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Вестник Пермского университета. Серия Биология. 2023. Вып. 1. С. 19-30. Bulletin of Perm University. Biology. 2023. Iss. 1. P. 19-30.
ЗООЛОГИЯ
Научная статья УДК 595.44(574)
doi: 10.17072/1994-9952-2023-1-19-30.
Новые данные о фауне пауков (Arachnida: Araneae) Западно - Казахстанской области
Сергей Леонидович Есюнин1 Алиби Аскарулы Кабдрахимов2
1 2 Пермский государственный национальный исследовательский университет, Пермь, Россия 1Н esyunin@psu.ru, https://orcid.org/0000-0003-3813-1316
2 kab.ali93@mail.ru, https://orcid.org/0000-0002-3721-1691
Аннотация. Регулярные арахно-фаунистические исследования Западно-Казахстанской обл. начались только в 90-х гг. ХХ в. На сегодняшний день опубликовано более 65 арахнологических работ по этому региону. Однако фауна пауков этого региона остается недостаточно изученной. Приведен список 77 видов пауков, собранных в 2021-2022 гг. в Западно-Казахстанской обл. Три вида - Mecynargus minutipalpis Gnelitsa, 2011, Neoscona spasskyi (Brignoli, 1983) и Rhysodromus hierosolymitanus (Levy, 1977) отмечены в Казахстане впервые. Сообщается о 19 новых для Западно-Казахстанской обл. видах и 86 новых для районов указаний для 55 видов. На основании полученных результатов общее количество видов пауков, зарегистрированных в Западно-Казахстанской обл., возросло до 274 видов из 130 родов и 23 семейств. Для трех малоизвестных видов - Evippa sjostedti Schenkel, 1936, M. minutipalpis и R. hierosolymitanus, а также одного паука-скакунчикаAttulus zimmermanni (Simon, 1877) приведены фотографии эпигин. Ключевые слова: фаунистика, Казахстан, пауки, новые указания, новые находки Для цитирования: Есюнин С. Л., Кабдрахимов А. А. Новые данные о фауне пауков (Arachnida: Araneae) Западно-Казахстанской области // Вестник Пермского университета. Сер. Биология. 2023. Вып. 1. С. 19-30. [На англ.]. http://dx.doi.org/10.17072/1994-9952-2023-1-19-30.
Благодарности: Гюлли Ш. Фарзалиевой (Пермский государственный университет, Пермь) за помощь в изготовлении цифровых фотографий. Особая благодарность Андрею В. Танасевичу (Институт экологии и эволюции им. А.Н. Северцова РАН, Москва, Россия) за определение самки Mecynargus minutipalpis, упомянутой в этой статье, и Дмитрию В. Логунову (Манчестер, Великобритания), который идентифицировал самку Rhysodromus hierosolymitanus и отредактировал английский язык окончательного варианта статьи.
ZOOLOGY
Original article
New data on the spider fauna of West Kazakhstan Region
(Arachnida: Araneae)
Sergei L. Esyunin Alibi A. Kabdrakhimov2
1 2 Perm State University, Perm, Russia
1H esyumn@psu.ru, https ://orcid. org/0000-0003 -3813-1316
2 kab.ali93@mail.ru, https://orcid.org/0000-0002-3721-1691
Abstract. Regular arachno-faunistic studies of West Kazakhstan Region started only in the 90s of the 20th century. To date, over 65 arachnological papers on this region have been published,. Yet, the spider fauna of West Kazakhstan Region remains incompletely studied. A list of 77 spider species collected in 2021-2022 from West Kazakhstan Region is provided. Three species - Mecynargus minutipalpis Gnelitsa, 2011, Neoscona spasskyi (Brignoli, 1983) and Rhysodromus hierosolymitanus (Levy, 1977), are recorded from Kazakhstan for the first time. Nineteen species are reported for West Kazakhstan Region for the first time, and 86 new district records are provided for 55 species. Baseed on the present results, a total number of spider species recorded from West Kazakhstan Region has raised to 274 species in 130 genera and 23 families. Three poorly known species, Evippa sjostedti Schenkel, 1936, M. minutipalpis and R. hierosolymitanus, as well as one jumping spider Attulus zimmermanni (Simon, 1877), are illustrated with photographs of their epigynes. Keywords: faunistics, Kazakhstan, spider, new records, new indications
--19
© Есюнин С. Л., Кабдрахимов А. А., 2023
For citacion: Esyunin S. L., Kabdrakhimov A. A. New data on the spider fauna of West Kazakhstan Region (Arachnida: Araneae). Bulletin of Perm University. Biology. Iss. 1 (2023): pp. 19-30. (In Russ.). http://dx.doi.org/10.17072/1994-9952-2023-1-19-30.
