC).
Oxymirus cursor (Linnaeus, 1758) Material: NNR: Sarov, 05-08.2013, 1 ex. (OA).
Pachyta quadrimaculata (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 20.06.2015, 1 ex. (PC).
Prionus coriarius (Linnaeus, 1758) Material: NNR: Sarov, 17.07.2016, 1 ex. (PC). Purpuricenus kaehleri (Linnaeus, 1758) RM: Temnikov distr., Russkoe Karaevo vill., 8-16.07.2019, 1 ex. (AR), Tarkhany vill., 16-25.07.2019, 25.07.-1.08.2019, 2 ex. (AR). NNR: Voznesenskoe distr., Shaprikha vill., 2-19.06.2019, 25.06.-9.07.2019, 4 ex. (AR).
Rhagium inquisitor (Linnaeus, 1758) Material: NNR: Sarov, 28.06.2015, 1 ex. (PC).
Rhagium mordax (De Geer, 1775) Material: RM: Tengushevo distr., Kleme-schshei vill., 14.05-21.06.2019, mixed forest, 1 ex. (ME). NNR: Voznesenskoe distr., Shaprikha vill., 2-19.06.2019, 19-24.06.2019, 16 ex. (AR), Alamasovo vill., 7.05.2016, 1 ex. (PC); Pervomaisk distr., 9 km E Bereschshino vill., 10-22.07.2019, 3 ex. (AR); Sarov, 3.05.2015, 1 ex. (PC).
Rusticoclytus rusticus (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 14.06.2015, 1 ex. (PC); Sarov, 25.07.2013, 1 ex. (AR).
Rutpela maculata (Poda, 1761) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 19.07.2015, 2.07.2016, 2 ex. (PC). Spondylis buprestoides (Linnaeus, 1758) Material: NNR: Sarov, 19.06.2015, 19.06.2016, 11.07.2017, 4 ex. (PC).
Stenocorus meridianus (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Shaprikha vill., 2-19.06.2019, 11 ex. (AR).
Stenurella bifasciata (O.F. Müller, 1776) Material: NNR: Voznesenskoe distr., Ala-masovo vill., 5.07.2014, 1 ex., Sarov, 1.07.2015, 1 ex. (PC); Sarov, 25.07.2013, 1 ex. (AR).
Stenurella melanura (Linnaeus, 1758) Material: NNR: Sarov, 05-08.2013, 2 ex. (OA); NNR: Sarov, 25.07.2013, 1 ex. (AR).
Stictoleptura rubra (Linnaeus, 1758) Material: NNR: Sarov, 28.06.2015, 1 ex.; Voznesenskoe distr., Alamasovo vill., 19.07.2015, 1 ex. (PC).
Chrysomelidae
Agelastica alni (Linnaeus, 1758) Material: RM: Elniki distr., Bolshoi Urkat vill., 13.05.2009, 1 ex. Temnikov distr., Kitsaev-ka vill., 12.05.2013, mixed forest, 4 ex. (AR).
Cassida nebulosa Linnaeus, 1758 Material: RM: Elniki distr., Novye Shaly vill., 26.05.2013, pine forest, 1 ex. Temnikov distr., Andreevka vill., 11.05.2013, the edge of the forest, 1 ex. (AR).
Cassida sanguinosa Suffrian, 1844 Material: RM: Elniki distr., Novye Shaly vill., 22.07.2017, 1 ex. (AR).
Cassida stigmatica Suffrian, 1844 Material: RM: Elniki distr., Novye Shaly vill., 19.07.2015, 1 ex. (AR).
Chrysolina hyperici (Forster, 1771) Material: RM: Elniki distr., Cherlyai vill., 23.07.2016, 1 ex. (AR).
Chrysolina varians (Schaller, 1783) Material: RM: Elniki distr., Svobodnyi sett., 31.07.2008, meadow, 1 ex.; Novye Shaly vill., 19.07.2015, 1 ex. (AR).
Chrysomela collaris Linnaeus, 1758 Material: RM: Temnikov distr., Kitsaevka vill., 12.05.2013, mixed forest, 1 ex. (AR).
Chrysomela populi Linnaeus, 1758 Material: RM: Elniki distr., Novye Shaly vill., 26.05.2013, pine forest, 1 ex. (AR). Cryptocephalus laetus Fabricius, 1792 Material: RM: Elniki distr., Malye Mordovs-kie Poshaty vill., 19.07.2015, 1 ex. (AR). Coptocephala unifasciata unifasciata (Scopoli, 1763) Material: RM: Elniki distr., Malye Mordovs-kie Poshaty vill., 22.07.2017, 2 ex.; Novye Shaly vill., 22.07.2017, 1 ex. (AR).
