CC BY
34© Амурский зоологический журнал III(2), 2011.128-162_
УДК 595.763.33 © Amurian zoological journal III(2), 2011.128-162
IRANIAN ROVE BEETLES (COLEOPTERA: STAPHYLINIDAE) N. Samin1, H. Zhou 2, S. Imam1
[Самин Н., Чжоу Х., Имани С. Каталог стафилинид Ирана (Coleoptera: Staphylinidae)]
'Department of Entomology, Islamic Azad University, Science and Research Branch, Tehran, Iran. E-mail: [email protected]; [email protected]
2Institute of Zoology, Chinese Academy of Sciences, Datun Rd., Chaoyang, Beijing, 100101, China; email: [email protected] 'Кафедра энтомологии, Исламский университет Азад, Научно-исследовательское отделение, Тегеран, Иран. E-mail: n_ [email protected]; [email protected]
2Институт зоологии, Академия наук Китайской Народной Республики, ул. Датун, пр. Чаоян, Пекин, 100101, Китай. E-mail: [email protected]
Key words: Rove beetle, Staphylinidae, fauna, catalogue, Iran Ключевые слова: стафилиниды, Staphylinidae, каталог, фауна, Иран
Summary. The catalogue of Iranian rove beetles (Coleoptera: Staphylinidae) which is based on detail study of all available published data is presented in this paper. Totally 716 species from four groups (Omaliine, Tachyporine, Oxyteline, Staphylinine) and 16 subfamilies (including, Aleocharinae, Habrocerinae, Leptotyphylinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Paederinae, Proteininae, Pselaphinae, Scaphidiinae, Scydmaeninae, Staphylininae, Steninae, Tachyporinae and Trichophyinae) are currently recognized as occurring in Iran.
Резюме. На основе тщательного изучения всех доступных публикаций составлен каталог стафилинид Ирана (Coleoptera: Staphylinidae). Всего к настоящему моменту для Ирана указывается 716 видов из 4 внесистематических групп (Omaliine, Tachyporine, Oxyteline, Staphylinine) и 16 подсемейств (Aleocharinae, Habrocerinae, Leptotyphylinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Paederinae, Proteininae, Pselaphinae, Scaphidiinae, Scydmaeninae, Staphylininae, Steninae, Tachyporinae и Trichophyinae).
INTRODUCTION
Rove beetles (Coleoptera: Staphylinidae) are the most diverse family in hexapoda with over than 56.768 valid species in 3.624 genera and 33 subfamilies. These beetles has worldwide distribution but their diversity is more various in tropical and temperate than in arctic regions, and also more diverse in moist forested areas than in drier areas or deserts [Newton and Thayer, 1992; Thayer, 2005]. Rove beetles are known from every type of habitat that beetles occur in, and their diets include just about everything except the living tissues of higher plants. Most rove beetles are predators of insects and other kinds of invertebrates, living in forest leaf litter and similar kinds of decaying plant matter. They are also commonly found under stones, and around freshwater margins. Several types are known to live on ocean shores that are submerged at high tide, several species have adapted to live as inquilines in ant and termite colonies, and some live in mutualistic relationships with mammals whereby they eat fleas and other parasites, benefiting the host. Most species are probably predators of other arthropods as far as known, but large numbers including several entire subfamilies are saprophagous on decaying plant matter (e.g., Osoriinae, Oxytelinae, Piestinae), fungivorous (e.g., Oxyporinae, Scaphidiinae), or have other specialized feeding habits such as pollenfeeding, but very few feed on green plants. A few species are of economic or medical importance, such as species of Paederus spp. [Newton and Chandler, 1989; Smetana, 1995; Herman, 2001]. Although rove beetles appetites for other insects would seem to make them obvious candidates for biological control of pests, and empirically they are believed to be important controls in the wild, experiments with using them have not been notably successful [White, 1983].
Ж
Iran forms a large part of the Iranian plateau, and covers an area of 1,623,779 km2 (Fig. 1). It is bordered in the north by the Caucasus Mts., Middle Asian natural regions and the Caspian Sea (-27 m below sea level); in the west by the Anatolian and Mesopotamian regions; in the east by the eastern part of the Iranian plateau (Afghanistan and adjacent west Pakistan) and the Baluch-Sindian region; and finally in the south by the Persian Gulf and Oman Sea, which are connected by the latter to the Indian Ocean. Seven desert plains and depressions give the landscape a completely different appearance: Dasht-e Kavir in Central Iran, Dasht-e Lut desert, Sistan and Jazmurian depressions in the southeast, Khuzestan plain in the southwest, Moghan steppe in the northwest and the Turkman-Sahra steppe in the northeast. Six main watershed areas are recognized within the country, but the central and southern watersheds in particular often are divided up into 21 by different authors. More than 20 larger lakes form part of the landscape of Iran, with the largest one, the saline Urumia Lake, covering 4,868 km2 in the northwest [Zehzad et al., 2002, Hangay et al., 2005]
The fauna of Iranian Staphylinidae was poorly studied so far and the newest checklist on this important and big family was prepared by Anlas and Newton [2010]. But the mentioned list does not include all the reported data and it lost some species; because there are some journals in Iran which are not available for all the researchers of the world. The objective of this paper is summarizing all the published data on Iranian Staphylinidae and arranging of all the species known from Iran to the present time in an impressive checklist.
MATERIALS AND METHODS
The published data on Staphylinidae family in Iran was
Fig. 1. Map of Iran with boundaries of provinces. Рис. 1. Карта Ирана с обозначением границ провинций.
summarized. The catalogue comprises the following data: the valid taxa name, published records with provincial distribution, or where this information is not available - "Not cited" or "no locality cited" are given. Iranian provinces are given in alphabetical order in the lists. In the catalogue, the family Staphylinidae is divided into four informal groups of subfamilies (Omaliine, Tachyporine, Oxyteline and Staphylinine Groups, following Lawrence & Newton [1982], and the subfamilies are presented according to Herman [2001] and Lobl & Smetana [2004]. Supertribe, tribe, subtribe, genus and species taxa are presented in alphabetical order. The classification generally follows Lobl & Smetana [2004] except for certain more recent changes, including at higher levels the reduction of Scydmaenidae to a subfamily of Staphylinidae and reduction of its former subfamilies to supertribes [Grebenikov & Newton, 2009] and changes in the tribal classification of Oxytelinae [Makranczy, 2006], as well as changes or additions at the generic and specific levels resulting from recent revisions.
RESULTS
The present catalogue includes 716 species from 170 genera of Iranian Staphylinidae. The list of species is given below with distributional data in Iran.
Omaliine Group
1. Subfamily Omaliinae Macleay 1825 Tribe Anthophagini Thomson 1859 Anthobium (Anthobium) atrocephalum (Gyllenhal 1827) Distribution in Iran: East Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Anthobium (Anthobium) melanocephalum (Illiger 1794)
Distribution in Iran: Golestan.
Iranian record: Ghahari et al. [2009c].
Deliphrosoma bakhtiyariense Zerche 1991
Distribution in Iran: Chahar-Mahal Bakhtiari.
Iranian record: Zerche [1991], Bordoni [1999b], Herman
[2001], Smetana [2004]. Endemic.
Deliphrosoma freudeorum Zerche 1996
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b].
Deliphrosoma linbergi (Scheerpeltz 1958)
Distribution in Iran: Khuzestan.
Iranian record: Ghahari et al. [2009d].
Deliphrosoma morvani (Jarrige 1971)
Distribution in Iran: Mazandaran.
Iranian record: Jarrige [1971], Zerche [1991], Bordoni
[1999b], Herman [2001], Smetana [2004]. Endemic.
Deliphrosomaprolongatum (Rottenberg 1873)
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b].
Deliphrosoma weiratheri (Scheerpeltz 1938)
Distribution in Iran: Mazandaran, Semnan.
Iranian record: Sakenin et al. [2008b, 2010].
Geodromicus asiaticus Bernhauer 1902
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Sakenin et al. [2008b, c].
Geodromicus iranicus Coiffait 1981
Distribution in Iran: Tehran.
Iranian record: Coiffait [1981a], Herman [2001], Smetana [2004]. Endemic.
Geodromicus jarrigeanus Fagel 1976
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b].
Geodromicus major (Motschulsky 1860)
Distribution in Iran: East Azarbaijan, Mazandaran.
Iranian record: Sakenin et al. [2008b], Ghahari et al. [2009b].
Geodromicus zwickianus Fagel 1976
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b].
Lesteva morvani Jarrige 1971
Distribution in Iran: Mazandaran.
Iranian record: Jarrige [1971], Smetana [2004]. Endemic.
Olophrum bodemeyeri Scheerpeltz 1929
Distribution in Iran: Lorestan.
Iranian record: Scheerpeltz [1929], Herman [2001], Smetana [2004]. Endemic. Olophrum fuscum (Gravenhorst 1806) Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1929], Herman [2001], Smetana [2004].
Olophrum puncticolle Eppelsheim 1880 Distribution in Iran: Mazandaran. Iranian record: Ghahari et al. [2009c]. Orochares villiersi Jarrige 1971 Distribution in Iran: Fars.
Iranian record: Jarrige [1971], Thayer [1993], Herman [2001], Smetana [2004]. Endemic.
Tribe Coryphiini Jakobson 1908 Coryphiodes jelineki Zerche 1990 Distribution in Iran: Tehran.
Zerche [1990, 1993], Herman [2001], Smetana [2004]. Endemic.
Tribe Eusphalerini Hatch 1957 Eusphalerum bothmeri Coiffait 1976 Distribution in Iran: Guilan.
Iranian record: Coiffait [1976], Zanetti [1993], Herman [2001], Smetana [2004].
Eusphalerum caucasicum caucasicum (Bernhauer 1908)
Distribution in Iran: Semnan.
Iranian record: Sakenin et al. [2010].
Eusphalerum celsum (Luze 1910)
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b].
Eusphalerum foveolatum (Luze 1910)
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b], Ghahari et al. [2009c].
Eusphalerumprimulae (Stephens 1834)
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b].
Eusphalerum rufoscutellatum (Eppelsheim 1881)
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Ghahari et al. [2009c], Sakenin et al. [2010].
Eusphalerum sareptanum (Eppelsheim 1878)
Distribution in Iran: Ardabil, East Azarbaijan. Iranian record: Ghahari et al. [2009b]. Eusphalerum transcaucasicum (Bernhauer 1902) Distribution in Iran: Mazandaran, Semnan. Iranian record: Sakenin et al. [2008b, 2010]. Eusphalerum sorbi (Gyllenhal 1810) Distribution in Iran: Golestan.
Iranian record: Zanetti [1993], Herman [2001], Smetana [2004]. Eusphalerum wittmeri Coiffait 1976 Distribution in Iran: Guilan.
Iranian record: Coiffait [1976], Zanetti [1993], Herman [2001], Smetana [2004]. Endemic.
Tribe Omaliini Macleay 1825 Dropephylla caucasica (Kolenati 1846) Distribution in Iran: Not cited. Iranian record: Anlaç & Newton [2010].
Dropephylla vilis (Erichson 1840)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Omalium caesum Gravenhorst 1806
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b].
Omalium cribriceps Fauvel 1900
Distribution in Iran: North Iran (no locality cited).
Iranian record: Fauvel [1900], Luze [1906], Herman [2001].
Omalium falsum Eppelsheim 1889
Distribution in Iran: East Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Omalium littorale Kraatz 1857
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b].
Omalium oxyacanthae Gravenhorst 1806
Distribution in Iran: Golestan.
Iranian record: Ghahari et al. [2009c].
Omalium rivulare (Paykull 1789)
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b].
Omalium saulcyi Fauvel 1875
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008b].
Phyllodrepa gracilicornis (Fairmaire & Laboulbène 1856)
Distribution in Iran: Semnan.
Iranian record: Sakenin et al. [2010].
2. Subfamily Proteininae Eeichson 1839 Megarthrus denticollis (Beck 1817) Distribution in Iran: East Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Tribe Proteinini Erichson 1839 Proteinus atomarius Erichson 1840 Distribution in Iran: Isfahan. Iranian record: Ghahari et al. [2009c].
3. Subfamily Pselaphinae Latreille 1802
Supertribe Batrisitae Reitter 1882 Tribe Batrisini Reitter 1882 Batrisodes insularis (Baudidi Selve 1870) Distribution in Iran: Chahar-Mahal Bakhtiari.
Iranian record: Ghahari et al. [2009c]. Batrisodes ruprechtii (Kolenati 1846) Distribution in Iran: Golestan, Guilan, Mazandaran. Iranian record: Besuchet [1981a], Löbl & Besuchet [2004].
Supertribe Clavigeritae Leach 1815 Tribe Clavigerini Leach 1815 Articerodes syriacus (Saulcy 1865) Distribution in Iran: Not cited.
Iranian record: Besuchet [1986], Löbl & Besuchet [2004].
Claviger caspicus Reitter 1882
Distribution in Iran: Not cited.
Iranian record: Löbl & Besuchet [2004].
Claviger colchicus Motschulsky 1837
Distribution in Iran: Not cited.
Iranian record: Löbl & Besuchet [2004].
Claviger katharinae Escherich 1897
Distribution in Iran: Mazandaran.
Iranian record: Ghahari et al. [2009c].
Claviger merkli Reitter 1885
Distribution in Iran: West Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Claviger testaceuspersicus B. Bodemeyer 1927
Distribution in Iran: North Iran (no locality cited).
Iranian record: B. Bodemeyer [1927], Löbl & Besuchet
[2004]. Endemic.
Supertribe Euplectitae Leconte 1861 Tribe Euplectini Leconte 1861
Euplectus caspicus Raffray 1910 Distribution in Iran: North Iran (no locality cited). Iranian record: Raffray [1910]. Endemic. Euplectus karstenii (Reichenbach 1816) Distribution in Iran: Not cited. Iranian record: Löbl & Besuchet [2004]. Euplectus occipitalis Reitter 1882 Distribution in Iran: Not cited. Iranian record: Löbl & Besuchet [2004]. Euplectus verticalis Reitter 1884 Distribution in Iran: Mazandaran. Iranian record: Ghahari et al. [2009c].
Tribe Trichonychini Reitter 1882 Amauronyx talyschensis (Reitter 1882) Distribution in Iran: Not cited. Iranian record: Löbl & Besuchet [2004]. Bibloplectus pauxillus Besuchet 1975 Distribution in Iran: Guilan, Mazandaran. Iranian record: Besuchet [1975a], Löbl & Besuchet [2004]. Bibloplectus spinosus Raffray 1914 Distribution in Iran: Central Alborz (no locality cited). Iranian record: Hematyian et al. [2010]. Bibloporus variicolor Reitter 1882 Distribution in Iran: North Iran (no locality cited). Iranian record: Reitter [1882], Löbl & Besuchet [2004]. Panaphantus atomus Kiesenwetter 1858 Distribution in Iran: Guilan, Lorestan, Tehran. Iranian record: Besuchet [1980], Löbl & Besuchet [2004], Ghahari et al. [2009c].
Plectophloeus nitidus (Fairmaire 1858) Distribution in Iran: Semnan.
Iranian record: Löbl & Besuchet [2004], Sakenin et al. [2010]. Plectophloeus nubigena nubigena (Reitter 1877) Distribution in Iran: East Azarbaijan, West Azarbaijan. Iranian record: Löbl & Besuchet [2004], Ghahari et al. [2009b]. Pseudoplectus perplexus (Jacquelin du val 1854) Distribution in Iran: Not cited. Iranian record: Löbl & Besuchet [2004]. Scotoplectus caspicus Besuchet 1975 Distribution in Iran: Ardabil.
Iranian record: Besuchet [1975b], Löbl & Besuchet [2004]. Trichonyx georgicus (Motschulsky 1845) Distribution in Iran: Not cited. Iranian record: Löbl & Besuchet [2004]. Zibus leiocephalus (Aube 1833) Distribution in Iran: East Azarbaijan. Iranian record: Löbl & Besuchet [2004], Ghahari et al. [2009b]. Supertribe Goniaceritae Reitter 1882 Tribe Brachyglutini Raffray 1904
Brachygluta afganistanica Löbl 1967
Distribution in Iran: Golestan, Khorasan, Mazandaran,
Tehran.
Iranian record: Löbl & Besuchet [2004], Sabella et al. [2004]. Brachygluta araxidis (Reitter 1889) Distribution in Iran: East Azarbaijan, West Azarbaijan, Fars, Guilan.
Iranian record: Löbl & Besuchet [2004], Sabella et al. [2004]. Brachygluta fossulata (Reichenbach 1816) Distribution in Iran: East Azarbaijan. Iranian record: Ghahari et al. [2009b]. Brachygluta foveola (Motschulsky 1840) Distribution in Iran: North Iran (no locality cited). Iranian record: Löbl & Besuchet [2004], Sabella et al. [2004]. Brachygluta furcata (Motschulsky 1835) Distribution in Iran: North Iran (no locality cited). Iranian record: Löbl & Besuchet [2004], Sabella et al. [2004]. Brachygluta giordanii Meggiolaro 1961 Distribution in Iran: Kermanshah. Iranian record: Meggiolaro [1961], Löbl & Besuchet [2004]. Endemic.
Brachygluta helferi longispina (Reitter 1884) Distribution in Iran: Northwest Iran (no locality cited). Iranian record: Löbl & Besuchet [2004], Sabella et al. [2004]. Brachygluta iranica (Saulcy 1876) Distribution in Iran: Tehran.
Iranian record: Saulcy [1876], Reitter [1882], Karaman [1961], Sabella et al. [2004]. Brachygluta larica Sabella 2004
Distribution in Iran: Hormozgan, Mazandaran, Sistan & Baluchestan.
Iranian record: Sabella et al. [2004]. Endemic. Brachygluta martina Bückle 2004 Distribution in Iran: Lorestan. Iranian record: Sabella et al. [2004]. Endemic. Brachygluta perissinottoi Besuchet 1969 Distribution in Iran: Fars.
Iranian record: Besuchet [1969], Löbl & Besuchet [2004]. Endemic.
Brachygluta richteri Besuchet 1961 Distribution in Iran: Khuzestan.
Iranian record: Besuchet [1961], Löbl & Besuchet [2004], Sabella et al. [2004].
Brachygluta spinicoxis spinicoxis (Motschulsky 1835) Distribution in Iran: Isfahan.