Acknowledgments: Gyulli Sh. Farzalieva (Perm State University, Perm) assisted in producing digital photographs. Special thanks go to Andrei V. Tanasevitch (A. N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia) for identifying the single female of Mecynargus minutipalpis and to Dmitri V. Logunov (Manchester, UK), who identified the female of Rhysodromus hierosolymitanus and edited the English of the final draft.
Introduction
Regular arachno-faunistic studies of West Kazakhstan Region started only in the 90s of the 20th century. To date, over 65 arachnological papers on this region have been published, of which the majority are taxonomic revisions or reviews of various spider groups. Faunistic studies on spiders of this region are scarce. First, it is worth mentioning a series of works on the fauna of the Dzhanybek field station [Михайлов, 1985; Peterkina, 2009; Питеркина, Михайлов, 2009]. A checklist of spiders of this station accounts for 200 species, of which 9 have been described as new: Devade kazakhstanica Esyunin et Efimik, 2000; Improphantes contus Tanasevitch et Piterkina, 2007; Silometopus crassipedis Tanasevitch et Piterkina, 2007; Talanites mikhailovi Platnick et Ov-tsharenko, 1991; Thanatus mikhailovi Logunov, 1996; Tibiaster djanybekensis Tanasevitch, 1987; Trichoncoides striganovae Tanasevitch et Piterkina, 2012; Trichoncus villius Tanasevitch et Piterkina, 2007; Walckenaeria stepposa Tanasevitch et Piterkina, 2007. Two other relatively well-studied sites are Taipak (Kalmykovo before 1993) and Zhanakazan (Novaya Kazanka before 1993) Villages, from which 40 species have been recorded and four described as new: Alopecosa sciophila Ponomarev, 2008; Micaria charitonovi Mikhailov et Ponomarev, 2008; Thanatus jaikensis Ponomarev, 2007; Zodariellum volgouralensis (Ponomarev, 2007) [Пономарев, 2022]. Yet, the spider fauna of West Kazakhstan Region remains incompletely studied. A total number of the spider species reported from West Kazakhstan Region to date is 252 [Пономарев, 2022]. Compared to arachnofaunas of neighboring Astrakhan', Volgograd, Rostov or Orenburg Regions [Пономарев, 2022; Esyunin, unpublished data], this number of species seems rather low. Thus, it is hardly surprising that relatively short field trips to West Kazakhstan Region by the second author have yielded 22 additional species (see below).
The purpose of this short faunistic note is to provide a list of the species collected during the 2021 and 2022 field trips to west Kazakhstan, with several poorly known species being illustrated and/or commented on.
Material and methods
The spiders reported herein were collected by the second author (AK) during his field trips to West Kazakhstan Region in the 2021 and 2022 summers. Details of all the localities visited are given below.
Localities (Fig. 1):