Cryptocephalus moraei (Linnaeus, 1758) Material: NNR: Sarov, 25.07.2013, 1 ex. (AR). Cryptocephalus solivagus Leonardi et Sassi, 2001 Material: RM: Elniki distr., Novye Shaly vill., 19.07.2015, 1 ex.; Malye Mordovskie Poshaty vill., 19.07.2015, 22.07.2017, 3 ex. (AR). NNR: Sarov, 25.07.2013, 1 ex. (AR). Gonioctena quinquepunctata (Fabricius, 1787) Material: RM: Temnikov distr., Kitsaevka vill., 12.05.2013, mixed forest, 2 ex. (AR). Gonioctena viminalis (Linnaeus, 1758) Material: RM: Elniki distr., Novye Shaly vill., 26.05.2013, pine forest, 2 ex. Temnikov distr., Veselyi sett., 25.05.2013, mixed forest, 2 ex. (AR).
Labidostomis longimana (Linnaeus, 1760) Material: RM: Elniki distr., Novoyamskaya Sloboda vill., 22.07.2017, 1 ex. (AR). Pachybrachis hieroglyphicus (Laicharting, 1781) Material: RM: Elniki distr., Svobodnyi sett., 31.07.2008, meadow, 1 ex. (AR); Novye Shaly vill., 22.07.2017, 1 ex. (AR). Phyllobrotica quadrimaculata (Linnaeus, 1758) Material: RM: Elniki distr., Novye Shaly vill., 19.07.2015, 1 ex. (AR).
Plagiodera versicolora (Laicharting, 1781) Material: RM: Temnikov distr., Kitsaevka vill., 12.05.2013, mixed forest, 1 ex. (AR).
Nemonychidae
Cimberis attelaboides (Fabricius, 1787) Material: RM: Elniki distr., Starye Shaly vill., 05-06.2014, pine forest, 1 ex.
Anthribidae
Dissoleucas niveirostris (Fabricius, 1798) Material: NNR: Voznesenskoe distr., Alamasovo vill., 3.05.2015, 1 ex. (PC).
Platyrhinus resinosus (Scopoli, 1763) Material: NNR: Voznesenskoe distr., Alamasovo vill., 3.05.2015, 1 ex. (PC); Sarov, 26.05.2016, 1 ex. (PC).
Platystomos albinus (Linnaeus, 1758) Material: NNR: Voznesenskoe distr., Alamasovo vill., 1.05.2014, 1 ex. (PC); 20.05.2017, 1 ex. (PC). RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 1 ex.
Attelabidae
Byctiscus betulae (Linnaeus, 1758) Material: RM: Elniki distr., Novye Shaly vill., 26.05.2013, pine forest, 1 ex.
Curculionidae
Chlorophanus viridis (Linnaeus, 1785) Material: RM: Elniki distr., Starye Pichin-gushi vill., 25.06.2012, 2 ex. (GB). Cryptorhynchus lapathi (Linnaeus, 1758) Material: RM: Elniki distr., Starye Pichin-gushi vill., 16.08.2008, 1 ex. (DK).
Curculio venosus (Gravenhorst, 1807) Material: RM: Temnikov distr., Tarkhany vill., 16-25.07.2019, 1 ex. (AR).
Eusomus ovulum Germar, 1823 Material: RM: Elniki distr., Malye Mordovs-kie Poshaty vill., 22.07.2017, 2 ex. (AR).
Hylastes cunicularius Erichson, 1836 Material: RM: Elniki distr., Novye Shaly vill., 05-06.2014, young pine forest, 1 ex. (AR).
Hylobius abietis (Linnaeus, 1758) Material: RM: Elniki distr., Novye Pichin-gushi vill., 04-06.2014, birch planting, 1 ex. (AR); Novye Shaly vill., 05-06.2014, 27 ex. (AR); Starye Shaly vill., 05-06.2014, 4 ex. (AR). NNR: Sarov, 05-08.2013, 15 ex. (OA).
Hylobius pinastri (Gyllenhal, 1813) Material: NNR: Sarov, 25.07.2013, 1 ex. (AR).