Iranian record: Saulcy [1876], Reitter [1882], Besuchet
[1999], Löbl & Besuchet [2004].
Brachygluta tumidipes Besuchet 1981
Distribution in Iran: Hormozgan, Kerman, Lorestan.
Iranian record: Besuchet [1981b], Löbl & Besuchet [2004],
Sabella et al. [2004].
Fagniezia spinipes (Reitter 1885)
Distribution in Iran: Not cited.
Iranian record: Löbl & Besuchet [2004].
Reichenbachia chevrieri (Aubé 1844)
Distribution in Iran: Mazandaran.
Iranian record: Ghahari et al. [2009a].
Rybaxis diabolica Reitter 1884
Distribution in Iran: Not cited.
Iranian record: Löbl & Besuchet [2004].
Rybaxis gigas (Baudi di Selve 1870)
Distribution in Iran: Not cited.
Iranian record: Reitter [1882], Löbl & Besuchet [2004].
Rybaxis longicornis (Leach 1817)
Distribution in Iran: Mazandaran.
Iranian record: Löbl & Besuchet [2004], Ghahari et al.
[2009a].
Tribatus creticus Reitter 1884
Distribution in Iran: East Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Trissemus dentipes (Baudi di Selve 1870)
Distribution in Iran: Semnan.
Iranian record: Sakenin et al. [2010].
Trissemus montanus (Saulcy 1876)
Distribution in Iran: Not cited.
Iranian record: Löbl & Besuchet [2004].
Trissemus mundulus Besuchet 1961
Distribution in Iran: Sistan & Baluchestan.
Iranian record: Besuchet [1961], Löbl & Besuchet [2004].
Endemic.
Tribe Bythinini Raffray 1890 Bryaxis anatolicus (Saulcy 1878) Distribution in Iran: Ardabil, East Azarbaijan. Iranian record: Ghahari et al. [2009b]. Bryaxis theanus (Reitter 1894) Distribution in Iran: Semnan. Iranian record: Sakenin et al. [2010]. Bythinus gracilis Motschulsky 1851 Distribution in Iran: North Iran (no locality cited). Iranian record: Karaman [1957], Löbl & Besuchet [2004]. Tychobythinus caviceps (Reitter 1882) Distribution in Iran: Not cited. Iranian record: Löbl & Besuchet [2004].
Tribe Tychini Raffray 1904 Tychus caspicus Besuchet & Sabella 1999 Distribution in Iran: Guilan, Mazandaran. Iranian record: Besuchet & Sabella [1999a], Löbl & Besuchet [2004]. Endemic. Tychus dubius Sabella & Kurbatov 2003 Distribution in Iran: Kermanshah. Iranian record: Sabella & Kurbatov [2003]. Endemic. Tychus holzschuhi Besuchet & Sabella 1999 Distribution in Iran: Ardabil.
Iranian record: Besuchet & Sabella [1999b], Löbl &
Besuchet [2004].
Tychus lederi Saulcy 1878
Distribution in Iran: Golestan.
Iranian record: Sabella & Kurbatov [2003], Löbl & Besuchet [2004].
Tychus persicus Besuchet & Sabella 1999
Distribution in Iran: Chahar-Mahal Bakhtiari.
Iranian record: Besuchet & Sabella [1999a], Löbl &
Besuchet [2004]. Endemic.
Tychus remaudierei Besuchet 1969
Distribution in Iran: Khorasan.
Iranian record: Besuchet [1969], Löbl & Besuchet [2004]. Tychus sengleti Besuchet & Sabella 1999 Distribution in Iran: Mazandaran. Iranian record: Besuchet & Sabella [1999b], Löbl & Besuchet [2004]. Endemic.
Supertribe Pselaphitae Latreille 1802 Tribe Ctenistini É. Blanchard 1845
Chennium antennatum Reitter 1882
Distribution in Iran: Not cited.
Iranian record: Löbl & Besuchet [2004].
Ctenisomorphus major (Raffray 1877)
Distribution in Iran: Not cited.
Iranian record: Löbl & Besuchet [2004].
Ctenistes marthae Reitter 1891
Distribution in Iran: Not cited.
Iranian record: Löbl & Besuchet [2004].
Ctenistes palpalis Reichenbach 1816
Distribution in Iran: Not cited.
Iranian record: Reitter [1882], Löbl & Besuchet [2004].
Ctenistes staudingerii (L. W. Schaufuss 1861) Distribution in Iran: Not cited.
Iranian record: Besuchet [1981b], Löbl & Besuchet [2004].
Desimia darius (Saulcy 1874)
Distribution in Iran: Not cited.
Iranian record: Löbl & Besuchet [2004].
Enoptostomus globulicornis (Motschulsky 1851)
Distribution in Iran: Not cited.
Iranian record: Reitter [1882], Löbl & Besuchet [2004].
Mentraphus pselaphodes Sharp 1883
Distribution in Iran: Not cited.
Iranian record: Besuchet [1981b], Löbl & Besuchet [2004].
Tribe Pselaphini Latreille 1802 Pselaphidius acuminatus (Motschulsky 1835) Distribution in Iran: Not cited.
Iranian record: Löbl & Besuchet [2004]. Pselaphus acuminatus Motschulsky 1835 Distribution in Iran: Semnan. Iranian record: Sakenin et al. [2010]. Pselaphus caspicus Reitter 1882 Distribution in Iran: Not cited. Iranian record: Löbl & Besuchet [2004]. Pselaphus striatus Löbl 1974 Distribution in Iran: Not cited. Iranian record: Löbl & Besuchet [2004].
Tachyporine Group
4. Subfamily Habrocerinae Mulsant & Rey 1877 Habrocerus capillaricornis (Gravenhorst 1806) Distribution in Iran: Isfahan, Mazandaran. Iranian record: Assing & Wunderle [1995, 1996], Smetana [2004], Ghahari et al. [2009c].
5. Subfamily Aleocharinae Fleming 1821 Tribe Aleocharini Fleming 1821 Aleochara adusta Eppelsheim 1890 Distribution in Iran: Guilan. Iranian record: Assing [2009l]. Aleochara bilineata Gyllenhal 1810 Distribution in Iran: Mazandaran, Tehran. Iranian record: Likovsky [1981], Smetana [2004]. Aleochara bipustulata (Linnaeus 1760) Distribution in Iran: East Azerbaijan, Fars, Lorestan, Mazandaran, Tehran.
Iranian record: Jarrige [1971], Likovsky [1981], Maus
[1998], Smetana [2004].
Aleochara brevilaminata Assing 2009
Distribution in Iran: Kermanshah, Khorasan.
Iranian record: Assing [2009b].
Aleochara brevipennis Gravenhorst 1806
Distribution in Iran: Semnan.
Iranian record: Sakenin et al. [2010].
Aleochara clavicornis Redtenbacher 1849
Distribution in Iran: Ardabil.
Iranian record: Smetana [2004], Tronquet [2009].
Aleochara cuniculorum Kraatz 1858
Distribution in Iran: Fars.
Iranian record: Assing [2009b].
Aleochara erythroptera Gravenhorst 1806
Distribution in Iran: Chahar-Mahal Bakhtiari.
Iranian record: Ghahari et al. [2009c].
Aleochara falsa Likovsky 1981
Distribution in Iran: Fars, Isfahan, Lorestan.
Iranian record: Likovsky [1981], Assing [2009b]. Endemic.
Aleochara gaudiuscula Tottenham 1939
Distribution in Iran: East Azerbaijan.
Iranian record: Likovsky [1981], Smetana [2004].
Aleochara haematoptera Kraatz 1858
Distribution in Iran: Tehran.
Iranian record: Likovsky [1981], Smetana [2004].
Aleochara ignipennis Fauvel 1900
Distribution in Iran: North Iran (no locality cited).
Iranian record: Fauvel [1900], Assing [2007a, 2009b].
Aleochara inconspicua Aubé 1850
Distribution in Iran: Tehran.
Iranian record: Likovsky [1981], Smetana [2004].
Aleochara intricate Mannerheim 1830
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Jarrige [1971], Likovsky [1981],
Smetana [2004].
Aleochara laevigata Gyllenhal 1810
Distribution in Iran: Lorestan, Mazandaran, Tehran.
Iranian record: Likovsky [1981], Smetana [2004].
Aleochara lanuginose Gravenhorst 1802
Distribution in Iran: Mazandaran.
Iranian record: Jarrige [1971], Likovsky [1981], Smetana
[2004].
Aleochara laticornis Kraatz 1856
Distribution in Iran: Golestan.
Iranian record: Likovsky [1981], Smetana [2004].
Aleochara lepidoptera Bernhauer 1901
Distribution in Iran: Mazandaran.
Iranian record: Likovsky [1981], Assing [2009b].
Aleochara leptocera Eppelsheim 1889
Distribution in Iran: Isfahan.
Iranian record: Assing [2009b].
Aleochara maculipennis Baudi di selve 1857
Distribution in Iran: Kermanshah, Lorestan.
Iranian record: Assing [2009b].
Aleochara milleri Kraatz 1862
Distribution in Iran: East Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Aleochara moesta Gravenhorst 1802
Distribution in Iran: Guilan, Kordestan, Lorestan,
Mazandaran, Tehran.
Iranian record: Scheerpeltz [1963b], Jarrige [1971],
Likovsky [1981], Smetana [2004].
Aleochara persica Likovsky 1981
Distribution in Iran: Mazandaran.
Iranian record: Likovsky [1981], Smetana [2004], Assing
[2009b].
Aleocharapuberula Klug 1834 Distribution in Iran: Mazandaran. Iranian record: Likovsky [1981], Smetana [2004]. Aleochara sarica Assing 2009 Distribution in Iran: Mazandaran. Iranian record: Assing [2009l]. Endemic. Aleochara spadicea (Erichson 1837) Distribution in Iran: North Iran (no locality cited). Iranian record: Scheerpeltz [1969]. Aleochara transcaspica Bernhauer 1936 Distribution in Iran: Khorasan.
Iranian record: Bernhauer [1936], Smetana [2004]. Endemic.
Aleochara tristis Gravenhorst 1806
Distribution in Iran: Mazandaran.
Iranian record: Jarrige [1971], Likovsky [1981], Smetana
[2004].
Aleochara verna Say 1833
Distribution in Iran: North Iran (no locality cited),
Mazandaran.
Iranian record: Maus [1998], Smetana [2004], Ghahari et al. [2009c].
Piochardia reitteri (Wasmann 1894)
Distribution in Iran: Tehran.
Iranian record: Likovsky [1981], Smetana [2004].
Tribe Athetini Casey 1910 Acrotona parvula (Mannerheim 1830) Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Acrotonapseudopygmaea (Scheerpeltz 1963) Distribution in Iran: Mazandaran. Iranian record: Ghahari et al. [2009d]. Acrotona pygmaea (Gravenhorst 1802) Distribution in Iran: Semnan. Iranian record: Sakenin et al. [2010]. Aloconota cambrica (Wollaston 1855) Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Aloconota gregaria (Erichson 1839) Distribution in Iran: Fars, Isfahan. Iranian record: Scheerpeltz [1961b], Jarrige [1971]. Aloconata iranica (Scheerpeltz 1961) Distribution in Iran: Khuzestan. Iranian record: Scheerpeltz [1961a]. Endemic. Aloconota luristanica Smetana 2004 Distribution in Iran: Lorestan.
Iranian record: G. Benick [1941], Smetana [2004]. Endemic.
Aloconota persica (Cameron 1940) Distribution in Iran: Kerman.
Iranian record: Cameron [1940], Smetana [2004]. Endemic. Amischa analis (Gravenhorst 1802) Distribution in Iran: Sistan & Baluchestan. Iranian record: Scheerpeltz [1961a], Smetana [2004].
Atheta aeneipennis (Thomson 1856)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Atheta atramentaria (Gyllenhal 1810)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Atheta castanoptera (Mannerheim 1840)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Atheta elongatula (Gravenhorst 1802)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Atheta judith (Saulcy 1865)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Atheta longicornis (Gravenhorst 1802)
Distribution in Iran: East Azarbaijan, Mazandaran.
Iranian record: Jarrige [1971], Ghahari et al. [2009b].
Atheta oblita (Erichson 1839)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Atheta oraria (Kraatz 1856)
Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1961d], Smetana [2004].
Atheta scrobicollis (Kraatz 1859) Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Atheta trinotata (Kraatz 1856) Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Atheta volans (Scriba 1859)
Distribution in Iran: East Azarbaijan, West Azarbaijan. Iranian record: Atamehr et al. [2004], Ghahari et al. [2009b].
Callicerus rigidicornis (Erichson 1839) Distribution in Iran: Mazandaran. Iranian record: Ghahari et al. [2009a]. Geostiba circellaris (Gravenhorst 1806) Distribution in Iran: Isfahan. Iranian record: Assing [2009f]. Geostiba huberi Pace 1983 Distribution in Iran: Khorasan.
Iranian record: Pace [1983], Smetana [2004], Assing [2005d]. Endemic.
Geostiba impressiventris Assing 2009 Distribution in Iran: Guilan. Iranian record: Assing [2009f]. Endemic. Geostiba sarica Assing 2009 Distribution in Iran: Mazandaran. Iranian record: Assing [2009f]. Endemic. Geostiba sengleti Pace 1983 Distribution in Iran: Golestan.
Iranian record: Pace [1983], Smetana [2004], Assing
[2005d, 2008g]. Endemic.
Liogluta longiuscula (Gravenhorst 1802)
Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1961d], Smetana [2004].
Mocyta fungi fungi (Gravenhorst 1806)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Nehemitropia lividipennis (Mannerheim 1830)
Distribution in Iran: Tehran.
Iranian record: Bohac [1981], Smetana [2004].
Ousipalia caesula (Erichson 1839)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Pseudosemiris kaufmanni (Eppelsheim 1887)
Distribution in Iran: East Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Tomoglossa laeta Eppelsheim 1884
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Tribe Falagriini Mulsant & Rey 1873 Anaulacaspis gratilla (Erichson 1839) Distribution in Iran: South Iran (no locality cited). Iranian record: Fagel [1969], Scheerpeltz [1963a], Smetana [2004].
Anaulacaspis lutzi (Reitter 1909)
Distribution in Iran: East Iran (no locality cited). Iranian record: Scheerpeltz [1960a], Smetana [2004].
Anaulacaspis naevula (Erichson 1839) Distribution in Iran: Kerman, Sistan & Baluchestan. Iranian record: Scheerpeltz [1961a, 1963a], Smetana [2004].
Anaulacaspispamphylica (Fagel 1969)
Distribution in Iran: Semnan.
Iranian record: Sakenin et al. [2010].
Anaulacaspispersica (Fagel 1969)
Distribution in Iran: North and South Iran (no locality cited).
Iranian record: Fagel [1969], Smetana [2004]. Endemic.
Anaulacaspis seclusa (Fagel 1969) Distribution in Iran: South Iran (no locality cited). Iranian record: Fagel [1969], Smetana [2004].
Cordalia fortepunctata Assing 2006
Distribution in Iran: Ardabil, Chahar-Mahal Bakhtiari, East
Azerbaijan, Fars, Guilan, Isfahan, Lorestan, Mazandaran,
Tehran.
Iranian record: Assing [2007e].
Cordalia obscura (Gravenhorst 1802)
Distribution in Iran: East Azerbaijan, Fars, Guilan,
Mazandaran.
Iranian record: Assing [2007e].
Falagria caesa Erichson 1837
Distribution in Iran: East Azarbaijan.
Iranian record: Smetana [2004], Atamehr et al. [2004].
Falagria collaris Reitter 1891
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Falagria splendens Kraatz 1858
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Falagrioma thoracica (Stephens 1832)
Distribution in Iran: East Azerbaijan, Guilan, Mazandaran.
Iranian record: Assing [2007e].
Myrmecopora effeminate Assing 2004
Distribution in Iran: Lorestan.
Iranian record: Assing [2007e, 2009d].
Myrmecopora uvida (Erichson 1840)
Distribution in Iran: East Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Tribe Homalotini Heer 1839 Caloderina hierosolymitana (Saulcy 1865) Distribution in Iran: Fars. Iranian record: Assing [2007e]. Leptusa armeniaca Pace 1989 Distribution in Iran: East Azerbaijan, West Azerbaijan. Iranian record: Assing [2009j].
Leptusa (Neopisalia) confinis Pace 1982 Distribution in Iran: Mazandaran. Iranian record: Sakenin et al. [2008d]. Leptusa delphiniformis Assing 2009 Distribution in Iran: Mazandaran. Iranian record: Assing [2009j]. Endemic. Leptusa flagrifera Assing 2009
Distribution in Iran: Guilan. Iranian record: Assing [2009j]. Endemic. Leptusa (Dysleptusa) fuliginosa (Aube 1850) Distribution in Iran: Mazandaran. Iranian record: Sakenin et al. [2008d]. Leptusa laeviuscula (Hochhuth 1849) Distribution in Iran: Mazandaran. Iranian record: Assing [2009j]. Leptusa persica Assing 2009 Distribution in Iran: Mazandaran. Iranian record: Assing [2009j]. Endemic. Leptusa piciformis Assing 2009 Distribution in Iran: Guilan. Iranian record: Assing [2009j]. Endemic. Leptusa sengleti Pace 1984 Distribution in Iran: Guilan.
Iranian record: Pace [1984, 1989], Assing [2009j]. Endemic.
Leptusa soriciformis Assing 2009 Distribution in Iran: Mazandaran. Iranian record: Assing [2009j]. Endemic.
Tribe Lomechusini Fleming 1821 Drusilla alutacea Reitter 1901 Distribution in Iran: Mazandaran. Iranian record: Assing [2008e]. Drusilla bulbata Assing 2008 Distribution in Iran: Mazandaran. Iranian record: Assing [2008e]. Endemic. Drusilla canaliculata (Fabricius 1787) Distribution in Iran: Ardabil, East Azerbaijan, Guilan, Mazandaran, Tehran. Iranian record: Assing [2005c, 2008e]. Drusilla ganglbaueri (Bernhauer 1906) Distribution in Iran: Khorasan, Semnan. Iranian record: Assing [2008e]. Drusilla heydeni (Eppelsheim 1887) Distribution in Iran: Khorasan, Mazandaran. Iranian record: Assing [2008e]. Drusilla persica Assing 2005
Distribution in Iran: Golestan, Khorasan, Mazandaran. Iranian record: Assing [2005c, 2008e]. Endemic.