1. Bokey Orda District, Khan Ordasy Village, 48°46'26"N, 47°26'02"E, IV.2021, VI-VHI.2022.
2. Bokey Orda District, Menay, 48°41'46"N 47°36'22"E, 9.V.2021.
3. Kaztal District, Sakryl Lake, 49°34'52"N 49°21'34"E, IV.2021, VIII.2022.
4. Zhanakala District, Aydin Lake near Zhanakazan Village, 48°56'35"N 49°35'22"E, IV and VI.2021, VI and VIII.2022.
5. Kaztal District, Saryshaganak Lake, 49°27'56"N 49°51'32"E, VI.2021, IV.2022.
6. Zhanakala District, Pyatimar Village, 49°28'32"N 50°31'51"E, VIII.2022.
7. Zhanakala District, Edil'sor Lake, 49°12'33"N 50°44'01"E, IV.2021, IV.2022.
8. Akzhaik District, Zhantemir Village, 49°05'02"N, 51°24'49"E, VIII.2021.
9. Akzhaik District, Bagyrlay River near Makat, 49°03'05"N, 51°41'18"E, VIII.2021.
10. Akzhaik District, Zhanama Lake near Tompak Village, 48°53'44"N, 51°52'32"E, VI.2021, VI.2022.
11. Akzhaik District, Glubinnoe Lake, 50°20'45"N, 51°03'45"E, VI.2021, V and VIII.2022.
12. Terekti District, Bobrovoe Lake near Shagatai Village, 50°26'20"N 51°08'189"E, VI.2022.
13. Terekti District, Shalkar Lake, 50°28'53"N 51°42'53"E, VI.2022.
14. Zhanakala District, Krugloozernoe Village near Uralsk Town, 51°04'31"N 51°16'24"E, VI.2022.
15. Terekti District, Podstepnoe Village, 51°08'17"N 51°29'20"E, V.2022.
16. Bayterek District, Brusyanoe Lake near Ozernoe Village, 51°21'22"N 51°50'53"E, VI.2021, V and VI.2022.
17. Karatobe District, 20 km E Karatobe Village, 49°41'27"N, 53°38'56"E, VI.2021.
18. Shyngyrlau District, Sulukol' Lake, 15 km NE Akshat Village, 50°29'13"N 54°17'15"E, V and VIII.2022.
Species distribution is discussed according to the World Spider Catalog [WSC, 2022]. In order to recognize
new records of studied taxa, all records were checked against the recently published monograph [Пономарев,
2022]. In the following text, the species recorded from West Kazakhstan Region for the first time are marked with asterisks (*).
Fig. 1. Collecting localities of spiders in West Kazakhstan Region [Точки сбора пауков в Западно-Казахстанской области]
Stacks of colour images were manually generated using an Olympus OMD EM-10 digital camera, with a Panasonic Lumix H-H025 25 mm f/1.7 lens mounted on a Zeiss microscope. Digital images were prepared using Photoshop CS6 image stacking software. Studied specimens are shared between the Zoological Museum of the Moscow State University, Moscow, Russia (curator: K.G. Mikhailov) and the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia (curator: D.V. Logunov).
New records and new regional indication
Agelenidae
Tegenaria domestica (Clerck, 1758)* Material. 1?, Bokey Orda District, Khan Ordasy Village, VI-VIII.2022, АК.
Records in Kazakhstan: East Kazakhstan Region [Савельева, 1970, 1976]. Range: cosmopolitan synanthropic.
Araneidae
Gibbaranea ullrichi (Hahn, 1835) Material. 1?, Bokey Orda District, Khan Ordasy Village, weeds on the sands, 3.IV.2021, АК.
Larinioides folium (Schranck, 1803) Material. 1?, Bayterek District, Brusyanoe Lake, 29.VI.2021, АК.
Larinioides ixobolus (Thorell, 1873) Material. 1^, Bayterek District, Brusyanoe Lake, 29.VI.2021, АК.
Mangora acalypha (Walckenaer, 1802) Material. 1?, Bokey Orda District, Khan Ordasy Village, VI-VIn.2022, АК
Neoscona adianta (Walckenaer, 1802) Material. 1?, Akzhaik District, Zhanama Lake, 23.VI.2021, АК.
Neoscona spasskyi (Brignoli, 1983)* Material. 1^, 1?, Bokey Orda District, Khan Ordasy Village, VI-VIII.2022, АК.
Distribution. Previously known from Tajikistan, Kyrgyzstan, Turkmenistan and Iran [Zamani, Marusik, Sestáko-vá, 2020]. New to Kazakhstan, with the current record representing the northernmost limit of the species range.
Dictynidae
Brigittea latens (Fabricius, 1775) Material. 1^, 3?, Shyngyrlau District, Sulukol' Lake, 17.V.2022, АК.
Gnaphosidae
Callilepis nocturna (Linnaeus, 1758) Material. 1?, Akzhaik District, Zhanama Lake, 23.VI.2022, АК.
Civizelotes caucasius (L. Koch, 1866) Material. 2?, Bokey Orda District, Khan Ordasy Village, VI-VIII.2022, АК; 1$, Akzhaik District, Zhanama Lake, 23.VI.2021, АК. 1?, Shyngyrlau District, Sulukol' Lake, 15.VIII.2022, АК.
Gnaphosa cumensis Ponomarev, 1981 Material. 2$$, Zhanakala District, Aydin Lake, 19.IV.2021, АК; 1?, same locality, 25.VI.2021, АК. 1$, Zhanakala District, Edil'sor Lake, bank, 19. IV.2021, АК.