Larinus obtusus Gyllenhal, 1835 Material: RM: Elniki distr., Malye Mordovs-
Table 1
Composition of Coleopterous insect fauna in the vicinity of the Mordovia Nature Reserve
Таблица 1
Состав фауны жесткокрылых насекомых в окрестностях Мордовского заповедника
Family Number of species
Carabidae 115
Hydrophilidae 2
Histeridae 1
Silphidae 7
Staphylinidae 6
Lucanidae 1
Geotrupidae 1
Scarabaeidae 15
Byrrhidae 2
Heteroceridae 1
Elateridae 12
Cantharidae 2
Ptinidae 1
Cleridae 2
Melyridae 3
Nitidulidae 10
Bothrideridae 1
Endomychidae 2
Coccinellidae 2
Melandryidae 2
Meloidae 4
Oedemeridae 3
Tenebrionidae 5
Cerambycidae 26
Chrysomelidae 18
Nemonychidae 1
Anthribidae 3
Attelabidae 1
Curculionidae 11
Total 260
kie Poshaty vill., 19.07.2015, 23.07.2016, 3 ex.; Cherlyai vill., 23.07.2016, 1 ex. (AR).
Phyllobius pyri (Linnaeus, 1758) Material: RM: Elniki distr., Novye Shaly vill., 26.05.2013, pine forest, 1 ex. (AR).
Sitona inops Schoenherr, 1832 Material: RM: Elniki distr., Novye Shaly vill., 19.07.2015, 1 ex. (AR).
Tanymecus palliatus (Fabricius, 1887) Material: RM: Elniki distr., Novye Shaly vill., 26.05.2013, pine forest, 1 ex. (AR).
Thus, as of today, 260 species from 29 fami-lies of Coleoptera are found in the vicinity of the Mordovia Nature Reserve (Table 1). The Carabidae
have been studied to the greatest extent, while the other families have not been sufficiently studied yet. Mostly common species have been recorded, but some rare taxa have also been found.
Acknowledgements
The authors express their gratitude to P. V. Cherenkov (Sarov, Russia) for
the photographic images, to K. V. Ma-karov (Moscow, Moscow Pedagogical State University) for the definition of Cylindera arenaria viennensis, and to A. G. Kirejtshuk (Saint Petersburg, Zoological Institute of the Russian Academy of Sciences) for the definition of Epu-raea biguttata.
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interactions. Climatic Change, vol. 15, no. 1-2, pp. 255-270. (In English) Redikortsev, V. V. (1938) Materialy k entomofaune Mordovskogo gosudarstvennogo zapovednika [Materials to the entomofauna of the Mordovia State Nature Reserve]. In: Fauna Mordovskogo gosudarstvennogo zapovednika im. P. G. Smidovicha: Nauchnye rezul'taty rabot Zoologicheskoj ekspeditsii pod rukovodstvom prof. S. S. Turova v 1936 g. [Fauna of the Mordovia State Nature Reserve: Scientific results of the Zoological expedition under the guidance of prof. S. S. Turov in 1936]. Moscow: Gudok Publ., pp. 137-146. (In Russian) Ruchin, A. B., Alekseev, S. K., Khapugin, A. A. (2019a) Post-fire fauna of carabid beetles (Coleoptera, Carabidae) in forests of the Mordovia State Nature Reserve (Russia). Nature Conservation Research, vol. 4, suppl. 1, pp. 11-20. https://dx.doi.org/10.24189/ncr.2019.009 (In English) Ruchin, A. B., Egorov, L. V. (2017) Obzor vidov nasekomykh Mordovskogo zapovednika, vklyuchaemykh v Krasnuyu knigu Rossijskoj Federatsii [Overview of insect species included in the Red Data Book of Russian Federation in the Mordovia State Nature Reserve]. Nature Conservation Research, vol. 2, suppl. 1, pp. 2-9. https://www.doi.org/10.24189/ncr.2017.016 (In Russian)