Pella similis (Märkel 1844) Distribution in Iran: Semnan. Iranian record: Sakenin et al. [2010]. Zyras fulgidus (Gravenhorst 1806) Distribution in Iran: Not cited.
Iranian record: Smetana [2004], Hlavac & Jaszay [2009].
Tribe Mesoporini Cameron 1959 Dictyon pumilio (Eppelsheim 1884) Distribution in Iran: Ardabil, Guilan. Iranian record: Bohac [1980], Assing [2010b].
Tribe Myllaenini Ganglbauer 1895 Myllaena intermedia Erichson 1837 Distribution in Iran: Mazandaran. Iranian record: Sakenin et al. [2008d], Ghahari et al. [2009a]. Tribe Oxypodini Thomson 1859
Amarochara umbrosa (Erichson 1837)
Distribution in Iran: Khorasan.
Iranian record: Assing [2010f].
Devia prospera (Erichson 1839)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Gnypeta carbonaria (Mannerheim 1830)
Distribution in Iran: East Azarbaijan.
Iranian record: Smetana [2004], Atamehr et al. [2004].
Gnypeta rubrior Tottenham 1939
Distribution in Iran: East Azerbaijan.
Iranian record: Assing [2009d].
Ocalea persica Coiffait 1973
Distribution in Iran: Kerman.
Iranian record: Coiffait [1973], Smetana [2004]. Endemic.
Oxypoda alternans (Gravenhorst 1802)
Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1961d], Smetana [2004].
Oxypoda brevicornis (Stephens 1832)
Distribution in Iran: Mazandaran.
Iranian record: Assing [2007e].
Oxypoda ignorata Zerche 1996
Distribution in Iran: Ghazvin, Hamadan.
Iranian record: Assing [2007e].
Oxypoda rubustior Assing 2009
Distribution in Iran: Semnan.
Iranian record: Assing [2009k]. Endemic.
Oxypoda vicina Kraatz 1858
Distribution in Iran: East Azerbaijan.
Iranian record: Jarrige [1971].
Oxypoda wankai Bernhauer 1936
Distribution in Iran: Fars.
Iranian record: Assing [2007e].
Ischnopoda umbratica (Erichson 1837)
Distribution in Iran: East Azarbaijan, Mazandaran.
Iranian record: Sakenin et al. [2008d], Ghahari et al. [2009a].
Tachyusa nitella Fauvel, 1895
Distribution in Iran: Mazandaran.
Iranian record: Pasnik [2006].
Tetralaucopora bicolorata Assing 2007
Distribution in Iran: Fars.
Iranian record: Assing [2007e]. Endemic.
Tetralaucopora longitarsis (Erichson 1839)
Distribution in Iran: Ardabil, Fars, Hamadan, Isfahan,
Kerman, Tehran.
Iranian record: Smetana [2004], Assing [2007e]. Tetralaucopora rubicunda (Erichson 1837) Distribution in Iran: Ardabil, East Azerbaijan, Mazandaran, Tehran.
Iranian record: Assing [2007e].
Tribe Pronomaeini Mulsant & Rey 1873 Pronomaea araxicola Reitter 1898 Distribution in Iran: Fars. Iranian record: Smetana [2004], Assing [2007h]. Pronomaea khnzoriani Semenov 2003 Distribution in Iran: Ardabil, Chahar-Mahal Bakhtiari, East
Azerbaijan, Fars, Guilan, Hamadan, Isfahan, Kermanshah, Khorasan, Kordestan, Lorestan, Mazandaran, Tehran. Iranian record: Smetana [2004], Assing [2007e, h].
Pronomaea procerula Assing 2007
Distribution in Iran: Kuhkuloyeh & Boyer Ahmad,
Kordestan.
Iranian record: Assing [2007e]. Endemic. Pronomaea rostrata Erichson 1837 Distribution in Iran: Semnan. Iranian record: Smetana [2004], Sakenin et al. [2010]. 6. Subfamily Tachyporinae Macleay 1825 Tribe Mycetoporini Thomson 1859 Bolitobius insignis Hochhuth 1849 Distribution in Iran: West Azarbaijan. Iranian record: Ghahari et al. [2009b]. Carphacis jarrigei Schülke 1994 Distribution in Iran: North Iran (no locality cited). Iranian record: Schülke [1994, 2000], Herman [2001], Smetana [2004]. Endemic. Ischnosoma longicorne (Maklin 1847) Distribution in Iran: East Azarbaijan. Iranian record: Ghahari et al. [2009b]. Ischnosoma splendidum (Gravenhorst 1806) Distribution in Iran: East Azarbaijan. Iranian record: Schülke [2008]. Lordithon thoracicus thoracicus (Fabricius 1777) Distribution in Iran: Tehran.
Iranian record: Smetana [2004], Anlaç & Newton [2010].
Lordithon trimaculatus (Fabricius 1793)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Lordithon trinotatus (Erichson 1839)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Mycetoporus baudueri Mulsant & Rey 1875
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Mycetoporus brucki (Pandellé 1869)
Distribution in Iran: Semnan.
Iranian record: Smetana [2004], Sakenin et al. [2010].
Mycetoporus confuses Eppelsheim 1889
Distribution in Iran: North Iran (no locality cited).
Iranian record: Eppelsheim [1889].
Mycetoporus lepidus (Gravenhorst 1806)
Distribution in Iran: Ardabil.
Iranian record: Smetana [2004], Ghahari et al. [2009b].
Mycetoporus liliputanus Luze 1901
Distribution in Iran: Fars.
Iranian record: Schülke [2008].
Mycetoporus reichei (Pandelle 1869)
Distribution in Iran: East Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Tribe Tachyporini Macleay 1825 Cilea silphoides (Linnaeus 1767) Distribution in Iran: Not cited. Iranian record: Herman [2001].
Coproporus colchicus Kraatz 1858
Distribution in Iran: Golestan.
Iranian record: Schülke [2007], Smetana [2004].
Lamprinus erythropterus Panzer 1796) Distribution in Iran: Hamadan. Iranian record: Schülke [2008]. Sepedophilus immaculatus (Stephens 1832) Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Sepedophilus littoreus (Linnaeus 1758) Distribution in Iran: Not cited. Iranian record: Herman [2001], Smetana [2004].
Sepedophilus luzei Schülke 1998
Distribution in Iran: Mazandaran.
Iranian record: Schülke [1998], Herman [2001], Smetana
[2004].
Sepedophilus pedicularius (Gravenhorst 1802) Distribution in Iran: Kerman.
Iranian record: Fauvel [1875], Scheerpeltz [1961a], Herman [2001].
Sepedophilus rufulus (Hochhuth 1849)
Distribution in Iran: Sistan & Baluchestan.
Iranian record: Scheerpeltz [1960a, 1963a], Herman
[2001], Smetana [2004].
Sepedophilus testaceus (Fabricius 1793)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Tachinus corticinus Gravenhorst 1802
Distribution in Iran: Lorestan.
Iranian record: Schülke [2008].
Tachinus fanynkae Hromadka 2001
Distribution in Iran: Mazandaran.
Iranian record: Hromadka [2001], Smetana [2004].
Endemic.
Tachinus fimetarius Gravenhorst 1802 Distribution in Iran: Not cited.
Iranian record: Ullrich [1975], Herman [2001], Smetana [2004].
Tachinus humeralis humeralis Gravenhorst 1802
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Tachinus rufipes (Linnaeus 1758)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Tachinus rufitarsis Hochhuth 1849
Distribution in Iran: Not cited.
Iranian record: Schülke [1989b].
Tachyporus gracilicornis Luze 1904
Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1960a], Herman [2001].
Tachyporus hypnorum (Fabricius 1775)
Distribution in Iran: Golestan, Lorestan, Sistan &
Baluchestan, Tehran.
Iranian record: Fauvel [1875], Scheerpeltz [1961a], Jarrige [1971], Herman [2001], Smetana [2004], Sakenin et al. [2008c], Anlaç & Newton [2010]. Tachyporus nitidulus (Fabricius 1781)
Distribution in Iran: Fars.
Iranian record: Fauvel [1875], Jarrige [1971], Herman
[2001], Smetana [2004].
Tachyporus pusillus Gravenhorst 1806
Distribution in Iran: Guilan.
Iranian record: Schülke [2008].
Tachyporus scitulus Erichson 1839
Distribution in Iran: Not cited.
Iranian record: Herman [2001], Smetana [2004].
Tachyporus solutus Erichson 1839 Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Tachyporus transversalis Gravenhorst 1806 Distribution in Iran: North Iran (no locality cited). Iranian record: Scheerpeltz [1963a].
7. Subfamily Trichophyinae Thompson 1859 Trichophya pilicornis (Gyllenhal 1810) Distribution in Iran: Mazandaran. Iranian record: Ghahari et al. [2009c].
Oxyteline Group
8. Subfamily Osoriinae Erichson 1839
Tribe Osoriini Erichson 1839 Leptotyphlopsis iranica Coiffait 1977 Distribution in Iran: Ilam.
Iranian record: Coiffait [1977a], Smetana [2004]. Endemic.
9. Subfamily Oxytelinae Fleming 1821
Tribe Blediini Âdam 20 Bledius akinini Eppelsheim 1888 Distribution in Iran: Not cited. Iranian record: Herman [1986, 2001], Smetana [2004].
Bledius angustus angustus Mulsant & Rey 1861 Distribution in Iran: Khuzestan, Sistan & Baluchestan. Iranian record: Scheerpeltz [1961a], Smetana [2004].
Bledius angustus pierrei Jarrige 1971 Distribution in Iran: Not cited.
Iranian record: Jarrige [1971], Smetana [2004]. Endemic.
Bledius atricapillus (Germar 1825)
Distribution in Iran: East Azarbaijan, West Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Bledius bicornis (Germar 1823)
Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1961b].
Bledius capra Fauvel 1875
Distribution in Iran: Hormozgan, Kerman.
Iranian record: Jarrige [1971], Smetana [2004].
Bledius carinicollis Fauvel 1875
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Bledius dilutipennis Motschulsky 1858
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Bledius dissimilis Erichson 1840
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Bledius fossor Heer 1839
Distribution in Iran: Not cited.
Iranian record: Fauvel [1872], Herman [2001], Smetana [2004].
Bledius furcatus (Olivier 1811) Distribution in Iran: Mazandaran. Iranian record: Anlaç & Newton [2010]. Bledius glasunovi Luze 1904
Distribution in Iran: Hormozgan, Sistan & Baluchestan. Iranian record: Scheerpeltz [1960b, 1961a, 1963a], Jarrige [1971], Herman [2001], Smetana [2004]. Bledius hinnulus Erichson 1840
Distribution in Iran: Hormozgan, Khuzestan, Sistan & Baluchestan.
Iranian record: Scheerpeltz [1960b, 1961a, 1963a], Fagel
[1970], Jarrige [1971], Herman [2001], Smetana [2004]. Bledius immarginatus Koch 1934
Distribution in Iran: Not cited.
Iranian record: Herman [2001], Smetana [2004].
Bledius klapperichi Coiffait 1981 Distribution in Iran: Guilan.
Iranian record: Coiffait [1981a], Herman [2001], Smetana [2004]. Endemic.
Bledius lindbergianus Scheerpeltz 1963 Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Bledius loeffleri Coiffait 1973 Distribution in Iran: Kerman.
Iranian record: Coiffait [1973], Herman [1986, 2001],
Smetana [2004]. Endemic.
Bledius opacus (Block 1799)
Distribution in Iran: Not cited.
Iranian record: Herman [2001], Smetana [2004].
Blediuspersicus Bernhauer 1940 Distribution in Iran: Not cited.
Iranian record: Bernhauer [1940], Herman [1970, 1986, 2001], Smetana [2004]. Endemic. Bledius richteri Scheerpeltz 1961 Distribution in Iran: Khuzestan.
Iranian record: Scheerpeltz [1961a], Herman [1970, 1986,
2001], Smetana [2004]. Endemic.
Bledius spectabilis Kraatz 1857
Distribution in Iran: Khuzestan, Hormozgan.
Iranian record: Scheerpeltz [1961a], Horion [1963], Jarrige
[1971], Herman [2001], Smetana [2004]. Bledius tibialis Heer 1839 Distribution in Iran: Ardabil.
Iranian record: Ghahari et al. [2009b]. Bledius tricornis (Herbst 1784) Distribution in Iran: Not cited. Iranian record: Herman [2001], Smetana [2004].
Bledius verres Erichson 1840 Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1961b], Herman [2001],
Smetana [2004].
Bledius vitulus Erichson 1840
Distribution in Iran: Hormozgan, Khuzestan.
Iranian record: Scheerpeltz [1960b, 1961a], Jarrige [1971],
Herman [2001], Smetana [2004].
Eppelsheimius miricollis (Fauvel 1898) Distribution in Iran: Not cited.
Iranian record: Jarrige [1971], Herman [1983, 2001], Smetana [2004].
Eppelsheimiuspirazzolii (Eppelsheim 1885) Distribution in Iran: Kerman, Hormozgan, Markazi. Iranian record: Scheerpeltz [1958b], Jarrige [1971], Herman [1983, 2001], Kashcheev [1988], Smetana [2004].
Tribe Coprophilini Heer 1839 Coprophiluspennifer (Motschulsky 1845) Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1960a, 1963a], Herman
[2001], Smetana [2004].
Coprophilus rufipennis (Reitter 1894)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Tribe Deleasterini Reitter 1909 Deleaster dichrous (Gravenhorst 1802) Distribution in Iran: Not cited.
Iranian record: Horion [1963], Herman [2001], Smetana [1967, 2004].
Tribe Oxytelini Fleming 1821 Anotylus complanatus (Erichson 1839) Distribution in Iran: Not cited.
Iranian record: Fauvel [1872], Scheerpeltz [1962], Horion
[1963], Herman [2001], Smetana [2004].
Anotylus hybridus (Eppelsheim 1878)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Anotylus intricatus (Erichson 1840)
Distribution in Iran: Mazandaran.
Iranian record: Fauvel [1872], Scheerpeltz [1962], Horion
[1963], Herman [2001], Smetana [2004], Anlaç & Newton
[2010].
Anotylus inustus (Gravenhorst 1806)
Distribution in Iran: East Azarbaijan, West Azarbaijan.
Iranian record: Fauvel [1872], Horion [1963], Jarrige
[1971], Herman [2001], Smetana [2004].
Anotylus luridipennis (Luze 1904)
Distribution in Iran: Isfahan.
Iranian record: Scheerpeltz [1960a], Jarrige [1971], Herman [2001], Smetana [2004]. Anotylus nitidulus (Gravenhorst 1802) Distribution in Iran: Golestan.
Iranian record: Fauvel [1872], Scheerpeltz [1960a, 1963a],
Jarrige [1971], Herman [2001], Smetana [2004].
Anotylus politus (Erichson 1840)
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008d].
Anotylus pumilus (Erichson 1839)
Distribution in Iran: Not cited.
Iranian record: Fauvel [1872], Scheerpeltz [1962], Horion [1963], Herman [2001], Smetana [2004]. Anotylus sculpturatus (Gravenhorst 1806) Distribution in Iran: Guilan, Hormozgan, Khorasan. Iranian record: Fauvel [1872], Jarrige [1971], Herman [2001], Smetana [2004].
Anotylus sexualis (Eppelsheim 1892) Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1962, 1963a], Herman [2001], Smetana [2004].
Anotylus speculifrons (Kraatz 1857)
Distribution in Iran: Not cited.
Iranian record: Fauvel [1872], Herman [2001].
Anotylus tetracarinatus (Block 1799) Distribution in Iran: Tehran.
Iranian record: Fauvel [1872], Herman [2001], Anlaç & Newton [2010].
Carpelimus anthracinus (Mulsant & Rey 1861)
Distribution in Iran: Kerman, Khuzestan.
Iranian record: Scheerpeltz [1961a], Gildenkov [2001b],
Herman [2001], Smetana [2004].
Carpelimus bilineatus Stephens 1834
Distribution in Iran: Fars.
Iranian record: Fauvel [1871], Herman [2001], Smetana [2004], Gildenkov [2009a]. Carpelimus corticinus (Gravenhorst 1806) Distribution in Iran: Kerman, Khuzestan, Northeastern Iran (no locality cited).
Iranian record: Scheerpeltz [1961a], Gildenkov [2001a], Herman [2001].
Carpelimus despectus despectus (Baudi di Selve 1870) Distribution in Iran: Khuzestan.
Iranian record: Scheerpeltz [1961a], Herman [2001], Smetana [2004].
Carpelimus exiguous (Erichson 1839)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Carpelimus gracilis (Mannerheim 1830
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Carpelimus gusarovi Gildenkov 1997
Distribution in Iran: Northeast Iran (no locality cited).
Iranian record: Gildenkov [2001a].
Carpelimus insularis (Kraatz 1858)
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Jarrige [1971], Gildenkov [2001a], Smetana
[2004].
Carpelimus iraniensis Herman 1970 Distribution in Iran: Khuzestan.
Iranian record: Gildenkov [2001a], Scheerpeltz [1961a],
Herman [1970, 2001], Smetana [2004].
Carpelimus nitidus (Baudi di Selve 1848)
Distribution in Iran: Not cited.
Iranian record: Coiffait [1981a], Smetana [2004].
Carpelimus obesus (Kiesenwetter 1844)
Distribution in Iran: Kerman, Khuzestan.
Iranian record: Scheerpeltz [1961a], Gildenkov [2001a],
Herman [2001], Smetana [2004].
Carpelimus pusillus (Gravenhorst 1802)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Carpelimus simplex (Motschulsky 1857)
Distribution in Iran: Southeast Iran (no locality cited)
Iranian record: Scheerpeltz [1960b], Herman [2001], Smetana [2004].
Carpelimus simplicicollis simplicicollis (Wollaston 1857)
Distribution in Iran: Khuzestan.
Iranian record: Gildenkov [2009b].
Carpelimus tener (Bernhauer 1902)
Distribution in Iran: Hormozgan.
Iranian record: Jarrige [1971], Herman [2001], Smetana [2004], Gildenkov [2009a]. Carpelimus transversicollis (Scheerpeltz 1947) Distribution in Iran: Northeast Iran (no locality cited). Iranian record: Gildenkov [2001a].