Gnaphosa dolosa O. Herman, 1879 Material. 2?, Shyngyrlau District, Sulukol' Lake, 17.V.2022, АК.
Gnaphosa licenti Schenkel, 1953 Material. 1$, Bayterek District, Brusyanoe Lake, 20.V.2022, АК.
Gnaphosa mongolica Simon, 1895 Material. 1?, Bokey Orda District, Khan Ordasy Village, VI-VIII.2022, АК.
Gnaphosa saurica Ovtsharenko, Platnick et Song, 1992 Material. 1$, 3?, Zhanakala District, Aydin Lake, 24.VI.2022, АК.
Gnaphosa taurica Thorell, 1875 Material. 1$, Akzhaik District, Zhanama Lake, 23.VI.2022, АК.
Marinarozelotes jaxartensis (Kroneberg, 1875)* Material. 1$, Zhanakala District, Edil'sor Lake, 13. IV.2022, АК.
Records in Kazakhstan: Atyrau [Пономарев, Цветков, 2004; Пономарев, Шматко, 2020], Mangystau [Zyuzin, Tarabaev, 1993] and Kostanay Regions [Пономарев, Брагина, 2014]. New to West Kazakhstan Region. The main part of the species range lies in the west-central-ancient Mediterranean; introduced in Hawaii, Central and South America, India, China and Japan.
Micaria gulliae Tuneva et Esyunin, 2003* Material. 2$$, 2??, Bokey Orda District, Khan Ordasy Village, weeds on the sands, 2-10.VI.2021, АК. Records in Kazakhstan: Atyrau [Пономарев, Цветков, 2006] and Kostanay Regions [Пономарев, Брагина, 2014]. New to West Kazakhstan Region. Outside Kazakhstan, the species is known from Astrakhan and Orenburg Regions of Russia [Tuneva, Esyunin, 2002; Пономарев, Алексеев, 2018].
Micaria rossica Thorell, 1875 Material. 1$, Kaztal District, Saryshaganak Lake, 25.VI.2021, АК; 1$, Terekti District, Shalkar Lake,
22.VI.2022, АК.
Nomisia aussereri (L. Koch, 1872) Material. 1?, Zhanakala District, Aydin Lake, 17.VIII.2022, АК; 1?, Akzhaik District, Zhantemir Village, 29.VIII.2021, АК; 2??, Akzhaik District, Bagyrlay River, 2021, АК
Zelotes mundus (Kulczynski, 1897)* Material. 1$, Akzhaik District, Glubinnoe Lake, overgrowth of common cocklebur (Xanthium strumarium),
23.VI.2021, АК.
Records in Kazakhstan: Atyrau [Пономарев, 2022], Kostanay [Пономарев, Брагина, Шматко, 2017] and East Kazakhstan Regions [Eskov, Marusik, 1995]. New to West Kazakhstan Region. Range: west-central Palearctic semiarid.
Linyphiidae
Mecynargus minutipalpis Gnelitsa, 2011* (Fig. 2B) Material. 1?, Zhanakala District, Aydin Lake, 24.VI.2022, АК.
Distribution. New to Kazakhstan. Earlier known from southern regions of Russian Plain [Gnelitsa, 2011; Ta-nasevitch, 2013; Пономарев, Абдурахманов, 2014; Пономарев, Шаповалов, Ивлиев, 2014; Пономарев, Брагина, Шматко, 2017; Polchaninova, Prokopenko, 2017; Пономарев, Алексеев, 2018]. Range: Eastern European semidesert [Kovblyuk, Kastrygina, 2015].
Microlinyphia impigra (O. Pickard-Cambridge, 1871)* Material. 1$, 3??, Terekti District, Shalkar Lake, 22.VI.2022, АК.
Records in Kazakhstan: East Kazakhstan Region [Eskov, Marusik, 1995]. New to West Kazakhstan Region. Range: circumholarctic temperate.
Microlinyphia pusilla (Sundevall, 1830) Material. 1?, Kaztal District, Saryshaganak Lake, 25.VI.2021, АК; 1?, Akzhaik District, Zhanama Lake, 23.VI.2021, АК
Lycosidae
Allohogna singoriensis (Laxmann, 1770) Material. 1?, Akzhaik District, Bagyrlay River, summer 2021, АК.