Ruchin, A. B., Egorov, L. V. (2018a) Zhestkokrylye (Insecta, Coleoptera), sobrannye fermentnymi kronovymi lovushkami v Mordovii. Soobshchenie 1. Mordovskij zapovednik [Beetles (Insecta, Coleoptera), collected using fermental crown trap in the Republic of Mordovia. Report 1. Mordovia State Nature Reserve]. In: L. V. Egorov (ed.). Nauchnye trudy gosudarstvennogo prirodnogo zapovednika "Prisurskii" [Scientific Proceedings of the State Nature Reserve "Prisursky"]. Vol. 33. Cheboksary: s. n., pp. 209-215. (In Russian) Ruchin, A. B., Egorov, L. V. (2018b) Discovery of Allonyx quadrimaculatus (Schaller, 1783) (Coleoptera Cleridae Clerinae) in Russia. REDIA, vol. 101, pp. 143-146. http://dx.doi.org/10.19263/ REDIA-101.18.19 (In English) Ruchin, A. B., Egorov, L. V. (2018c) Leptura aurulenta (Coleoptera, Cerambycidae), a new record of a very rare species in Russia. Nature Conservation Research, vol. 3, no. 1, pp. 88-91. https://www.doi. org/10.24189/ncr.2018.003 (In English) Ruchin, A. B., Egorov, L. V. (2018d) On distribution of Mimela holosericea (Fabricius, 1787) (Insecta, Scarabaeoidea, Scarabaeidae, Rutelinae) in Russia and adjacent territories. Journal of Entomological and Acarological Research, vol. 50, no. 2, article 7390. https://www.doi.org/10.4081/jear.2018.7390 (In English) Ruchin, A. B., Egorov, L. V. (2018e) Zhestkokrylye (Insecta, Coleoptera), sobrannye fermentnymi kronovymi lovushkami v Mordovii. Soobshchenie 4. Razlichnye lesnye biotsenozy [Beetles (Insecta, Coleoptera), collected using fermental crown trap in the Republic of Mordovia. Report 4. Different forest biocoenoses]. In: L. V. Egorov (ed.). Nauchnye trudy gosudarstvennogo prirodnogo zapovednika "Prisurskii" [Scientific Proceedings of the State Nature Reserve "Prisursky"]. Vol. 33. Cheboksary: s. n., pp. 222-226. (In Russian)
Ruchin, A. B., Egorov, L. V. (2018f) Fauna of longicorn beetles (Coleoptera: Cerambycidae) of Mordovia. Russian Entomological Journal, vol. 27, no. 2, pp. 161-177. https://www.doi.org/10.15298/ rusentj.27.2.07 (In English) Ruchin, A. B., Egorov, L. V. (2019) Contribution to the study of the Cantharoidea (Coleoptera: Drilidae, Lycidae, Lampyridae, Cantharidae) in the Republic of Mordovia (Russia). Entomology and Applied Science Letters, vol. 6, no. 2, pp. 1-12. (In English) Ruchin, A. B., Egorov, L. V., Alekseev, S. K. (2013) Annotirovannyj spisok zhukov-mertvoedov (Coleoptera, Silphidae) Mordovii [The annotated list of carrion beetles (Coleoptera, Silphidae) in Mordovia]. Izvestiya vysshikh uchebnykh zavedenij. Povolzhskij region. Estestvennye nauki — University Proceedings. Volga Region. Natural Sciences, no. 2 (2), pp. 28-41. (In Russian) Ruchin, A. B., Egorov, L. V., Khapugin, A. A. et al. (2020) The use of simple crown traps for the insects collection. Nature Conservation Research, vol. 5, no. 1, pp. 87-108. https://dx.doi.org/10.24189/ ncr.2020.008 (In English)
Ruchin, A. B., Egorov, L. V., Sazhnev, A. S. et al. (2019a) Present distribution of Protaetiafieberi (Kraatz, 1880) (Insecta, Coleoptera, Scarabaeidae) in the European part of Russia. Biharean Biologist, vol. 13, no. 1, pp. 12-16, article e181206. (In English) Ruchin, A. B., Egorov, L. V., Semishin, G. B. (2018) Fauna of click beetles (Coleoptera: Elateridae) in the interfluve of Rivers Moksha and Sura, Republic of Mordovia, Russia. Biodiversitas, vol. 19, no. 4, pp. 1352-1365. https://www.doi.org/10.13057/biodiv/d190423 (In English) Ruchin, A. B., Egorov, L. V., Sazhnev, A. S., Ishin, R. N. (2019b) Metoecus paradoxus (Linnaeus, 1760) (Coleoptera: Ripiphoridae) — novyj vid v faune Respubliki Mordoviya, Nizhegorodskoj, Saratovskoj i Tambovskoj oblastej [Metoecus paradoxus (Linnaeus, 1760) (Coleoptera: Ripiphoridae), a new species for the fauna of Republic of Mordovia, Nizhniy Novgorod, Saratov and Tambov Provinces]. Eversmanniya — Eversmannia, no. 59-60, pp. 59-60. (In Russian) Ruchin, A. B., Egorov, L. V., Semishin, G. B. (2019c) Ladybird beetles fauna (Coleoptera: Coccinellidae) of the Republic of Mordovia, Russia. Biodiversitas, vol. 20, no. 2, pp. 316-327. https://www.doi. org/10.13057/biodiv/d200203 (In English) Ruchin, A. B., Khapugin, A. A. (2019). Red Data Book invertebrates in a protected area of European Russia. Acta Zoologica Academiae Scientiarum Hungaricae, vol. 65, no. 4, pp. 349-370. https:// www.doi.org/10.17109/AZH.65A349.2019 (In English) Ruchin, A. B., Kurmaeva, D. K. (2010) On rare insects of Mordovia included in the Red Book of the Russian Federation. Entomological Review, vol. 90, no. 6, pp. 712-717. https://www.doi.org/10.1134/ S0013873810060060 (In English)