Carpelimus troglodytes troglodytes (Erichson 1840) Distribution in Iran: Kerman, Khuzestan. Iranian record: Scheerpeltz [1961a], Herman [2001], Smetana [2004].
Oxytelus incisus Motschulsky 1858 Distribution in Iran: Hormozgan.
Iranian record: Fauvel [1872], Jarrige [1971], Herman [2001].
Oxytelus piceus (Linnaeus 1767) Distribution in Iran: Guilan, Mazandaran. Iranian record: Jarrige [1971], Herman [2001]. Oxytelus sculptus Gravenhorst 1806 Distribution in Iran: Tehran.
Iranian record: Fauvel [1872], Bohac [1981], Herman [2001].
Platystethus arenarius (Geoffroy 1785)
Distribution in Iran: Mazandaran.
Iranian record: Jarrige [1971], Herman [2001], Smetana
[2004].
Platystethus capito capito Heer 1839
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Platystethus cornutus cornutus (Gravenhorst 1802)
Distribution in Iran: Kordestan, North Iran (no locality cited).
Iranian record: Kolenati [1846], Fauvel [1872], Scheerpeltz
[1960a, 1963a], Jarrige [1971], Herman [2001], Smetana
[2004].
Platystethus degener (Mulsant & Rey 1878) Distribution in Iran: Fars, East Azarbaijan. Iranian record: Atamehr et al. [2004], Schülke [2009]. Platystethus nitens (C. R. Sahlberg 1832) Distribution in Iran: Not cited.
Iranian record: Fauvel [1872], Herman [2001], Smetana [2004].
Platystethus nodifrons Mannerheim 1830
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008d], Ghahari et al. [2009a].
Platystethus rufospinus Hochhuth 1851
Distribution in Iran: Not cited.
Iranian record: Fauvel [1872], Herman [2001], Smetana [2004].
Platystethus spinosus Erichson 1840 Distribution in Iran: Not cited.
Iranian record: Fauvel [1872], Scheerpeltz [1955], Horion [1963], Herman [2001], Smetana [2004].
Thinobius brevipennis Kiesenwetter 1850 Distribution in Iran: Semnan. Iranian record: Sakenin et al. [2010]. Thinobius brundini Scheerpeltz 1959 Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Thinobius nodicornis Eppelsheim 1884 Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Thinodromus arcuatus (Stephens 1834) Distribution in Iran: West Azarbaijan. Iranian record: Horion [1963], Herman [2001], Smetana [2004], Ghahari et al. [2009b]. Thinodromus coloripennis (Scheerpeltz 1963) Distribution in Iran: Khorasan. Iranian record: Gildenkov [2001b], Smetana [2004]. Thinodromus dilatatus (Erichson 1839) Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Thinodromus mannerheimii (Kolenati 1846) Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Thinodromus pilosellus (Eppelsheim 1890) Distribution in Iran: Not cited. Iranian record: Gildenkov [2001b], Smetana [2004]. 10. Subfamily Scaphidiinae Latreille 1807 Tribe Scaphisomatini Casey 1894 Baeocera schirmeri Reitter 1881 Distribution in Iran: Not cited. Iranian record: Löbl [2004].
Scaphisoma assimile assimile Erichson 1845 Distribution in Iran: Not cited. Iranian record: Löbl [2004].
Scaphisoma boleti dilutum Reitter 1885 Distribution in Iran: East Azarbaijan. Iranian record: Löbl [2004], Ghahari et al. [2009b]. Scaphisoma laeviusculum Reitter 1898 Distribution in Iran: Guilan, Golestan. Iranian record: Löbl [1970, 2004]. Scaphisoma subalpinum subalpinum Reitter 1881 Distribution in Iran: Not cited. Iranian record: Löbl [1970, 2004].
Staphylinine Group
11. Subfamily Leptotyphylinae Fauvel 1874 Tribe Entomoculiini Coiffait 1957 Kenotyphlus pisidicus Coiffait 1957 Distribution in Iran: Isfahan. Iranian record: Ghahari et al. [2009c].
12. Subfamily Oxyporinae Fleming 1821 Oxyporus rufus rufus (Linnaeus 1758) Distribution in Iran: Not cited. Iranian record: Bernhauer [1923], Herman [2001].
13. Subfamily Scydmaeninae LEACH 1815 Supertribe Mastigitae Fleming 1821 Tribe Leptomastacini Casey 1897 Leptomastax hyrcana Castellini 1996
Distribution in Iran: Mazandaran.
Iranian record: Castellini [1996], Vit [2004]. Endemic.
Supertribe Scydmaenitae Leach 1815 Tribe Chevrolatiini Reitter 1882 Chevrolatia egregia Reitter 1881 Distribution in Iran: Tehran. Iranian record: Orousset [2008].
Tribe Cyrtoscydmini Schaufuss 1889 Euconnus goleanus Franz 1981 Distribution in Iran: Golestan. Iranian record: Franz [1981], Davies [2004]. Endemic.
14. Subfamily Steninae Macleay 1825 Stenus aereus Solsky 1871
Distribution in Iran: East Azarbaijan, West Azarbaijan.
Iranian record: Puthz [1970], Ryvkin [1990], Herman
[2001], Smetana [2004].
Stenus affaber Baudi di selve 1848
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenus alienigenus Puthz 1964
Distribution in Iran: Kerman.
Iranian record: Puthz [2006].
Stenus arabicus Saulcy 1865
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenus araxis Ryvkin 1990
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenus armeniacus Puthz 1967
Distribution in Iran: Markazi.
Iranian record: Puthz [1973], Herman [2001], Smetana [2004].
Stenus asiaticus Bernhauer 1940
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Bernhauer [1940], Jarrige [1971], Puthz
[1979b], Ryvkin [1990], Herman [2001], Smetana [2004].
Stenus assequens assequens Rey 1884
Distribution in Iran: Guilan.
Iranian record: Puthz [1977].
Stenus ater Mannerheim 1830
Distribution in Iran: Mazandaran.
Iranian record: Smetana [2004], Ghahari et al. [2009c].
Stenus atratulus Erichson 1839
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenus barbarae Hromadka 1979
Distribution in Iran: Guilan, Tehran.
Iranian record: Hromadka [1979], Puthz [1979b, 1980b],
Bohac [1980], Ryvkin [1990], Herman [2001], Smetana
[2004]. Endemic.
Stenus bidenticulatus Puthz 1970
Distribution in Iran: Not cited.
Iranian record: Herman [2001], Smetana [2004].
Stenus binotatus Ljungh 1804 Distribution in Iran: West Azarbaijan. Iranian record: Ghahar et al. [2009b].
Stenus brunnipes brunnipes Stephens 1833
Distribution in Iran: Khorasan, Mazandaran.
Iranian record: Puthz [2001], Smetana [2004].
Stenus callidus Baudi di selve 1848
Distribution in Iran: Chahar-Mahal Bakhtiari, East
Azarbaijan, Fars, Guilan, Hamadan, Ilam, Isfahan,
Kermanshah, Khuzhestan, Kordestan, Kuhkuloyeh &
Boyer Ahmad, Lorestan, Mazandaran, Tehran, West
Azarbaijan.
Iranian record: Smetana [2004], Puthz [2008].
Stenus canescens Rosenhauer 1856
Distribution in Iran: Semnan.
Iranian record: Sakenin et al. [2010].
Stenus caspius Puthz 1972
Distribution in Iran: Guilan.
Iranian record: Bohac [1980], Herman [2001].
Stenus circularis Gravenhorst 1802
Distribution in Iran: Golestan.
Iranian record: Ghahari et al. [2009c].
Stenus claritarsis Puthz 1971
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenus confrater Eppelsheim 1890
Distribution in Iran: Guilan.
Iranian record: Puthz [1972b], Ryvkin [1990], Herman
[2001], Smetana [2004], Puthz [2009].
Stenus cordatoides Puthz 1972
Distribution in Iran: Not cited.
Iranian record: Puthz [1972b], Herman [2001].
Stenus crassus Stephens 1833
Distribution in Iran: East Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Stenus cyaneus Baudi de Selve 1848
Distribution in Iran: East Azarbaijan.
Iranian record: Sakenin et al. [2008c].
Stenus darius Puthz 2009
Distribution in Iran: Guilan.
Iranian record: Puthz [2009]. Endemic.
Stenus derwisch Puthz 1981
Distribution in Iran: Guilan.
Iranian record: Puthz [1981], Ryvkin [1990], Herman [2001], Smetana [2004], Puthz [2009]. Endemic. Stenus erythrocnemus Eppelsheim 1884 Distribution in Iran: Fars, Guilan, Ilam, Kerman, Kermanshah, Khuzestan, Kuhkuloyeh & Boyer Ahmad, Tehran.
Iranian record: Puthz [1970], Ryvkin [1990], Herman
[2001], Smetana [2004], Assing [2005e].
Stenus fornicatus Stephens 1833
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenus ganglbaueri Bernhauer 1905
Distribution in Iran: Mazandaran.
Iranian record: Puthz [1972a, 1983], Ryvkin [1990],
Herman [2001], Smetana [2004].
Stenus guilanensis Puthz 1979
Distribution in Iran: Guilan.
Iranian record: Puthz [1979a], Ryvkin [1990], Herman
[2001], Smetana [2004]. Endemic.
Stenus guttula P. Müller 1821
Distribution in Iran: Sistan & Baluchestan.
Iranian record: Scheerpeltz [1961a].
Stenus heinzianus Puthz 1970
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Puthz [1970, 1973, 1983], Ryvkin [1990],
Herman [2001], Smetana [2004].
Stenus ignotus Eppelsheim 1890
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenus incrassatus Erichson 1839
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenus kambyses Puthz 2009
Distribution in Iran: Guilan.
Iranian record: Puthz [2009]. Endemic.
Stenus latifrons Erichson 1839
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenus limicola Korge 1967
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenus maculiger Weise 1875
Distribution in Iran: Mazandaran.
Iranian record: Ghahari et al. [2009c].
Stenus martensi Puthz 1983
Distribution in Iran: Mazandaran.
Iranian record: Puthz [1983, 2009], Ryvkin [1990], Herman
[2001], Smetana [2004]. Endemic.
Stenus medus Puthz 1981
Distribution in Iran: East Azarbaijan, Guilan, Tehran.
Iranian record: Puthz [1981], Ryvkin [1990], Herman
[2001], Smetana [2004].
Stenus melanarius melanarius Stephens 1833
Distribution in Iran: Not cited.
Iranian record: Herman [2001], Smetana [2004].
Stenus melanopus Marsham 1802
Distribution in Iran: Not cited.
Iranian record: Fauvel [1873], Herman [2001], Smetana [2004].
Stenus mongolicus Eppelsheim 1889
Distribution in Iran: Khorasan, Guilan.
Iranian record: Puthz [1980a, 2009], Herman [2001],
Smetana [2004], Ghahari et al. [2009d].
Stenus morio Gravenhorst 1806
Distribution in Iran: East Azarbaijan, West Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Stenus nodipes Puthz 1972
Distribution in Iran: Ilam, Kermanshah, Kordestan.
Iranian record: Puthz [2009].
Stenus ochropus Kiesenwetter 1858
Distribution in Iran: East Azarbaijan.
Iranian record: Puthz [1983], Ryvkin [1990], Herman
[2001], Smetana [2004].
Stenus orientis Puthz 1967
Distribution in Iran: Fars, Kermanshah, Khuzestan.
Iranian record: Puthz [2006].
Stenuspallitarsis Stephens 1833
Distribution in Iran: Guilan.
Iranian record: Jarrige [1971], Herman [2001].
Stenuspaludicola Kiesenwetter 1858
Distribution in Iran: Ardabil.
Iranian record: Ghahari et al. [2009b].
Stenus parcior Bernhauer 1929
Distribution in Iran: Lorestan.
Iranian record: Puthz [2008].
Stenusperipherus Korge 1971
Distribution in Iran: Guilan.
Iranian record: Puthz [2008].
Stenus persicus Puthz 1981
Distribution in Iran: East Azarbaijan, Kermanshah.
Iranian record: Puthz [1981], Ryvkin [1990], Herman
[2001]. Endemic.
Stenus pieperi Puthz 1983
Distribution in Iran: Mazandaran.
Iranian record: Puthz [1983, 2009], Ryvkin [1990], Herman
[2001], Smetana [2004]. Endemic.
Stenuspiscator Saulcy 1865
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Stenusplanifronsplanifrons Rey 1884
Distribution in Iran: Semnan.
Iranian record: Sakenin et al. [2010].
Stenus planifrons robustus L. Benick 1914
Distribution in Iran: Not cited.
Iranian record: Ryvkin [1990], Herman [2001], Smetana [2004].
Stenusproprius L. Benick 1921
Distribution in Iran: Khorasan.
Iranian record: Puthz [1972b, 2003], Smetana [2004].
Stenuspseudofossulatus Scheerpeltz 1960
Distribution in Iran: East Azarbaijan.
Iranian record: Sakenin et al. [2008c].
Stenuspusillus Stephens 1833
Distribution in Iran: Isfahan.
Iranian record: Ghahari et al. [2009c].
Stenus ressli Puthz 1979
Distribution in Iran: Mazandaran.
Iranian record: Puthz [1979a, 2009], Ryvkin [1990],
Herman [2001], Smetana [2004]. Endemic.
Stenus schah Puthz 1981
Distribution in Iran: Fars, Lorestan, Tehran.
Iranian record: Puthz [1981], Ryvkin [1990], Herman
[2001], Smetana [2004]. Endemic.
Stenus similis (Herbst 1784)
Distribution in Iran: Not cited.
Iranian record: Ménétriés [1832], Herman [2001], Smetana [2004].
Stenus simonae Hromadka 1979 Distribution in Iran: Guilan. Iranian record: Bohac [1980].
Comment: The presence of this species in Iran is doubtful
because it is known only from Macedonia [Smetana 2004, Anlas & Newton 2010].
Stenus skoraszewskyi Korge 1971 Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Stenus stigmula Erichson 1840 Distribution in Iran: East Azarbaijan. Iranian record: Ghahari et al. [200b]. Stenus turbulentus Bondroit 1912 Distribution in Iran: Semnan. Iranian record: Sakenin et al. [2010]. Stenus turk Puthz 1972 Distribution in Iran: Not cited.
Iranian record: Ryvkin [1990], Herman [2001], Smetana [2004].
Stenus viti Puthz 1977 Distribution in Iran: Guilan.
Iranian record: Puthz [1977], Ryvkin [1990], Herman [2001], Smetana [2004]. Endemic. Stenus wittmeri Puthz 1972 Distribution in Iran: Golestan.
Iranian record: Puthz [1972b], Ryvkin [1990], Herman
[2001], Smetana [2004]. Endemic.
Stenus xerxes Puthz 2009
Distribution in Iran: Mazandaran
Iranian record: Puthz [2009]. Endemic.
15. Subfamily Paederinae Fleming 1821 Tribe Paederini Fleming 1821 Subtribe Astenina Hatch 1957 Astenus gracilis (Paykull 1789) Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Astenus indicus (Kraatz 1859) Distribution in Iran: Not cited. Iranian record: Scheerpeltz [1963b]. Astenus klapperichi Coiffait 1980 Distribution in Iran: Mazandaran. Iranian record: Coiffait [1980, 1984], Smetana [2004]. Endemic.
Astenus lithocharoides (Solsky 1874)
Distribution in Iran: Kordestan, North Iran (no locality
cited).
Iranian record: Fauvel [1900], Scheerpeltz [1963a], Coiffait [1971b, 1984], Jarrige [1971], Smetana [2004]. Astenus lyonessius (Joy 1908)
Distribution in Iran: East Azarbaijan, Golestan, Semnan. Iranian record: Sakenin et al. [2008c, d, 2010].
Astenus melanurus (Küster 1853)
Distribution in Iran: Hormozgan, Kerman, Mazandaran.
Iranian record: Scheerpeltz [1961a], Coiffait [1971b,
1984], Jarrige [1971], Smetana [2004], Sakenin et al.
[2008c, d], Anlaç & Newton [2010].
Astenus morvani Jarrige 1971
Distribution in Iran: East Azarbaijan, Mazandaran.
Iranian record: Jarrige [1971], Coiffait [1984], Smetana
[2004].
Astenus pictipennis Fauvel 1900
Distribution in Iran: Kordestan, North and Central Iran (no locality cited).
Iranian record: Fauvel [1900], Scheerpeltz [1963a],
Coiffait [1971b, 1984], Jarrige [1971], Smetana [2004].
Astenus procerus (Gravenhorst 1806)
Distribution in Iran: West Azarbaijan, Zanjan.
Iranian record: Sakenin et al. [2008c, d].
Astenus thaboris (Saulcy 1865)
Distribution in Iran: Mazandaran.
Iranian record: Ghahari et al. [2009c].
Astenus theodoridesi Jarrige 1971
Distribution in Iran: Kordestan.
Iranian record: Jarrige [1971], Coiffait [1984], Smetana [2004]. Endemic.
Astenus thoracicus (Baudi di Selve 1857) Distribution in Iran: Isfahan, Mazandaran. Iranian record: Sakenin et al. [2008c, d], Anlaç & Newton [2010].
Subtribe Cryptobiina Casey 1905 Afrobium arabicum Assing 2008 Distribution in Iran: Hormozgan. Iranian record: Assing [2008c]. Cryptobium fracticorne (Paykull 1800) Distribution in Iran: East Azarbaijan. Iranian record: Ghahari et al. [2009b]. Homaeotarsus chaudoirii Hochhuth 1851 Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Homaeotarsus iranoturcestanicus (Scheerpeltz 1956) Distribution in Iran: Guilan, Sistan & Baluchestan. Iranian record: Scheerpeltz [1961a, 1963a], Jarrige [1971], Smetana [2004].
Ochthephilum besucheti (Bordoni 1980) Distribution in Iran: Mazandaran. Iranian record: Assing [2009c]. Ochthephilum egregium (Reitter 1884) Distribution in Iran: Guilan, Mazandaran. Iranian record: Coiffait [1984], Smetana [2004]. Ochthephilum fracticorne (Paykull 1800) Distribution in Iran: East Azarbaijan, Khuzestan. Iranian record: Scheerpeltz [1961a], Smetana [2004], Sakenin et al. [2008c, d]. Ochthephilum permutatum Assing 2009 Distribution in Iran: Guilan. Iranian record: Assing [2009c]. Ochthephilum turkestanicum (Korge 1968) Distribution in Iran: East Azerbaijan, Fars, Golestan, Hormozgan, Khorasan, Semnan.