Alopecosa sciophila Ponomarev, 2008 Material. 2??, Kaztal District, Saryshaganak Lake, 14.IV.2022, АК. Alopecosa taeniopus (Kulczynski, 1895)
Material. 1$, Shyngyrlau District, Sulukol' Lake, bank, 15.IV.2021, АК.
Bogdocosa kronebergi(Andreeva, 1976) Material. 1$, 3??, Bokey Orda District, Khan Ordasy Village, VI-VIn.2022, АК; 1?, Akzhaik District, Zhanama Lake, 23.VI.2021, АК; 1$, 3??, Akzhaik District, Zhanama Lake, 23.VI.2022, АК.
Evippa sjostedti Schenkel, 1936* (Fig. 2A) Material. 1?, Bokey Orda District, Khan Ordasy Village, pine forest, 05.VIII.2022, АК.
Records in Kazakhstan: Atyrau Region [Пономарев, Цветков, 2004]. New to West Kazakhstan Region. Range: central-ancient Mediterranean (= Central Asian) semiarid.
Fig. 2. Epigynes of Evippa sjostedti Schenkel, 1936 (A), Mecynargus minutipalpis Gnelitsa, 2011 (B), Rhysodromus hierosolymitanus (Levy, 1977) (D) and Attulus zimmermanni (Simon, 1877) (C)
[Эпигины Evippa sjostedti Schenkel, 1936 (A), Mecynargus minutipalpis Gnelitsa, 2011 (B), Rhysodromus hierosolymitanus (Levy, 1977) (D) и Attulus zimmermanni (Simon, 1877) (C)]
Evippa turkmenica Sternbergs, 1979* Material. 1?, Bokey Orda District, Khan Ordasy Village, summer 2021, АК.
Records in Kazakhstan: Atyrau [Цветков, Пономарев, Ханов, 2006] and Mangystau Regions [Пономарев, Абдурахманов, 2014]. New to West Kazakhstan Region. Range: central-ancient Mediterranean (= Central Asian) arid.
Lycosapraegrandis C.L. Koch, 1836 Material. 1$, Kaztal District, Sakryl Lake, bank, 18.VIII.2022, АК.
Pardosa agrestis (Westring, 1861) Material. 1$, Zhanakala District, Aydin Lake, 24.VI.2022, АК; 1$, Akzhaik District, Glubinnoe Lake, overgrowth of common cocklebur (Xanthium strumarium), 23.VI.2021, АК; 1$, same locality, 18.V.2022, АК;
1^, Shyngyrlau District, Sulukol' Lake, 17.V.2022, АК; 2??, Shyngyrlau District, Sulukol' Lake, 15.VIII.2022, АК.
Pardosa italica Tongiorgi, 1966* Material. 3??, Zhanakala District, Pyatimar Village, 16.VIII.2022, АК.
Records in Kazakhstan: Atyrau [Пономарев, Цветков, 2004], Kostanay [Пономарев, Брагина, 2014], Akmola [Триликаускас, Любечанский, 2020], East Kazakhstan Region [Fomichev, Marusik, 2013] and Mangystau Regions [Zyuzin, Tarabaev, 1993]. New to West Kazakhstan Region. Range: trans-ancient Mediterranean sub-boreal-semiarid.
Pardosa jaikensis Ponomarev, 2007 Material. 1?, Kaztal District, Saryshaganak Lake, 5.VIII.2021, АК; 1?, Akzhaik District, Glubinnoe Lake, 18.V.2022, АК.
Pardosa luctinosa Simon, 1876* Material. 1^, Kaztal District, Sakryl Lake, bank, 18. IV.2021, АК. 1^, Zhanakala District, Edil'sor Lake, bank, 19.IV.2021, АК. 1?, same locality, 4.VIII.2021, АК; 1?, Terekti District, Shalkar Lake, 22.VI.2022, АК. Records in Kazakhstan: Kostanay [Пономарев, Брагина, 2014], Atyrau [Пономарев, Абдурахманов, 2014] and Zhetysu Regions [Спасский, Шнитников, 1937]. New to West Kazakhstan Region. Range: trans-ancient Mediterranean subboreal-semiarid.
Pardosa mikhailovi Ballarin, Marusik, Omelko et Koponen, 2012* Material. 1^, Bokey Orda District, Menay, 9.V.2021, АК.
Notes. Previously known only from the type locality in Almaty Region [Ballarin et al., 2012]. New to West Kazakhstan Region, with the current material representing the westernmost record of the species.