Semenov, V. B. (2015) Dopolnenie k faune zhukov-stafilinid (Coleoptera, Staphylinidae) Mordovskogo gosudarstvennogo prirodnogo zapovednika [Additions to the fauna of staphylinid beetles (Coleoptera, Staphylinidae) of the Mordovia State Nature Reserve]. Trudy Mordovskogo gosudarstvennogo prirodnogo zapovednika imeni P. G. Smidovicha — Proceedings of the Mordovia State Nature Reserve, no. 14, pp. 358-365. (In Russian) Semenov, V. B. (2016) Novye dannye po faune zhukov-stafilinid (Coleoptera, Staphylinidae) Mordovii [New data on the fauna of Staphylinidae (Coleoptera) of Mordovia]. Trudy Mordovskogo gosudarstvennogo prirodnogo zapovednika imeni P. G. Smidovicha — Proceedings of the Mordovia State Nature Reserve, no. 16, pp. 431-434. (In Russian) Semenov, V. B. (2017) Materialy k poznaniyu zhukov-stafilinid (Coleoptera, Staphylinidae) Mordovskogo zapovednika [Materials for the knowledge of staphylinid beetles (Coleoptera, Staphylinidae) of the Mordovia State Nature Reserve]. Trudy Mordovskogo gosudarstvennogo prirodnogo zapovednika imeni P. G. Smidovicha — Proceedings of the Mordovia State Nature Reserve, no. 18, pp. 190-205. (In Russian)
Spitzer, L., Meca, P., Benes, J. (2014) Nova lokalita sviznika piscinneho (Cylindera arenaria viennensis) (Coleoptera: Carabidae) na severni Morave [New locality of the Viennese Tiger Beetle (Cylindera arenaria viennensis) (Coleoptera: Carabidae) in the northern Moravia]. Acta Musei Beskidensis, vol. 6, pp. 131-134. (In Czech) Tomaszewska, W., Egorov, L. V., Ruchin, A. B., Vlasov, D. V. (2018) First record of Clemmus troglodytes (Coleoptera: Coccinelloidea, Anamorphidae) for the fauna of Russia. Nature Conservation Research, vol. 3, no. 3, pp. 103-105. https://www.doi.org/10.24189/ncr.2018.016 (In English) Uligova, T. S., Gedgafova, F. V., Gorobtsova, O. N. et al. (2019) Lugovye biogeotsenozy subal'pijskogo poyasa Kabardino-Balkarskogo gosudarstvennogo vysokogornogo zapovednika (Tsentral'nyj Kavkaz) [Meadow biogeocenoses in the subalpine belt of the Kabardino-Balkaria State High-Mountain Reserve (Central Caucasus)]. Nature Conservation Research, vol. 4, no. 2, pp. 29-47. http://dx.doi. org/10.24189/ncr.2019.012 (In Russian) Xavier da Silva, M., Paviolo, A., Tambosi, L. R., Pardini, R. (2018) Effectiveness of protected areas for biodiversity conservation: Mammal occupancy patterns in the Igua^u National Park, Brazil. Journal Nature Conservation, vol. 41, pp. 51-62. https://doi.org/10.1016/j.jnc.2017.11.001 (In English)
For citation: Ruchin, A. B., Egorov, L. V., Alekseev, S. K., Semishin, G. B., Esin, M. N. (2021) Notes on the fauna
of beetles (Insecta, Coleoptera) adjacent to the territory of the Mordovia State Nature Reserve. Amurian Zoological Journal, vol. XIII, no. 1, pp. 12-35. https://www.doi.org/10.33910/2686-9519-2021-13-1-12-35
Received 22 August 2020; reviewed 20 November 2020; accepted 12 December 2020.
Для цитирования: Ручин, А. Б., Егоров, Л. В., Алексеев, С. К., Семишин, Г. Б., Есин, М. Н. (2021) Материалы
к фауне жесткокрылых (Insecta, Coleoptera) окрестностей Мордовского заповедника. Амурский зоологический журнал, т. XIII, № 1, с. 12-35. https://www.doi.org/10.33910/2686-9519-2021-13-1-12-35 Получена 22 августа 2020; прошла рецензирование 20 ноября 2020; принята 12 декабря 2020.