Iranian record: Coiffait [1984], Smetana [2004], Assing [2007e, 2008f, 2009c].
Subtribe Dolicaonina Casey 1905 Leptobium assingi Bordoni 1994 Distribution in Iran: Isfahan. Iranian record: Sakenin et al. [2008c]. Leptobium convexicolle Assing 2009 Distribution in Iran: Mazandaran. Iranian record: Assing [2009g]. Endemic.
Leptobium gracile (Gravenhorst 1802)
Distribution in Iran: Ardabil, Fars, Golestan, Guilan,
Khuzestan.
Iranian record: Scheerpeltz [1961a], Coiffait [1982], Smetana [2004], Sakenin et al. [2008c, d], Assing [2005b, 2009a].
Leptobium illyricum (Erichson 1840)
Distribution in Iran: East Azarbaijan.
Iranian record: Smetana [2004], Assing [2005b, 2009a].
Subtribe Lathrobiina Laporte 1835 Achenium debile Erichson 1840 Distribution in Iran: Hormozgan.
Iranian record: Jarrige [1971], Coiffait [1982], Smetana [2004].
Achenium iranicum Coiffait 1982
Distribution in Iran: Sistan & Baluchestan.
Iranian record: Coiffait [1982], Smetana [2004]. Endemic.
Achenium khouzestanicum Bohac 1981 Distribution in Iran: Khuzestan.
Iranian record: Bohac [1981], Coiffait [1982], Smetana [2004]. Endemic.
Achenium meridionale Coiffait 1982 Distribution in Iran: Kerman.
Iranian record: Coiffait [1982], Smetana [2004]. Endemic. Achenium mesopotamicum Coiffait 1971 Distribution in Iran: Not cited. Iranian record: Coiffait [1971a, 1982], Smetana [2004].
Acheniumpersicum Coiffait 1972 Distribution in Iran: Kerman.
Iranian record: Coiffait [1972b, 1982], Smetana [2004]. Endemic.
Acheniumpicinum Fauvel 1875
Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1961c].
Achenium planum Erichson 1840
Distribution in Iran: Not cited.
Iranian record: Coiffait [1982], Smetana [2004].
Achenium quadriceps Eppelsheim 1889
Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1961b, 1963a].
Achenium richteri Scheerpeltz 1961
Distribution in Iran: Fars, Hormozgan, Kerman, Khuzestan.
Iranian record: Scheerpeltz [1961a], Jarrige [1971],
Coiffait [1982], Smetana [2004]. Endemic.
Achenium scimbaloides Koch 1937
Distribution in Iran: East Azarbaijan, Guilan.
Iranian record: Sakenin et al. [2008c, d].
Domene stilicina (Erichson 1840)
Distribution in Iran: Chahar-Mahal Bakhtiari, Fars,
Isfahan, Khorasan, Mazandaran.
Iranian record: Assing [2007e], Sakenin et al. [2008c, d],
Anlaç & Newton [2010].
Lathrobium caspicum Koch 1938
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Koch [1938], Smetana [2004], Assing
[2008f, 2009i].
Lathrobium castaneipenne Kolenati 1846
Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Lathrobium fulvipenne (Gravenhorst 1806) Distribution in Iran: Ardabil.
Iranian record: Horion [1965], Coiffait [1982], Smetana
[2004], Assing [2007e].
Lathrobium furcatum Czwalina 1888
Distribution in Iran: East Azarbaijan.
Iranian record: Horion [1965], Coiffait [1982], Smetana
[2004], Ghahari et al. [2009b], Assing [2009i].
Lathrobium impressifrons Eppelsheim 1884
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Assing [2007e, 2009h].
Lathrobium kuntzeni Koch 1939
Distribution in Iran: Not cited.
Iranian record: Koch [1939], Smetana [2004], Assing [2008f].
Lathrobium serriae Assing 2007
Distribution in Iran: Khorasan.
Iranian record: Assing [2007e]. Endemic.
Lobrathium farsicum Assing 2007
Distribution in Iran: Fars, Hormozgan.
Iranian record: Assing [2007f]. Endemic.
Lobrathium rugipenne (Hochhuth 1851)
Distribution in Iran: East Azarbaijan, Khuzestan.
Iranian record: Sakenin et al. [2008c, d], Ghahari et al. [2009b].
Micrillus pallidus (Reitter 1887)
Distribution in Iran: Chahar-Mahal Bakhtiary.
Iranian record: Sakenin et al. [2008d].
Micrillus testaceus (Erichson 1840)
Distribution in Iran: Mazandaran.
Iranian record: Anlaç & Newton [2010].
Platydomene stoeckleini (Koch 1937)
Distribution in Iran: Fars.
Iranian record: Assing [2003, 2007e].
Pseudobium anatolicum Assing 2006
Distribution in Iran: Fars.
Iranian record: Assing [2007c].
Pseudobium cypriacum Jarrige 1949
Distribution in Iran: Chahar-Mahal Bakhtiari, East
Azerbaijan, Fars.
Iranian record: Assing [2007c].
Pseudobium iranicum Bohac 1988
Distribution in Iran: Fars, Khorasan, Tehran.
Iranian record: Bohac [1988], Smetana [2004] Assing
[2007c, e]. Endemic.
Pseudobium richteri (Scheerpeltz 1961)
Distribution in Iran: Hormozgan, Sistan & Baluchestan.
Iranian record: Scheerpeltz [1961a], Coiffait [1984],
Smetana [2004] Assing [2007c].
Pseudolathra tennenbaumi (Bernhauer 1932)
Distribution in Iran: Guilan, Hamedan.
Iranian record: Sakenin et al. [2008c, d].
Scymbalium anale (Nordmann 1837)
Distribution in Iran: East Azarbaijan, Mazandaran,
Semnan.
Iranian record: Sakenin et al. [2008c, d], Anlaç & Newton
[2010].
Scymbalium minimum Eppelsheim 1888 Distribution in Iran: East Azarbaijan. Iranian record: Ghahari et al. [2009b]. Tetartopeus persicus Coiffait 1972 Distribution in Iran: Not cited.
Iranian record: Coiffait [1972b, 1982], Bordoni [1982], Smetana [2004], Assing [2008f, Assing, 2010e], Anlaç [2009a].
Tetartopeus stylifer (Reitter 1909)
Distribution in Iran: East Azerbaijan, Fars, Guilan, Isfahan, Kerman, Mazandaran, Zanjan.
Iranian record: Assing [2004c, 2007e, 2009i, Assing, 2010e]. Throbalium iranicum Scheerpeltz 1961 Distribution in Iran: Hormozgan, Khuzestan. Iranian record: Scheerpeltz [1961a], Jarrige [1971], Bordoni [1982], Coiffait [1982], Smetana [2004]. Endemic.
Subtribe Medonina Casey 1905 Hypomedon debilicornis (Wollaston 1857) Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Lithocharis fontinalis (Peyerimhoff 1929) Distribution in Iran: Hormozgan. Iranian record: Jarrige [1971]. Lithocharis mateui Coiffait 1968 Distribution in Iran: Hormozgan. Iranian record: Assing [2007e]. Lithocharis ochracea (Gravenhorst 1802) Distribution in Iran: Tehran. Iranian record: Bohac [1981]. Luzea graeca (Kraatz, 1857)
Distribution in Iran: Fars, Isfahan, Kordestan, West Azarbaijan.
Iranian record: Assing [2010a].
Luzea infirma (Erichson 1840)
Distribution in Iran: Semnan.
Iranian record: Scheerpeltz [1960a], Assing [2010a].
Luzea nigritula (Erichson 1840)
Distribution in Iran: Not cited.
Iranian record: Coiffait [1984], Smetana [2004].
Luzea shavrini Assing 2010 Distribution in Iran: Lorestan. Iranian record: Assing [2010a]. Medon frischi Assing 2005 Distribution in Iran: Chahar-Mahal Bakhtiari. Iranian record: Assing [2005a, 2007d]. Endemic. Medon fusculoides Coiffait 1970 Distribution in Iran: Chahar-Mahal Bakhtiari, East Azerbaijan, Fars, Isfahan, Kerman, Kermanshah, Kordestan, Lorestan, West Azarbaijan, Yazd. Iranian record: Assing [2004b, 2005a, 2006a, 2007d, 2007g, 2009h], Assing [2010c]. Medon fusculus (Mannerheim 1830) Distribution in Iran: Golestan, Tehran, West Azarbaijan. Iranian record: Sakenin et al. [2008c], Anlaç & Newton [2010].
Medon lydicus Bordoni 1980
Distribution in Iran: Guilan, Lorestan.
Iranian record: Sakenin et al. [2008c, d].
Medon maronitus (Saulcy 1865)
Distribution in Iran: East Azerbaijan, Golestan, Guilan.
Iranian record: Assing [2004b, 2007d, 2009h].
Medon paradisiacus Assing 2004
Distribution in Iran: East Azerbaijan, Tehran.
Iranian record: Assing [2007d].
Medon pythonissa (Saulcy 1865)
Distribution in Iran: East Azarbaijan, Isfahan.
Iranian record: Sakenin et al. [2008c, d].
Medon semiobscurus (Fauvel 1875)
Distribution in Iran: East Azarbaijan, Guilan, Khorasan.
Iranian record: Assing [2006a], Sakenin et al. [2008c, d],
Ghahari et al. [2009b]. Endemic.
Medon sparsiventris Eppelsheim 1889
Distribution in Iran: Golestan, Guilan, Mazandaran, West
Azarbaijan.
Iranian record: Bernhauer [1923], Scheerpeltz [1958a],
Coiffait [1970, 1976], Schülke [1989a], Smetana [2004],
Assing [2004b, 2005a, 2009h], Assing [2010c].
Pseudomedon dido (Saulcy 1865)
Distribution in Iran: Fars.
Iranian record: Assing [2009e].
Pseudomedon obscurellus (Erichson 1840)
Distribution in Iran: Golestan, Mazandaran, Semnan,
Tehran
Iranian record: Assing [2009e]. Pseudomedon obsoletus (Nordmann 1837) Distribution in Iran: Golestan, Mazandaran. Iranian record: Smetana [2004], Sakenin et al. [2008c, d]. Sunius adanensis (Lokay 1919)
Distribution in Iran: Kerman, Kordestan, Kuhkuloyeh &
Boyer Ahmad, Lorestan, Semnan.
Iranian record: Assing [2006b, 2010d], Sakenin et al. [2010].
Sunius fulgocephalus (Coiffait 1970)
Distribution in Iran: East Azerbaijan, Guilan, Golestan,
Khorasan, Kordestan, Mazandaran, Semnan, Tehran, West
Azarbayjan.
Iranian record: Assing [2008a, 2010d]. Sunius iranicus Assing 2002
Distribution in Iran: Fars, Isfahan, Kerman, Kuhkuloyeh & Boyer Ahmad.
Iranian record: Assing [2002, 2008a, 2010d]. Endemic.
Sunius khnzoriani (Coiffait 1970)
Distribution in Iran: Hamadan, Isfahan, Mazandaran, West
Azarbayjan.
Iranian record: Assing [2008a, 2010d]. Sunius melanocephalus (Fabricius 1793) Distribution in Iran: Kordestan, Mazandaran. Iranian record: Smetana [2004], Assing [2008a], Sakenin et al. [2008c, d].
Sunius picinus (Bernhauer 1902)
Distribution in Iran: North Iran [No locality cited), Isfahan. Iranian record: Scheerpeltz [1963a], Ghahari et al. [2009d]. Sunius praecisus Assing 2008 Distribution in Iran: Khorasan.
Iranian record: Assing [2008a, 2008d, 2010d].
Subtribe Paederina Fleming 1821 Paederidus albipilis (Solsky 1871) Distribution in Iran: North East Iran (no locality cited), Mazandaran.
Iranian record: Scheerpeltz [1960a, 1961c, 1963a], Ghahari et al. [2009a].
Paederidus rubrothoracicus (Goeze 1777) Distribution in Iran: Isfahan. Iranian record: Sakenin et al. [2008c, d]. Paederidus ruficollis (Fabricius 1777) Distribution in Iran: Isfahan.
Iranian record: Smetana [2004], Ghahari et al. [2009c].
Paederus balachowskyi (Jarrige 1971)
Distribution in Iran: Sistan & Baluchestan.
Iranian record: Jarrige [1971], Coiffait [1982], Smetana
[2004], Li & Zhou [2008].
Paederus balcanicus Koch 1938
Distribution in Iran: Mazandaran.
Iranian record: Tirgari & Nikbakhtzadeh [2002],
Nikbakhtzadeh & Tirgari [2008].
Paederus fuscipes fuscipes Curtis 1826
Distribution in Iran: Fars, Golestan, Guilan, Hormozgan,
Kerman, Khuzestan Mazandaran, Tehran, Zanjan.
Iranian record: Scheerpeltz [1961a], Jarrige [1971],
Bohac [1981], Nikbakhtzadeh & Sadeghiani [1999],
Nikbakhtzadeh [2002], Zargari et al. [2003], Smetana
[2004], Abbasipor and Taghavi [2005], Nikbakhtzadeh &
Tirgari [1999, 2008], Sakenin et al. [2008c, d], Ghahari et
al. [2009c], Anlaç & Newton [2010].
Paederus limnophilus Erichson 1840
Distribution in Iran: Ardabil.
Iranian record: Nikbakhtzadeh & Tirgari [2008].
Paederus littoralis ilsae Bernhauer 1932
Distribution in Iran: Ardabil, Fars, Guilan, Kerman,
Kordestan, Mazandaran.
Iranian record: Scheerpeltz [1957, 1961a, 1963a], Jarrige [1971], Coiffait [1982], Nikbakhtzadeh & Sadeghiani [1999], Tirgari & Nikbakhtzadeh [2002], Zargari et al. [2003], Smetana [2004], Abbasipor and Taghavi [2005], Nikbakhtzadeh & Tirgari [1999, 2008], Ghahari et al. [2009c]. Paederus littoralis littoralis Gravenhorst 1802 Distribution in Iran: Ardabil.
Iranian record: Zargari et al. [2003], Nikbakhtzadeh & Tirgari [2008].
Paederus mesopotamicus Eppelsheim 1889 Distribution in Iran: Kordestan. Iranian record: Smetana [2004], Li & Zhou [2008]. Paederus nigricornis Bernhauer 1911 Distribution in Iran: Sistan & Baluchestan. Iranian record: Nikbakhtzadeh & Tirgari [2008].
Paederus riparius (Linnaeus 1758)
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Jarrige [1971], Zargari et al. [2003],
Smetana [2004], Abbasipor and Taghavi [2005].
Paederus tamulus Erichson 1840
Distribution in Iran: Sistan & Baluchestan.
Iranian record: Scheerpeltz [1961a], Smetana [2004], Li & Zhou [2008].
Subtribe Scopaeina Mulsant & Rey 1878 Scopaeus alaniensis Coiffait 1969 Distribution in Iran: Mazandaran, West Azarbaijan. Iranian record: Sakenin et al. [2008c, d]. Scopaeus asiaticus Bernhauer 1915 Distribution in Iran: Mazandaran. Iranian record: Scheerpeltz [1963a], Ghahari et al., 2009a. Scopaeus azerbaidzhanus Gusarov 1994 Distribution in Iran: Mazandaran, Tehran. Iranian record: Gusarov [1994], Frisch et al. [2002], Frisch [2003, 2006a], Smetana [2004]. Scopaeus bicolor Baudi di Selve 1848 Distribution in Iran: Guilan, Mazandaran. Iranian record: Sakenin et al. [2008c, d]. Scopaeus cameroni Coiffait 1968
Distribution in Iran: Chahar-Mahal Bakhtiary, Mazandaran. Iranian record: Sakenin et al. [2008c, d]. Scopaeus chalcodactylus (Kolenati 1846) Distribution in Iran: Ardabil, East Azerbaijan, Isfehan, Golestan, Guilan, Hamadan, Khorasan, Mazandaran, Semnan, Tehran.
Iranian record: Frisch [2006a, 2007a]. Scopaeus debilis Hochhuth 1851
Distribution in Iran: Chahar-Mahal Bakhtiari, East Azerbaijan, Fars, Guilan, Hamadan, Khorasan, Lorestan, Tehran, Zanjan.
Iranian record: Frisch [1999a, 2006a], Frisch et al. [2002],
Smetana [2004], Sakenin et al. [2008c, d].
Scopaeus ebneri Scheerpeltz 1929
Distribution in Iran: Chahar-Mahal Bakhtiari, Fars,
Hamadan, Isfehan, Kermanshah, Kuhkuloyeh & Boyer
Ahmad, Lorestan.
Iranian record: Frisch et al. [2002], Frisch [2003, 2006a],
Smetana [2004].
Scopaeus farsensis Frisch 2007
Distribution in Iran: Fars.
Iranian record: Frisch [2007a]. Endemic.
Scopaeus femursetosus Frisch 2006
Distribution in Iran: Chahar-Mahal Bakhtiari, Fars, Isfahan,
Hamadan, Kuhkuloyeh & Boyer Ahmad, Lorestan.
Iranian record: Frisch [2006a]. Endemic.
Scopaeus filiformis Wollaston 1867
Distribution in Iran: Fars, Hormozgan, Kerman, Khuzestan,
Sistan & Baluchestan.
Iranian record: Scheerpeltz [1961a], Jarrige [1971], Coiffait [1984], Frisch [1999a, 2006a], Frisch et al. [2002]. Scopaeus gracilis (Sperk 1835)
Distribution in Iran: Chahar-Mahal Bakhtiari, East Azerbaijan, Fars, Golestan, Guilan, Isfahan, Kerman, Kermanshah, Khorasan, Kuhkuloyeh & Boyer Ahmad, Mazandaran, Tehran.
Iranian record: Scheerpeltz [1970], Frisch [1998, 2006a, 2007b], Frisch et al. [2002], Smetana [2004], Sakenin et al. [2008c, d].
Scopaeus hyrcanus Frisch 2006
Distribution in Iran: Guilan, Golestan, Mazandaran. Iranian record: Frisch [2006b]. Endemic.
Scopaeus iranensis Frisch 2006
Distribution in Iran: Kermanshah, Lorestan.