Pardosa nebulosa (Thorell, 1872) Material. 1?, Akzhaik District, Zhanama Lake, 23.VI.2021, АК;1^, 2??, Akzhaik District, Zhanama Lake, 23.VI.2022, АК; 2??, Bayterek District, Brusyanoe Lake, 20.V.2022, АК; 4??, Terekti District, Podstepnoe Village, 16.V.2022, АК; 2??, Terekti District, Bobrovoe Lake, 20.VI.2022, АК.
Pardosa plumipes (Thorell, 1875) Material. 1^, Zhanakala District, Krugloozernoe Village, 01-10.VI.2022, АК; 1^, 2??, Akzhaik District, Glubinnoe Lake, 18.V.2022, АК; 1?, Akzhaik District, Glubinnoe Lake, 19.VIII.2022, АК; 2??, Bayterek District, Brusyanoe Lake, 20.V.2022, АК; 1^, 2??, Shyngyrlau District, Sulukol' Lake, 17.V.2022, АК
Xerolycosa miniata (C.L. Koch, 1834) Material. 1?, Bokey Orda District, Khan Ordasy Village, VI-VIn.2022, АК; 1^, Terekti District, Bobrovoe Lake, 20.VI.2022, АК.
Oxyopidae
Oxyopes heterophthalmus (Latreille, 1804) Material. 1?, Bayterek District, Brusyanoe Lake, 29.VI.2021, АК
Oxyopes lineatus Latreille, 1806 Material. 3^, 3??, Bokey Orda District, Khan Ordasy Village, VI-VIn.2022, АК; 2??, Akzhaik District, Zhanama Lake, 23.VI.2022, АК; 1?, Bayterek District, Brusyanoe Lake, 25.VIII.2021, АК; 2?, same locality, 21.VI.2022, АК; 2??, Terekti District, Bobrovoe Lake, 20.VI.2022, АК.
Oxyopes maracandensis Charitonov, 1946 Material. 2??, Bokey Orda District, Khan Ordasy Village, pine forest, 05.VIII.2022, АК.
Philodromidae
Philodromuspoecilus (Thorell, 1872)* Material. 2??, Terekti District, Bobrovoe Lake, 20.VI.2022, АК.
Records in Kazakhstan: Almaty [Спасский, Шнитников, 1937] and East Kazakhstan Regions [Савельева, 1970]. New to West Kazakhstan Region. Range: trans-Palearctic temperate.
Rhysodromus hierosolymitanus (Levy, 1977)* (Fig. 2D) Material. 1?, Terekti District, Shalkar Lake, 22.VI.2022, АК.
Distribution. Earlier known from Iran, Israel and the United Arab Emirates [Torabi, Moradmand, Muster, 2019]. New to Kazakhstan, with the current record representing the northernmost limit of the species range.
Rhysodromus histrio (Latreille, 1819) Material. 1?, Bokey Orda District, Khan Ordasy Village, VI-VIn.2022, АК
Rhysodromus timidus (Szita et Logunov, 2008)* Material. 1?, Zhanakala District, Aydin Lake, 24.VI.2022, АК.
Records in Kazakhstan: Atyrau and Almaty Regions [Szita, Logunov, 2008]. New to West Kazakhstan Region. Outside of Kazakhstan, it is known from Dagestan of Russia [Ponomarev, Dvadnenko, 2013] and Pakistan [Logunov, Ballarin, Marusik, 2011]. Thanatus atratus Simon, 1875
Material. 2^, 4??, Bokey Orda District, Khan Ordasy Village, VI-VHI.2022, АК; 1^, Zhanakala District, Edil'sor Lake, 13. IV.2022, АК; 1^, 1?, Akzhaik District, Zhanama Lake, 23.VI.2021, АК; 1?, Karatobe District, 20 km E Karatobe Village, 28.VI.2021, АК; 1?, Akzhaik District, Glubinnoe Lake, 18.V.2022, АК.
Thanatusformicinus (Clerck, 1758)* Material. 1?, Bayterek District, Brusyanoe Lake, 20.V.2022, АК.
Records in Kazakhstan: Kostanay [Пономарев, Брагина, Шматко, 2017], Pavlodar, Almaty [Lyakhov, 1999] and East-Kazakhstan Regions [Савельева, 1970; Fomichev, Marusik, 2013]. New to West Kazakhstan Region. Range: circum-Holarctic polysonal.