Iranian record: Frisch [2006a]. Endemic.
Scopaeus khnzoriani Coiffait 1968
Distribution in Iran: Ardabil, Chahar-Mahal Bakhtiari,
East Azerbaijan, Fars, Isfahan, Kermanshah, Lorestan.
Iranian record: Frisch [2006a, 2009].
Scopaeus kurdistanicoides Frisch 2002 Distribution in Iran: Tehran. Iranian record: Coiffait [1984], Frisch [2006a]. Scopaeus laevigatus (Gyllenhal 1827) Distribution in Iran: East Azarbaijan, Golestan. Iranian record: Sakenin et al. [2008c, d]. Scopaeus minutoides Coiffait 1969 Distribution in Iran: West Azarbaijan. Iranian record: Sakenin et al. [2008c]. Scopaeus pakistanensis Frisch 2007 Distribution in Iran: Sistan & Baluchestan. Iranian record: Frisch [2007b]. Scopaeuspersicus Frisch 1994
Distribution in Iran: Chahar-Mahal Bakhtiari, Fars, Ghazvin, Hamadan, Isfahan, Kermanshah, Kuhkuloyeh & Boyer Ahmad, Lorestan.
Iranian record: Frisch [1994, 1999b, 2006a], Frisch et al.
[2002], Smetana [2004]. Endemic.
Scopaeus qohrudensis Frisch 2008
Distribution in Iran: Isfahan, Yazd.
Iranian record: Frisch [2008]. Endemic.
Scopaeus schillhammeri Frisch 1994
Distribution in Iran: Tehran.
Iranian record: Frisch [1999b, 2006a], Frisch et al. [2002],
Smetana [2004].
Scopaeus serriae Frisch 2006
Distribution in Iran: Lorestan.
Iranian record: Frisch [2006a]. Endemic.
Scopaeus signifer Fauvel 1899
Distribution in Iran: Lorestan.
Iranian record: Frisch [1997], Frisch et al. [2002], Smetana [2004].
Scopaeus stramineus Frisch 2007
Distribution in Iran: Fars, Hormozgan.
Iranian record: Frisch [2007b]. Endemic.
Scopaeus zagrosensis Frisch 2006
Distribution in Iran: Chahar-Mahal Bakhtiari, Fars,
Kuhkuloyeh & Boyer Ahmad, Lorestan.
Iranian record: Frisch [2006a]. Endemic.
Subtribe Stilicina Casey 1905 Rugilus angustatus (Geoffroy 1785) Distribution in Iran: Mazandaran, Semnan. Iranian record: Sakenin et al. [2008c, d]. Rugilus iranicus (Coiffait 1981) Distribution in Iran: Mazandaran. Iranian record: Coiffait [1981b, 1984], Smetana [2004]. Endemic.
Rugilus korbi (Fauvel 1900)
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Bohac [1980], Rougemont [1988], Smetana
[2004].
Rugilus longicollis (Fauvel 1900) Distribution in Iran: North Iran (no locality cited). Iranian record: Fauvel [1900], Coiffait [1984], Smetana [2004].
Rugilus orbiculatus (Paykull 1789)
Distribution in Iran: Mazandaran, Semnan.
Iranian record: Smetana [2004], Sakenin et al. [2008c, d].
Rugilus rossii (Zanetti 1977)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Rugilus rufipes Germar 1836
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Tribe Pinophilini Nordmann 1837 Subtribe Pinophilina Nordmann 1837 Mimopinophilus siculus (Kraatz 1857) Distribution in Iran: Khorasan.
Iranian record: Bernhauer [1935], Jarrige [1971], Smetana [2004].
Mimopinophilus zarudnyi (Semenov 1909)
Distribution in Iran: Northeast Iran (no locality cited),
Khuzestan, Chahar-Mahal Bakhtiari.
Iranian record: Semenov [1909], Coiffait [1978], Assing
[2004a], Smetana [2004].
Pinophilus rufipennis Sharp 1874
Distribution in Iran: Kerman, Khuzestan, Sistan &
Baluchestan.
Iranian record: Scheerpeltz [1961a], Smetana [2004].
Subtribe Procirrina Bernhauer & Schubert 1912 Oedichirus reitteri Bernhauer 1908 Distribution in Iran: Not cited. Iranian record: Scheerpeltz [1961b].
16. Subfamily Staphylininae Latreille 1802 Tribe Othiini Thomson 1859 Atrecus affinis (Paykull 1789) Distribution in Iran: Mazandaran. Iranian record: Ghahari et al. [2009c]. Othius laeviusculus Stephens 1833 Distribution in Iran: Fars, Guilan, Mazandaran. Iranian record: Fauvel [1873], Assing [1999], Herman [2001], Smetana [2004], Anlaç & Newton [2010].
Tribe Platyprosopini Lynch Arribalzaga 1884 Platyprosopus bagdadensis Stierlin 1867 Distribution in Iran: Khuzestan.
Iranian record: Scheerpeltz [1961a], Herman [2001], Smetana [2004].
Platyprosopus elongates Mannerheim 1830 Distribution in Iran: Northeast Iran (no locality cited). Iranian record: Scheerpeltz [1961b, 1961c, 1963a].
Platyprosopus hierochonticus Reiche & Saulcy 1856 Distribution in Iran: East Azarbaijan. Iranian record: Ghahari et al. [2009b].
Tribe Staphylinini Latreille 1802
Subtribe Philonthina Kirby 1837 Bisnius cephalotes (Gravenhorst 1802) Distribution in Iran: Not cited. Iranian record: Smetana [2004]. Bisnius fimetarius (Gravenhorst 1802) Distribution in Iran: North Iran (no locality cited). Iranian record: Herman [2001], Smetana [2004]. Bisnius piochardi (Fauvel 1875) Distribution in Iran: Not cited.
Iranian record: Coiffait [1974], Herman [2001], Smetana [2004].
Bisnius reitteri (Eppleisheim 1889)
Distribution in Iran: Ghazvin.
Iranian record: Ghahari et al. [2009c].
Bisnius schillhammeri Hromadka 2001
Distribution in Iran: Chahar-Mahal Bakhtiari.
Iranian record: Hromadka [2001], Smetana [2004].
Endemic.
Bisnius sordidus (Gravenhorst 1802) Distribution in Iran: South Iran (no locality cited). Iranian record: Fauvel [1874], Gridelli [1924a], Herman [2001], Smetana [2004].
Erichsonius rivularis rivularis (Kiesenwetter 1858) Distribution in Iran: Not cited.
Iranian record: Coiffait [1974], Herman [2001], Smetana [2004].
Erichsonius subopacus (Hochhuth 1851)
Distribution in Iran: Guilan.
Iranian record: Bohac [1980].
Gabrius appendiculatus Sharp 1910
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Gabrius astutoides (Strand 1946)
Distribution in Iran: Golestan.
Iranian record: Sakenin et al. [2008d].
Gabrius lividipes (Baudidi Selve 1848)
Distribution in Iran: Guilan.
Iranian record: Ghahari et al. [2009c].
Gabrius nigritulus (Gravenhorst 1802)
Distribution in Iran: Mazandaran.
Iranian record: Fauvel [1874], Gridelli [1924a], Herman
[2001], Smetana [2004], Anlaç & Newton [2010].
Gabrius subnigritulus Joy 1913
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Gabrius tornus Joy 1913
Distribution in Iran: Not cited.
Iranian record: Smetana [2004], Assing [2009d].
Gabrius toxotes Joy 1913
Distribution in Iran: Mazandaran.
Iranian record: Ghahari et al. [2009c].
Gabronthus maritimus (Motschulsky 1858)
Distribution in Iran: Mazandaran, South Iran (no locality
cited).
Iranian record: Scheerpeltz [1963b], Anlaç & Newton [2010]. Gabronthus thermarum (Aube 1850) Distribution in Iran: East Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Neobisnus lathrobioides (Baudi di selve 1848)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Neobisnius procerulus procerulus (Gravenhorst 1806)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Neobisnius prolixus (Erichson 1840)
Distribution in Iran: Mazandaran.
Iranian record: Fauvel [1874], Herman [2001] Smetana
[2004], Anlaç & Newton [2010].
Philonthus aeneipennis Boheman 1858
Distribution in Iran: Not cited.
Iranian record: Scheerpeltz [1960b], Herman [2001], Smetana [2004].
Philonthus atratus (Gravenhorst 1802) Distribution in Iran: North Iran (no locality cited). Iranian record: Fauvel [1874], Gridelli [1922a], Herman [2001], Smetana [2004]. Philonthus biskrensis Fagel 1957
Distribution in Iran: Khuzestan, Northeast Iran (no locality cited).
Iranian record: Scheerpeltz [1961a], Gusarov [1995], Coiffait [1967, 1974, 1976], Herman [2001], Smetana [2004].
Philonthus broumandi Bohac 1981 Distribution in Iran: Khuzestan.
Iranian record: Bohac [1981], Herman [2001] Smetana [2004]. Endemic.
Philonthus carbonarius (Gravenhorst 1802) Distribution in Iran: Not cited.
Iranian record: Ménétriés [1832], Fauvel [1874], Herman [2001], Smetana [2004]. Philonthus caucasicus Nordmann 1837 Distribution in Iran: Kordestan.
Iranian record: Gridelli [1922a], Jarrige [1971], Coiffait
[1974], Herman [2001], Smetana [2004].
Philonthus cognatus Stephens 1832
Distribution in Iran: Fars, Mazandaran.
Iranian record: Jarrige [1971], Herman [2001], Smetana
[2004].
Philonthus concinnus (Gravenhorst 1802) Distribution in Iran: Mazandaran, North and South Iran (no locality cited).
Iranian record: Gridelli [1922a], Jarrige [1971], Herman
[2001], Smetana [2004].
Philonthus cruentatus (Gmelin 1790)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Philonthus debilis (Gravenhorst 1802)
Distribution in Iran: North Iran (no locality cited).
Iranian record: Fauvel [1874], Gridelli [1922a], Herman
[2001], Smetana [2004].
Philonthus dimidiatipennis Erichson 1840
Distribution in Iran: Guilan, Khuzestan, North Iran (no
locality cited), Mazandaran.
Iranian record: Gridelli [1922a], Scheerpeltz [1961a,
1961b, 1963a], Herman [2001], Smetana [2004], Sakenin et al. [2008c], Ghahari et al. [2009d]. Philonthus discoideus (Gravenhorst 1802) Distribution in Iran: Sistan & Baluchestan. Iranian record: Scheerpeltz [1961a], Herman [2001], Smetana [2004].
Philonthus ebeninus ebeninus (Gravenhorst 1802)
Distribution in Iran: Golestan, South Iran (no locality cited).
Iranian record: Gridelli [1922a], Jarrige [1971], Herman
[2001], Smetana [2004].
Philonthus infirmus Erichson 1840
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Philonthus intermedius (Lacordaire 1835)
Distribution in Iran: Guilan, Mazandaran, South Iran (no
locality cited).
Iranian record: Fauvel [1874], Gridelli [1922a], Herman [2001], Smetana [2004], Ghahari et al. [2009c], Anlaç & Newton [2010].
Philonthus irakoiranicus Scheerpeltz 1961 Distribution in Iran: Khuzestan.
Iranian record: Scheerpeltz [1961a], Coiffait [1974], Bohac
[1981], Herman [2001], Smetana [2004].
Philonthus khouzestanicus Bohac 1981
Distribution in Iran: Hormozgan, Khuzestan, Sistan &
Baluchestan.
Iranian record: Bohac [1981], Smetana [2004]. Endemic.
Philonthus laminatus (Creutzer 1799)
Distribution in Iran: Mazandaran.
Iranian record: Faldermann [1835], Gemminger & Harold
[1868], Jarrige [1971], Herman [2001], Smetana [2004].
Philonthus minutus Boheman 1848
Distribution in Iran: East Azarbaijan.
Iranian record: Ghahri et al. [2009b].
Philonthus nitidicollis (Lacordaire 1835)
Distribution in Iran: Mazandaran, Tehran.
Iranian record: Gridelli [1924a], Coiffait [1974], Herman
[2001], Smetana [2004], Anlaç & Newton [2010].
Philonthus parvicornis (Gravenhorst 1802)
Distribution in Iran: Mazandaran.
Iranian record: Jarrige [1971], Herman [2001], Smetana
[2004].
Philonthus picimanus (Ménétriés 1832)
Distribution in Iran: Mazandaran.
Iranian record: Jarrige [1971], Herman [2001], Smetana
[2004].
Philonthuspolitus (Linnaeus 1758) Distribution in Iran: Northwest Iran (no locality cited). Iranian record: Ménétriés [1832], Gridelli [1922a], Herman [2001], Smetana [2004].
Philonthus punctus punctus (Gravenhorst 1802) Distribution in Iran: Golestan, Northwest Iran (no locality cited).
Iranian record: Ménétriés [1832], Faldermann [1835], Herman [2001], Smetana [2004], Ghahari et al. [2009c]. Philonthus punctus rapillyi Jarrige 1971 Distribution in Iran: Fars.
Iranian record: Jarrige [1971], Herman [2001], Smetana [2004]. Endemic.
Philonthus quisquiliarius quisquiliarius (Gyllenhal 1810) Distribution in Iran: Not cited. Iranian record: Herman [2001].
Philonthus rotundicollis (Ménétriés 1832) Distribution in Iran: East Azarbaijan, Mazandaran. Iranian record: Faldermann [1835], Fauvel [1874], Herman [2001], Smetana [2004], Sakenin et al. [2008c, d], Ghahari et al. [2009a].
Philonthus rubripennis Stephens 1832
Distribution in Iran: Not cited.
Iranian record: Gridelli [1924a], Smetana [2004].
Philonthus rufipes (Stephens 1832) Distribution in Iran: Ghazvin. Iranian record: Ghahari et al. [2009c]. Philonthus salinus Kiesenwetter 1844 Distribution in Iran: Guilan.
Iranian record: Jarrige [1971], Herman [2001], Smetana [2004].
Philonthus schaeuffelei Scheerpeltz 1961 Distribution in Iran: Khuzestan.
Iranian record: Scheerpeltz [1961a], Coiffait [1974], Smetana [2004].
Philonthus splendens splendens (Fabricius 1793) Distribution in Iran: East Azarbaijan, West Azarbaijan. Iranian record: Smetana [2004], Ghahari et al. [2009b]. Philonthus succicola Thomson 1860 Distribution in Iran: Golestan.
Iranian record: Jarrige [1971], Herman [2001], Smetana [2004].
Philonthus tenuicornis Mulsant & Rey 1853
Distribution in Iran: Guilan.
Iranian record: Jarrige [1971], Herman [2001].
Philonthus turbidus Erichson 1840
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Philonthus varians (Paykull 1789)
Distribution in Iran: Mazandaran, Tehran.
Iranian record: Gridelli [1922a], Jarrige [1971], Smetana
[2004].
Philonthus velatipennis Solsky 1870
Distribution in Iran: Golestan, Hormozgan, Mazandaran,
Sistan & Baluchestan.
Iranian record: Jarrige [1971], Coiffait [1974], Smetana
[2004], Ghahari et al. [2009c].
Philonthus ventralis (Gravenhorst 1802)
Distribution in Iran: Not cited.
Iranian record: Herman [2001], Smetana [2004].
Rabigus abauriae (Gridelli 1924)
Distribution in Iran: North Iran (no locality cited).
Iranian record: Gridelli [1924a], Coiffait [1974], Herman
[2001], Smetana [1965, 2004].
Rabigus formosus (Motschulsky 1860)
Distribution in Iran: North Iran (no locality cited).
Iranian record: Scheerpeltz [1963a].
Rabigus tenuis (Fabricius 1793)
Distribution in Iran: Tehran.
Iranian record: Gridelli [1924a], Herman [2001], Smetana [2004].
Subtribe Quediina Kraatz 1857 Heterothops dissimilis (Gravenhorst 1802) Distribution in Iran: Sistan & Baluchestan. Iranian record: Scheerpeltz [1961a], Herman [2001], Stourac [2002], Smetana [2004]. Heterothops gestroi Gridelli 1922 Distribution in Iran: North Iran (no locality cited). Iranian record: Gridelli [1922b, 1924a], Coiffait [1978], Herman [2001], Smetana [2004]. Heterothops iranicus Stourac 2000 Distribution in Iran: Sistan & Baluchestan. Iranian record: Stourac [2000], Herman [2001], Smetana [2004]. Endemic.
Heterothops nigerrimus Bernhauer 1914
Distribution in Iran: Chahar-Mahal Bakhtiari, North Iran
(no locality cited).
Iranian record: Scheerpeltz [1963a], Sakenin et al. [2008c]. Heterothopspraevius Erichson 1839 Distribution in Iran: Not cited.
Iranian record: Fauvel [1874], Herman [2001], Smetana [2004].
Quedius acuminatus acuminatus Hochhuth 1849 Distribution in Iran: North Iran (no locality cited). Iranian record: Fauvel [1874], Gridelli [1925], Smetana [1962, 2004], Horion [1965], Herman [2001]. Quedius amplissimus Bernhauer 1912 Distribution in Iran: Kordestan.
Iranian record: Jarrige [1971], Herman [2001], Smetana [2004].
Quedius atricapillus Reitter 1900
Distribution in Iran: Ardabil.
Iranian record: Ghahari et al. [2009b].
Quedius bucharensis Bernhauer 1918
Distribution in Iran: Khorasan.
Iranian record: Bernhauer [1918], Gridelli [1924b],
Herman [2001], Smetana [2004].
Quedius cohaesus Eppelsheim 1888
Distribution in Iran: North Iran (no locality cited).
Iranian record: Coiffait [1977b, 1978], Herman [2001],
Solodovnikov [2004].
Quedius coloratus Fauvel 1875
Distribution in Iran: Isfahan.
Iranian record: Ghahari et al. [2009c].
Quedius gridellii Scheerpeltz 1933
Distribution in Iran: East Azarbaijan, North and South Iran
(no locality cited).
Iranian record: Gridelli [1922c, 1925], Coiffait [1978], Herman [2001], Smetana [1971, 2004]. Quedius hypocrita Korge 1971 Distribution in Iran: Guilan.
Iranian record: Korge [1971], Coiffait [1978], Herman [2001], Smetana [2004]. Endemic. Quedius iranicus Coiffait 1976 Distribution in Iran: Fars.
Iranian record: Coiffait [1976, 1978], Bordoni & Oromi
[1998], Herman [2001], Smetana [2004]. Endemic. Quedius kermanensis Coiffait 1973 Distribution in Iran: Kerman.
Iranian record: Coiffait [1973, 1978], Herman [2001], Smetana [2004]. Endemic. Quedius levicollis (Brullé 1832) Distribution in Iran: Not cited.
Iranian record: Horion [1965], Faldermann [1835], Herman [2001].
Quedius limbatus (Heer 1839) Distribution in Iran: Guilan.
Iranian record: Coiffait [1977b, 1978], Herman [2001],
Solodovnikov [2002], Smetana [2004].
Quedius lindbergi Scheerpeltz 1958
Distribution in Iran: Mazandaran.
Iranian record: Sakenin et al. [2008c].
Quedius lucidulus Erichson 1839
Distribution in Iran: West Azarbaijan.
Iranian record: Ghahri et al. [2009b].
Quedius macchabaeus Saulcy 1865 Distribution in Iran: South Iran (no locality cited). Iranian record: Fagel [1968b], Herman [2001], Smetana [2004].
Quedius microcapillatus Korge 1971 Distribution in Iran: East Azerbaijan. Iranian record: Korge [1971], Smetana [2004]. Quedius mixtus Eppelsheim 1890
Distribution in Iran: Guilan, Mazandaran, East Azarbaijan.
Iranian record: Korge [1971], Herman [2001], Smetana [2004].
Quedius nemoralis nemoralis Baudi di selve 1848
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Quedius nitipennis (Stephens 1833)
Distribution in Iran: Chahar-Mahal Bakhtiari.
Iranian record: Ghahari et al. [2009c].
Quedius nivicola Kiesenwetter 1858
Distribution in Iran: Not cited.
Iranian record: Fagel [1968a], Coiffait [1978], Herman
[2001], Smetana [2004].
Quedius obliqueseriatus Eppelsheim 1889
Distribution in Iran: Guilan.
Iranian record: Korge [1971], Herman [2001].
Quedius ochripennis (Ménétriés 1832)
Distribution in Iran: Northwest Iran (no locality cited),
East Azarbaijan, Zanjan.
Iranian record: Ménétriés [1832], Faldermann [1835],
Scheerpeltz [1958a, 1960a, 1961d], Herman [2001],
Sakenin et al. [2008c], Ghahari et al. [2009b, d].
Quedius orientalis Korge 1971
Distribution in Iran: Kermanshah.
Iranian record: Korge [1971], Coiffait [1978], Herman
[2001], Smetana [2004]. Endemic.
Quedius persicus Korge 1971
Distribution in Iran: Mazandaran.
Iranian record: Korge [1971], Coiffait [1978], Herman [2001].
Quedius riparius Kellner 1843
Distribution in Iran: East Azarbaijan.
Iranian record: Korge [1971], Herman [2001], Smetana [2004]. Quedius scintillans (Gravenhorst 1806) Distribution in Iran: Not cited.
Iranian record: Fauvel [1874], Gridelli [1925], Horion [1965], Coiffait [1978], Herman [2001], Smetana [2004]. Quedius semiobscurus (Marsham 1802) Distribution in Iran: Guilan.
Iranian record: Horion [1965], Korge [1971], Herman
[2001], Smetana [2004].
Quedius semirufus Korge 1971
Distribution in Iran: East Azarbaijan.
Iranian record: Korge [1971], Coiffait [1978], Herman
[2001], Solodovnikov [2005].
Quedius suramensis Eppelsheim 1880
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Quedius vicinus (Ménétriés 1832)
Distribution in Iran: Guilan, Kermanshah, Mazandaran,
Northwest Iran (no locality cited).
Iranian record: Ménétriés [1832], Faldermann [1835],
Jarrige [1971], Korge [1971], Coiffait [1978], Bohac
[1988], Herman [2001], Smetana [2004], Ghahari et al.
[2009c].
Subtribe Staphylinina Latreille 1802 Creophilus maxillosus maxillosus (Linnaeus 1758) Distribution in Iran: Mazandaran, Northwest Iran (no locality cited).
Iranian record: Ménétriés [1832], Fauvel [1874], Herman [2001], Smetana [2004], Ghahari et al. [2009d]. Dinothenarus flavocephalus adonis (Coiffait 1956) Distribution in Iran: Fars.
Iranian record: Jarrige [1971], Coiffait [1974], Smetana [2004]. Dinothenarus sibiricus (Gebler 1830) Distribution in Iran: Mazandaran, West Azarbaijan. Iranian record: Scheerpeltz [1963a, 1966], Herman [2001], Smetana [2004], Sakenin et al. [2008c], Ghahari et al. [2009].
Emus hirtus (Linnaeus 1758) Distribution in Iran: Tehran.
Iranian record: Jarrige [1971], Herman [2001], Smetana [2004].
Nudobius lentus (Gravenhorst 1806)
Distribution in Iran: East Azarbaijan, West Azarbaijan.
Iranian record: Ghahari et al. [2009b].
Ocypus curtipennis Motschulsky 1849
Distribution in Iran: Not cited.
Iranian record: Coiffait [1964, 1971c, 1974], Herman
[2001], Smetana [2004].
Ocypus fulvipennis Erichson 1840
Distribution in Iran: Semnan.
Iranian record: Sakenin et al. [2010].
Ocypus helleni (G. Müller 1926)
Distribution in Iran: East Azerbaijan, Khorasan.
Iranian record: G. Müller [1943], Scheerpeltz [1963a],
Smetana [1971, 2004], Herman [2001].
Ocypus matilei Jarrige 1971
Distribution in Iran: East Azarbaijan, Mazandaran.
Iranian record: Jarrige [1971], Coiffait [1974], Herman [2001], Smetana [1971, 2004], Smetana & Davies [2000]. Endemic.
Ocypus milleri robustus Coiffait 1964 Distribution in Iran: Not cited.
Iranian record: Coiffait [1964, 1974], Herman [2001], Smetana & Davies [2000], Smetana [2004]. Ocypus mus (Brullé 1832)
Distribution in Iran: Guilan, Mazandaran, North Iran (no locality cited).
Iranian record: Herman [2001], Smetana [1971, 2004],
Anlaç & Newton [2010].
Ocypus nitens nitens (Schrank 1781)
Distribution in Iran: Guilan.
Iranian record: Coiffait [1974], Herman [2001], Smetana [1971, 2004], Jarrige [1971]. Ocypus picipennis caucasicus (G. Müller 1926) Distribution in Iran: East Azarbaijan, Guilan, Kordestan, Mazandaran, Tehran.
Iranian record: Jarrige [1971], Smetana [1971, 2004], Herman [2001].
Ocypus picipennis picipennis (Fabricius 1793)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Ocypus quadraticeps (Ménétriés 1832)
Distribution in Iran: Northwest Iran (no locality cited).
Iranian record: Ménétriés [1832], Faldermann [1835],
Herman [2001], Smetana [2004].
Ocypus sericeicollis (Ménétriés 1832)
Distribution in Iran: Mazandaran.
Iranian record: Faldermann [1835], Herman [2001],
Smetana [2004], Anlaç & Newton [2010].
Ocypus syriacus Baudi di Selve 1848
Distribution in Iran: Semnan.
Iranian record: Sakenin et al. [2010].
Ontholestus callistus (Hochhuth 1849)
Distribution in Iran: Golestan, Guilan.
Iranian record: Smetana [1959], Coiffait [1974], Bohac
[1980], Herman [2001], Smetana [2004].
Ontholestus murinus (Linnaeus 1758)
Distribution in Iran: Fars, Mazandaran.
Iranian record: Jarrige [1971], Herman [2001], Smetana [2004].
Physetops giganteus Semenov 1906
Distribution in Iran: Golestan.
Iranian record: Sakenin et al. [2008c].
Physetops tataricus (Pallas 1773)
Distribution in Iran: East Azarbaijan, Golestan, Kerman,
Khorasan, Sistan & Baluchestan.
Iranian record: Scheerpeltz [1963a], Coiffait [1974], Herman [2001], Smetana [2004], Solodovnikov & Grebennikov [2005].
Platydracus flavopunctatus (Latreille 1804) Distribution in Iran: Mazandaran. Iranian record: Jarrige [1971], Herman [2001], Smetana [1971, 2004].
Platydracus stercorarius stercorarius (Olivier 1795) Distribution in Iran: East Azarbaijan, Guilan, Mazandaran,
Semnan.
Iranian record: Jarrige [1971], Herman [2001], Smetana [1971, 2004], Sakenin et al. [2010]. Staphylinus caesareus caesareus Cederhjelm 1798 Distribution in Iran: East Azarbaijan, Golestan, Guilan, Mazandaran
Iranian record: Fauvel [1874], Jarrige [1971], Herman
[2001], Smetana [1971, 2004], Anlaç & Newton [2010].
Staphylinus erythropterus erythropterus Linnaeus 1758
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Tasgius ater (Gravenhorst 1802)
Distribution in Iran: Not cited.
Iranian record: Faldermann [1835], Herman [2001], Smetana [2004].
Tasgius doriae (G. Müller 1932) Distribution in Iran: North Iran (no locality cited). Iranian record: G. Müller [1932], Coiffait [1964, 1974], Herman [2001], Smetana [2004]. Endemic. Tasgius gracilicornis (Hochhuth 1849) Distribution in Iran: East Azarbaijan, Mazandaran. Iranian record: G. Müller [1932], Herman [2001], Smetana [1971, 2004].
Tasgius pedator pedator (Gravenhorst 1802)
Distribution in Iran: Northwest Iran (no locality cited).
Iranian record: Ménétriés [1832], Faldermann [1835],
Herman [2001], Smetana [2004].
Tasgius pliginskii (Bernhauer 1915)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Tasgiusprotensus (Ménétriés 1832)
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Tasgius rubripennis (Reiche & Saulcy 1856)
Distribution in Iran: Not cited.
Iranian record: Coiffait [1974], Herman [2001], Smetana [2004].
Tasgius winkleri (Bernhauer 1906) Distribution in Iran: Semnan. Iranian record: Sakenin et al. [2010].
Tribe Xantholinini Erichson 1839 Calontholinus fasciatus (Hochhuth 1849) Distribution in Iran: Guilan.
Iranian record: Jarrige [1971], Bordoni [1975], Herman
[2001], Smetana [2004].
Gauropterus bucharicus Bernhauer 1905
Distribution in Iran: Fars.
Iranian record: Assing [2007b].
Gauropterus fulgidus (Fabricius 1787)
Distribution in Iran: Fars.
Iranian record: Herman [2001], Smetana [2004], Assing [2007b].
Gauropterus punctulatus (Paykull 1789) Distribution in Iran: East Azarbaijan. Iranian record: Ghahari et al. [2009b]. Gauropterus sanguinipennis (Kolenati 1846) Distribution in Iran: Fars, Guilan.
Iranian record: Jarrige [1971], Coiffait [1972a], Herman [2001], Smetana [2004], Bordoni [1978, 2005], Assing [2007b].
Gauropterus sanguinipes (Reitter 1889)
Distribution in Iran: Mazandaran.
Iranian record: Jarrige [1971], Herman [2001], Smetana
[2004], Bordoni [2005], Assing [2007b, 2009d].
Gauropterus semenovi Kirshenblat 1951
Distribution in Iran: Kerman.
Kirshenblat [1951], Coiffait [1972a], Herman [2001],
Smetana [2004], Bordoni [2005]. Endemic.
Gyrohypnus angustatus Stephens 1833
Distribution in Iran: Ardabil, East Azerbaijan, Guilan.
Iranian record: Assing [2007b].
Gyrohypnuspunctulatus (Paykull 1789)
Distribution in Iran: East Azarbaijan.
Iranian record: Fauvel [1873], Herman [2001], Smetana
[2004].
Hypnogyra laevissimus (Reitter 1898)
Distribution in Iran: Mazandaran.
Iranian record: Reitter [1898], Bordoni [1975, 1999a,
2008], Herman [2001], Smetana [2004].
Leptacinus batychrus (Gyllenhal 1827)
Distribution in Iran: Ardabil, East Azerbaijan, Tehran.
Iranian record: Fauvel [1873], Herman [2001], Smetana
[2004], Assing [2007b].
Leptacinus klapperichi Coiffait 1981
Distribution in Iran: Tehran.
Iranian record: Coiffait [1981a], Herman [2001], Smetana [2004]. Endemic.
Stenistoderus cephalotes cephalotes (Kraatz 1858)
Distribution in Iran: Fars, Khuzestan.
Iranian record: Scheerpeltz [1961a], Herman [2001],
Smetana [2004], Assing [2007b].
Xantholinus araxis Reitter 1898
Distribution in Iran: Fars, Kermanshah, Kordestan.
Iranian record: Jarrige [1971], Bordoni [1975], Herman
[2001], Smetana [2004].
Xantholinus audrasi Coiffait 1956
Distribution in Iran: Ardabil, Guilan, Mazandaran, Tehran.
Iranian record: Bordoni [1973], Korge [1973], Herman
[2001], Smetana [2004], Assing [2007b].
Xantholinus azarbaijanicus Korge 1973
Distribution in Iran: East Azarbaijan.
Iranian record: Korge [1973], Bordoni [1999a], Herman
[2001], Smetana [2004].
Xantholinus crassicornis Hochhuth 1851
Distribution in Iran: East Azarbaijan, Mazandaran,
Semnan.
Iranian record: Coiffait [1966], Bordoni [1983, 1985], Herman [2001], Smetana [2004], Assing [2007b], Sakenin et al. [2010].
Xantholinus fortepunctatus Motschulsky 1860 Distribution in Iran: Golestan, Guilan, Khorasan, Mazandaran.
Iranian record: Jarrige [1971], Korge [1973], Bordoni [1975, 1999a], Herman [2001], Smetana [2004].
Xantholinus linearis linearis (Olivier 1795) Distribution in Iran: Not cited.
Iranian record: Fauvel [1873], Herman [2001], Smetana [2004].
Xantholinus longiventris Heer 1839
Distribution in Iran: Not cited.
Iranian record: Smetana [2004].
Xantholinus reitteri Coiffait 1966
Distribution in Iran: Guilan, Mazandaran.
Iranian record: Coiffait [1973], Bordoni [1999a], Herman
[2001], Smetana [2004], Assing [2007b].
Xantholinus rufipennis Erichson 1839
Distribution in Iran: Mazandaran.
Iranian record: Anlaç & Newton [2010].
Zeteotomus brevicornis (Erichson 1839)
Distribution in Iran: Mazandaran.
Iranian record: Ghahari et al. [2009a].
Zeteotomus iranicus (Coiffait 1976)
Distribution in Iran: Golestan.
Iranian record: Coiffait [1976], Herman [2001], Smetana [2004]. Endemic.
DISCUSSION
The fauna of Iranian rove beetles is studied in this paper. The result indicates that the fauna of Iranian Staphylinidae is very diverse and several new records are added to the list annually. Iran is a large country and with attention to its various geographical climates, we estimate that 1500 staphylinid species can be present in this country.
ACKNOWLEDGEMENT
The authors are indebted to Dr. A. Newton and Dr. H. Frank of USA and V. Assing of Germany for sending the necessary papers. The research was supported by Islamic Azad University (Science and Research Branch of Tehran).
REFERENCES
Abbasipor H., Taghavi A., 2005. Study of rove beetles, Paederus spp. (Col.: Staphylinidae), the agents of skin dermatitis in north of Iran// Proceedings of the Fifth International Conference on Urban Pests. 487 p. Anlas S., 2009a. On the genus Tetartopeus Czwalina 1888 of Turkey. A new species and additional records (Coleoptera, Staphylinidae, Paederinae)// Zootaxa 2031. P. 63-67. Anlas S., 2009b. Distributional checklist of the Staphlinidae (Coleoptera) of Turkey, with new and additional records// Linzer biologische Beiträge 41 (1). P. 215-342. Anlas S., Newton A.F., 2010. Distributional checklist of the Staphylinidae (Coleoptera) of Iran, with new and additional records// Linzer biol. Beitr. 42/1. P. 335-388 Assing V., 1999. A revision of Othius Stephens (Coleoptera, Staphylinidae). VIII. Further records, new species, and a new synonym// Linzer biologische Beiträge 31 (2). P. 661-691.
Assing V., 2002. New species of Sunius Curtis from China and Iran (Coleoptera: Staphylinidae, Paederinae)// Linzer biologische Beiträge 34 (1). P. 289-296. Assing V., 2003. Two new species of Platydomene Ganglbauer from Romania and Turkey, with a redescription of P stoeckleini (Koch), (Coleoptera:
Staphylinidae, Paederinae)// Linzer biologische Beiträge 35 (1). P. 5-12.
Assing V., 2004a. New species and records of Staphylinidae from Turkey II (Insecta: Coleoptera: Staphylinidae)// Beiträge zur Entomologie 54 (1). P. 53-73.
Assing V., 2004b. A revision of the Medon species of the Eastern Mediterranean and adjacent regions (Insecta: Coleoptera: Staphylinidae: Paederinae)// Bonner Zoologische Beiträge 52. P. 33-82.
Assing V., 2004c. New species and records of Staphylinidae from Turkey III (Insecta: Coleoptera)// Linzer biologische Beiträge 36 (2). P. 669-733.
Assing V., 2005a. A revision of Western Palaearctic Medon. IV. A new species and new records from Iran, and a catalogue of species (Coleoptera, Staphylinidae, Paederinae)// Linzer biologische Beiträge 37 (2). P. 1027-1034.
Assing V, 2005b. A revision of the genus Leptobium Casey (Coleoptera: Staphylinidae: Paederinae)// Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 673. P. 1-182.
Assing V., 2005c. On the western Palaearctic species of Drusilla Leach, with special reference to the species of the eastern Mediterranean (Coleoptera: Staphylinidae, Aleocharinae)// Koleopterologische Rundschau 75. P. 111-149.
Assing V, 2005d. A revision of the species of Geostiba Thomson and Tropimenelytron Pace of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae, Aleocharinae)// Linzer biologische Beiträge 37 (2). P. 903-1006.
Assing V., 2005e. On the taxonomy and biogeography of Stenus (s.str.) erythrocnemus Eppelsheim and related species (Insecta: Coleoptera: Staphylinidae)// Bonner Zoologische Beiträge 53 (2004). P. 303-310.