Thanatus jaikensis Ponomarev, 2007 Material. 1?, Bokey Orda District, Khan Ordasy Village, VI-VIn.2022, АК.
Thanatus mikhailovi Logunov, 1996 Material. 2^, 3?, Shyngyrlau District, Sulukol' Lake, 17.V.2022, АК.
Thanatus mongolicus (Schenkel, 1936) Material. 1^, 1?, Akzhaik District, Zhanama Lake, 23.VI.2021, АК.
Thanatus oblongiusculus (Lucas, 1846) Material. 1^, Shyngyrlau District, Sulukol' Lake, 17.V.2022, АК.
Thanatus vulgaris Simon, 1870 Material. 1?, Akzhaik District, Glubinnoe Lake, 19.VIII.2022, АК.
Tibellus oblongus (Walckenaer, 1802) Material. 1?, Terekti District, Bobrovoe Lake, 20.VI.2022, АК.
Pisauridae
Pisaura mirabilis (Clerck, 1758) Material. 2??, Zhanakala District, Krugloozernoe Village, 01-10.VI.2022, АК; 1?, Terekti District, Bobrovoe Lake, 20.VI.2022, АК.
Salticidae
Aelurillus m-nigrum Kulczynski, 1891 Material. 2??, Shyngyrlau District, Sulukol' Lake, shores, 15. IV.2021, АК.
Aelurillus v-insignitus (Clerck, 1758) Material. 1^, Bokey Orda District, Khan Ordasy Village, VI-VIII.2022, АК.
Attulus zimmermanni (Simon, 1877) (Fig. 2C) Material. 1?, Bayterek District, Brusyanoe Lake, 20.V.2022, АК.
Heliophanuspotanini Schenkel, 1963* Material. 2??, Bokey Orda District, Khan Ordasy Village, weeds on the sands, 3. IV.2021, АК; 2^, 1?, same locality, VI-VIII.2022, АК.
Records in Kazakhstan: Atyrau [Пономарев, 2022], Zhetysu, Turkistan, Almaty, East Kazakhstan [Logunov, Marusik, 2000] and Mangystau Regions [Zyuzin, Tarabaev, 1993]. New to West Kazakhstan Region. Range: central-ancient Mediterranean (= Central Asian) semiarid.
Mendoza canestrinii (Ninni, 1868)* Material. 1?, Zhanakala District, Aydin Lake, 19.IV.2021, АК;1^, Terekti District, Bobrovoe Lake, 20.VI.2022, АК.
Records in Kazakhstan: Akmola and Turkistan Regions [Logunov, Rakov, 1998]. New to West Kazakhstan Region. Range: trans-Palearctic subboreal-subtropic.
Pellenes allegrii Caporiacco, 1935* Material. 1^, Shyngyrlau District, Sulukol' Lake, 17.V.2022, АК.
Records in Kazakhstan: Kostanay Region [Пономарев, Брагина, Шматко, 2017]. New to West Kazakhstan Region. Range: central-ancient Mediterranean (= Central Asian) semiarid.
Pseudicius encarpatus (Walckenaer, 1802) Material. 1^, Akzhaik District, Zhanama Lake, 23.VI.2022, АК.
Salticus tricinctus (C.L. Koch, 1846)* Material. 1^, Akzhaik District, Zhanama Lake, 23.VI.2022, АК.
Records in Kazakhstan: Atyrau [Пономарев, 2022], Turkistan, Almaty [Logunov, Rakov, 1998] and Mangystau Regions [Пономарев, Абдурахманов, 2014]. New to West Kazakhstan Region. Range: east-central-ancient Mediterranean semiarid.
Theridiidae
Asagena phalerata (Panzer, 1801) Material. 1?, Shyngyrlau District, Sulukol' Lake, 17.V.2022, АК.
Enoplognatha mordax (Thorell, 1875) Material. 1^, Terekti District, Shalkar Lake, 22.VI.2022, АК. Latrodectus tredecimguttatus (P. Rossi, 1790)
Material. 1?, Akzhaik District, Zhantemir Village, 29.VIII.2021, АК; 1 immature б1, Akzhaik District, Glu-binnoe Lake, 18.V.2022, АК.
Steatoda albomaculata (DeGeer, 1778) Material. 1?, Zhanakala District, Aydin Lake, 24.VI.2022, АК.
Steatoda bipunctata (Linnaeus, 1758)* Material. 1?, Akzhaik District, Glubinnoe Lake, 18.V.2022, АК.
Records in Kazakhstan: Almaty Region [Спасский, Шнитников, 1937]. New to West Kazakhstan Region. Range: trans-Palearctic polyzonal; introduced in Canada and south America.
Steatoda grossa (C. L. Koch, 1838)* Material. 1б, 1?, Kaztal District, Saryshaganak Lake, 14. IV.2022, АК.
Records in Kazakhstan: East Kazakhstan [Савельева, 1970, 1976] and Almaty Regions [Темрешев и др., 2016]. New to West Kazakhstan Region. Range: cosmopolitan synanthropic.
Theridion innocuum Thorell, 1875 Material. 1?, Bokey Orda District, Khan Ordasy Village, VI-VIII.2022, АК.
Thomisidae
Bassaniodes robustus (Hahn, 1832) Material. 1б, Akzhaik District, Zhanama Lake, 23.VI.2021, АК.
Ebrechtella tricuspidate (Fabricius, 1775) Material. 1 immature б, Bayterek District, Brusyanoe Lake, 25.VIII.2021, АК.
Psammitis ninnii fusciventris (Crome, 1965) Material. 1б, Akzhaik District, Zhanama Lake, 23.VI.2021, АК; 1?, Karatobe District, 20 km E Karatobe Village, 28.VI.2021, АК.
Synemaplorator (O. Pickard-Cambridge, 1872)* Material. 1?, Shyngyrlau District, Sulukol' Lake, 17.V.2022, АК.
Records in Kazakhstan: Atyrau [Пономарев, 2022], Mangystau [Zyuzin, Tarabaev, 1993] and Almaty Regions [Спасский, Шнитников, 1937; Marusik, Logunov, 1990]. New to West Kazakhstan Region. Range: east-central-ancient Mediterranean semiarid.
Thomisus onustus Walckenaer, 1805 Material. 1?, Bayterek District, Brusyanoe Lake, 20.V.2022, АК; 1?, Bokey Orda District, Khan Ordasy Village, pine forest, 05.VIII.2022, АК.
Titanoecidae
Nurscia albosignata (Simon, 1874) Material. 1б, 1?, Bokey Orda District, Khan Ordasy Village, VI-VIII.2022, АК.
Uloboridae
Uloborus walckenaerius Latreille, 1806 Material. 1б, 3??, Bokey Orda District, Khan Ordasy Village, among the pines, 05.VIII.2022, АК
Conclusion
The present species list consists of 77 species belonging to 14 families, of which one genus (Mecynargus Kulczynski, 1894; Linyphiidae) and three species - M. minutipalpis, N. spasskyi, R. hierosolymitanus - are new to the spider fauna of Kazakhstan, and 19 species are newly found in West Kazakhstan Region. Besides, 86 new district records were provided for 55 species.
Of the studied material, two species records (N. spasskyi, R. hierosolymitanus) represent the northernmost limits of the corresponding species' ranges, those of specie M. minutipalpis - the easternmost range limit, and that P. mikhailovi - the south-westernmost limit.
Prior to this study 252 spider species were known from West Kazakhstan Region [Пономарев, 2022]. Baseed on the present results, a total number of spider species recorded from West Kazakhstan Region has raised to 274 species in 130 genera and 23 families.
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Статья поступила в редакцию 27.12.2022; одобрена после рецензирования 11.01.2023; принята к публикации 22.02.2023.
The article was submitted 27.12.2022; approved after reviewing 11.01.2023; accepted for publication 22.02.2023. Информация об авторах
С. Л. Есюнин - д-р биол. наук, доцент, профессор кафедры зоологии беспозвоночных и водной экологии; А. А. Кабдрахимов - аспирант кафедры зоологии беспозвоночных и водной экологии. Information about the authors
S. L. Esyunin - Doctor of Biology, Associate Professor, Professor of the Department of Invertebrate Zoology and Aquatic Ecology; A. A. Kabdrakhimov - post-graduate student of the Department of Invertebrate Zoology and Aquatic Ecology.
Вклад авторов:
Есюнин С. Л. - концепция исследования, оформление текста, обработка полевого материала.
Кабдрахимов А. А. - изучение литературных источников, оформление первого варианты текста, сбор полевого материала. Contribution of the authors:
Esyunin S. L. - research concept, text design, processing of field material.
Kabdrakhimov A. A. - study of literary sources, design of the first version of the text, collection of field material.