Assing V., 2006a. A revision of Western Palaearctic Medon: the species of the Atlantic Islands, the Western Mediterranean, and Europe, except for the southeast (Insecta: Coleoptera: Staphylinidae: Paederinae)// Bonner Zoologische Beiträge 54 (2005). P. 25-95.
Assing V, 2006b. On the Turkish species of Sunius. VII. Five new micropterous species from western Anatolia and additional records (Coleoptera: Staphylinidae, Paederinae)// Beiträge zur Entomologie 56 (2). P. 297-315.
Assing V, 2007a. New species and additional records of Staphylinidae from Turkey V (Coleoptera)// Stuttgarter Beiträge zur Naturkunde Serie A (Biologie) 700. P. 1-64.
Assing V, 2007b. On the Xantholinini of Turkey and adjacent regions (Coleoptera: Staphylinidae, Staphylininae)// Zootaxa 1474. P. 1-54.
Assing V., 2007c. On the genus Pseudobium Mulsant & Rey II. A new species from Pakistan and additional records (Coleoptera: Staphylinidae: Aleocharinae)// Linzer biologische Beiträge 39 (1). P. 15-21.
Assing V., 2007d. A revision of Western Palaearctic Medon. V. A new species from Iran and additional records (Coleoptera: Staphylinidae, Paederinae)// Linzer biologische Beiträge 39 (1). P. 23-32.
Assing V., 2007e. New species and additional records of Paederinae and Aleoharinae from Iran (Coleoptera, Staphylinidae)// Deutsche Entomologische Zeitschrift
54 (2). P. 179-193.
Assing V, 2007f. A revision ofPalaearctic Lobrathium Mulsant & Rey. III. New species, new synonyms, and additional records (Coleoptera: Staphylinidae, Paederinae)// Linzer biologische Beiträge 39 (2). P. 731-755.
Assing V., 2007g. A revision of Western Palaearctic Medon. VI. A new species from Turkey and additional records (Coleoptera: Staphylinidae, Paederinae)// Linzer biologische Beiträge 39 (2). P. 791-797.
Assing V, 2007h. A revision of the species of Pronomaea Erichson of the Western Palaearctic region, including Middle Asia (Coleoptera: Staphylinidae, Aleocharinae, Pronomaeini)// Beiträge zur Entomologie 57. P. 367-396.
Assing V., 2008a. A revision of the Sunius species of the Western Palaearctic region and Middle Asia (Coleoptera: Staphylinidae, Paederinae)// Linzer biologische Beiträge 40 (1). P. 5-135.
Assing V, 2008b. A revision of Othiini. XVI. Four new species of Othius from the Himalaya and China, and additional records (Coleoptera: Staphylinidae, Staphylininae)// Koleopterologische Rundschau 78. P. 245-263.
Assing V, 2008c. On the Cryptobiina of the Arabian peninsula (Coleoptera: Staphylinidae, Paderinae)// Zootaxa 1892. P. 53-64.
Assing V., 2008d. Four new species and additional records of Palaearctic Sunius, with two new synonymies (Coleoptera: Staphylinidae: Paederinae)// Beiträge zur Entomologie 58 (2). P. 455-470.
Assing V., 2008e. A revision of the Western Palaearctic and Middle Asian species of Drusilla Leach IV. A new species from Iran and additional records (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)// Entomologische Blätter für Biologie und Systematik der Käfer 103/104. P. 51-58.
Assing V., 2008f. On the taxonomy and zoogeography of some Palaearctic Paederinae and Xantholinini (Coleoptera: Staphylinidae)// Linzer biologische Beiträge 40 (2). P. 1237-1294.
Assing V., 2008g. New species and additional records of Geostiba from the Eastern Mediterranean region and from Middle Asia (Coleoptera: Staphylinidae: Aleocharinae)// Linzer biologische Beiträge 40 (2). P. 1327-1335.
Assing V., 2009a. A revision of Leptobium Casey IV. Three new species and additional records (Coleoptera: Staphylinidae: Paederinae)// Stuttgarter Beiträge zur Naturkunde Serie A, Neue Serie 2. P. 227-236.
Assing V., 2009b. On the taxonomy and zoogeography of some Palaearctic Aleochara species of the subgenera Xenochara Mulsant & Rey and Rheochara Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae)// Beiträge zur Entomologie 59 (1). P. 33-101.
Assing V, 2009c. On the Western Palaearctic and Middle Asian species of Ochthephilum Stephens, with notes on Cryptobium koltzei Eppelsheim (Coleoptera: Staphylinidae: Paederinae: Cryptobiina)// Linzer biologische Beiträge 41 (1). P. 397-426.
Assing V., 2009d. On the Staphylinidae of Turkey. VI. Thirteen new species and additional records (Coleoptera)// Koleopterologische Rundschau 79. P. 117-172.
Assing V., 2009e. On the Pseudomedon species of
the Palaearctic region (Coleoptera: Staphylinidae: Paederinae)// Linzer biologische Beiträge 41 (2). P. 1175-1189.
Assing V., 2009f. A revision of Geostiba of the Western Palaearctic region. XIX. New species from Turkey and Iran and additional records, with an updated key and catalogue of the species of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae: Aleocharinae)// Linzer biologische Beiträge 41 (2). P. 1191-1246.
Assing V, 2009g. A revision of Leptobium CASEY III. A new species from Iran and additional records (Coleoptera: Staphylinidae: Paederinae)// Linzer biologische Beiträge 41 (2). P. 1247-1252.
Assing V., 2009h. A revision of Western Palaearctic Medon. VII. A new species from southern Turkey and additional records (Coleoptera: Staphylinidae: Paederinae)// Linzer biologische Beiträge 41 (2). P. 1253-1268.
Assing V., 2009i. New species and additional records of Lathrobium and Tetartopeus from the Palaearctic region (Coleoptera: Staphylinidae: Paederinae)// Linzer biologische Beiträge 41 (2). P. 1269-1283.
Assing V., 2009j. New species, new synonymies, and additional records of Leptusa from Turkey and Iran (Coleoptera: Staphylinidae: Aleocharinae)// Linzer biologische Beiträge 41 (2). P. 1285-1305.
Assing V, 2009k. On some Oxypoda species of the subgenus Sphenoma Mannerheim (Coleoptera, Staphylinidae, Aleocharinae)// Linzer biologische Beiträge 41 (2). P. 1307-1315.
Assing V., 2009l. A revision of Ceranota Stephens, subgenus of Aleochara Gravenhorst (Coleoptera: Staphylinidae: Aleocharinae)// Beiträge zur Entomologie 59 (2). P. 355-422.
Assing V., 2010a. A revision of the genus Luzea (Coleoptera, Staphylinidae, Paederinae)// Dtsch. Entomol. Z. 57 (1). P. 117-135.
Assing V., 2010b. The second species of Dictyon Fauvel from the Western Palaearctic region, with additional records of D. pumilio (Eppelsheim) (Coleoptera: Staphylinidae: Aleocharinae)// Linzer biol. Beitr. 42/1. P. 485-488.
Assing V., 2010c. A revision of Palaearctic Medon. VIII. A new species from Nepal and additional records (Coleoptera: Staphylinidae: Paederinae)// Linzer biol. Beitr. 42/1. P. 489-498.
Assing V., 2010d. A revision of Palaearctic Sunius. XII. New species, new synonymies, and additional records (Coleoptera: Staphylinidae: Paederinae)// Linzer biol. Beitr. 42/2. P. 1045-1061.
Assing V., 2010e. Two new species of Tetartopeus from Turkey (Coleoptera: Staphylinidae: Paederinae)// Linzer biol. Beitr. 42/2. P. 1063-1071.
Assing V, 2010f. A revision of Amarochara of the Holarctic region. IV. Three new species from China, a new synonymy, additional records, and an updated key to species (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini)// Linzer biol. Beitr. 42/2. P. 1139-1154.
Assing V, Wunderle P., 1995. A revision of the species of the subfamily Habrocerinae (Coleoptera: Staphylinidae) of the world// Revue Suisse de Zoologie 102 (2). P. 307-359.
Assing V., Wunderle P., 1996. A revision of the species of the subfamily Habrocerinae of the world. Supplement I. (Coleoptera: Staphylinidae)// Beiträge zur Entomologie 46 (2). P. 373-378.
Atamehr A., Kamali K., Ostovan H., 2004. Fauna of aquatic beetles (Coleoptera) in Tabriz region// Proceedings of 16th Iranian Plant Protection Congress, Tabriz University, p. 123.
Benick G. 1941. Neue paläarktische Arten der Untergattung Aloconota Thomson. (Gattung Atheta, Col. Staph.)// Mitteilungen der Münchner Entomologischen Gesellschaft 31 (1). P. 184-196.
Bernhauer M., 1918. Neue Quedius Arten der palaarktischen Fauna// Verhandlungen der KaiserlichKöniglichen Zoologisch-Botanischen Gesellschaft in Wien 68. P. 92-96.
Bernhauer M., 1923. Neue Staphyliniden der palaearktischen Fauna// Koleopterologische Rundschau 10 1922. P. 122-128.
Bernhauer M., 1935. Neuheiten der paläarktischen Staphylinidenfauna. I// Koleopterologische Rundschau 21. P. 39-48.
Bernhauer M., 1936. Neuheiten der paläarktischen Staphylinidenfauna III// Koleopterologische Rundschau 22. P. 50-58.
Bernhauer M., 1940. Neuheiten der paläarktischen Staphylinidenfauna (Col., Staph.)// Itteilungen der Münchner Entomologischen Gesellschaft 30 (2). P. 622-642.
Bernhauer M., Schubert K. 1910, 1911, 1912, 1914, 1916. Staphylinidae I-II-III-VV. In: Schenkling, S. (ed.), Coleopterorum Catalogus. Pars 19, 29, 40, 57, 67. Junk Publ. P. 1-498.
Bernhauer M., Scheerpeltz O., 1926. Staphylinidae VI. In: Schenkling S. (ed.), Coleopterorum Catalogus. Pars 82. Junk Publ. P. 499-988.
Besuchet C., 1961. Deux psélaphides nouveaux d'Iran (Coleoptera)// Stuttgarter Beiträge zur Naturkunde 51. P. 1-3.
Besuchet C., 1969. Psélaphides paléarctiques. Espéces nouvelles et notes synonymiques. III (Coleoptera)// Revue Suisse de Zoologie 76 (1). P. 397-420.
Besuchet C., 1975a. Bibloplectus nouveaux de la région méditerranéenne orientale (Coleoptera, Pselaphidae)// Mitteilungen der Schweizerischen Entomologischen Gesellschaft 48. P. 31-36.
Besuchet C., 1975b. Revision du genre Scotoplectus Reitt. (Col., Pselaphidae)// Mitteilungen der Schweizerischen Entomologischen Gesellschaft 48. P. 397-404.
Besuchet C., 1980. Le genre Panaphantus Kiesw. (Col., Pselaphinae)// Mitteilungen der Schweizerischen Entomologischen Gesellschaft 53. P. 151-155.
Besuchet C., 1981a. Contribution à l'étude des Batrisodes paléarctiques (Coleoptera, Pselaphidae)// Revue Suisse de Zoologie 88 (1). P. 275-296.
Besuchet C., 1981b. Insects of Saudi Arabia. Coleoptera: Fam. Pselaphidae// Fauna of Saudi Arabia 3. P. 243-250.
Besuchet C., 1986. Synonymes et homonyme nouveaux de quelques genres de Psélaphides (Coleoptera)// Revue Suisse de Zoologie 93 (1). P. 257-264.
Besuchet C., 1999. Psélaphides paléarctiques.
Notes taxonomiques et faunistiques (Coleoptera, Staphylinidae, Pselaphinae)// Revue Suisse de Zoologie 106 (1). P. 45-67.
Besuchet C., Sabella G., 1999a. Nuove specie di Tychus della regione paleartica (Coleoptera, Staphylinidae, Pselaphinae)// Mitteilungen der Schweizerischen Entomologischen Gesellschaft 72. P. 235-258.
Besuchet C., Sabella G., 1999b. Descrizione di nuove specie di Tychus della regione paleartica con revisione dei Tychus del gruppo armeniacus (Coleoptera: Staphylinidae: Pselaphinae)// Annales de la Société Entomologique de France (N.S.) 35. P. 303-318.
Bodemeyer B. von, 1927. Ueber meine Entomologische Reisen nach Kleinasien (1911), Ost-Sibirien, Schilka und Amur (1912), Tunis, Oasis Gafsa, Khroumerie (1913) und Iran, das Elbursgebirge (1914). Bd. IV// Iran, das Elbursgebirge. Stuttgart: Alfred Kerner, 96 pp., 1 pl.
Bohac J., 1980. New or interesting finds of Staphylinidae from palearctic region (Coleoptera, Staphylinidae)// Acta Faunistica Entomologica Musei Nationalis Pragae 16. P. 85-87.
Bohac J., 1981. Results of the Czechoslovak-Iranian entomological expeditions to Iran. Coleoptera, Staphylinidae. Three new species of staphylinids from Iran. Acta Entomologica Musei Nationalis Pragae 40. P. 355-358.
Bohac J. 1988. Descriptions of new species of the subfamily Paederinae (Coleoptera, Staphylinidae) from the Palaearctic region// Acta Entomologica Bohemoslovaca 85. P. 434-443.
Bordoni A., 1973. Studi sulla sistematica e la geonemia degli Xantholinus. VII. Nuove entità del Mediterraneo e della Persia del Museo Civico di Storia Naturale di Milano// Atti Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 114 (1). P. 71-80.
Bordoni A., 1975. Studi sulla sistematica e la geonemia degli Xantholinus. VIII. Le specie eurocentroasiatiche e caucasiche in particolare. Revisione di tipi e descrizione di nuove entità// Memorie della Società Entomologica Italiana 53 (1974). P. 56-96.
Bordoni A., 1978. Staphilinidae dell'Asia Minore. Quinta nota: Entita' raccolte in grotta e descrizione di nuove specie (Coleoptera)// Fauna Ipogea di Turchia, Quaderni di Speleologia, Circolo Speleologico Romano 3. P. 55-67.
Bordoni A., 1982. Studi sui Paederinae -V- Intorno ad alcuni Lathrobiini poco noti, revisione dei tipi e descrizione di una nuova specie italiana (Col. Staphylinidae)// Frustula Entomologica, Nuova Serie 2 (1979). P. 31-56.
Bordoni A., 1983. Studi sulla sistematica degli Xantholinus Berthold 1827, XIII. Una nuova specie dell'Iran (Insecta: Coleoptera: Staphylinidae)// Senckenbergiana Biologica 63 (1982). P. 61-63.
Bordoni A., 1985. Note sulla morfologia di alcuni Xantholinini europei (Col., Staphylinidae)// Frustula Entomologica, Nuova Serie 6 (1983). P. 81-88.
Bordoni A., 1999a. Note sulla sinonimia di alcune specie e sottogeneri del genere Xantholinus Dejean (sic), 1821 e designazione di lectotipi (Coleoptera: Staphylinidae)// Opuscula Zoologica Fluminensia 171. P. 1-8.
Bordoni A., 1999b. Una nuova specie di Deliphrosoma della Turchia (Coleoptera, Staphylinidae)// Fragmenta
Entomologica 31 (2). P. 195-203.
Bordoni A., 2005. Sui Gauropterus della Regione Paleartica e in particolare su quelli. descritti da Kirschenblatt, conservati nel Museo Zoologico di San Pietroburgo (Coleoptera, Staphylinidae)// Bollettino della Società Entomologica Italiana 137 (3). P. 205-213.
Bordoni A., 2008. Lepidophallus Coifaith is a synonym of Megalinus Mulsant & Rey (Coleoptera, Staphylinidae, Xantholinini)// Onychium 6. P. 54-59.
Bordoni A., Oromi P. 1998. Coleoptera Staphylinidae. pp. 1147-1162. In: Juberthie, C. & V. Decu, Encyclopaedia Biospeologica.Vol. 3. Bucarest: Société de Biospéologie.
Cameron M., 1940. New species of Oriental Staphylinidae (Coleoptera)// The Entomologist's Monthly Magazine 76. P. 249-253.
Castellini G., 1996. Revisione del genere Leptomastax Pirazzoli, 1855 (Coleoptera, Scydmaenidae)// Atti del Museo Civico di Storia Naturale (Grosseto) 15 (Supplement). P. 1-137.
Coifaith H., 1964. Note sur les Ocypus (sensu lato) avec description de formes nouvelles// Bulletin de la Société d'Histoire Naturelle de Toulouse 99. P. 81-106.
Coifaith H., 1966. Novye Xantholinini iz sovetskogo soiuza (Coleoptera, Staphylinidae)// Zoologicheskii Zhurnal 45. P. 195-202.
Coifaith H., 1967 Tableau de détermination des Philonthus de la région Paléarctique occidentale (Col., Staphylinidae)// Annales de la Société Entomologique de France (N.S.) 3 (2). P. 381-450.
Coifaith H., 1970. Staphylinides nouveaux ou mal connus de la région paléarctique occidentale// Bulletin de la Société d'Histoire Naturelle de Toulouse 106. P. 99-111.
Coifaith H., 1971a. Le genre Achenium (Staphylinidae)// Nouvelle Revue d'Entomologie 1(3). P. 11-42.
Coifaith H., 1971b. Nouveaux Astenus de la région paléarctique occidentale// Nouvelle Revue d'Entomologie 1 (2). P. 177-199.
Coifaith H., 1971c. Staphylinides nouveaux ou mal connus de Bulgarie// Nouvelle Revue d'Entomologie 3. P. 279-286.
Coifaith H., 1972a. Coléoptères Staphylinidae de la région paléarctique occidentale. I. Généralités. Sousfamilles: Xantholininae et Leptotyphlinae// Revue d'Entomologie, Supplément 2 (2). P. 1-651.
Coifaith H., 1972b. Paederinae nouveaux ou mal connus de la région paléarctique occidentale// Nouvelle Revue d'Entomologie 2 (2). P. 131-150.
Coifaith H., 1973. Staphylinides nouveaux ou mal connus d'Europe, d'Afrique du nord et du Moyen-Orient. Les sousgenres du genre Bledius// Nouvelle Revue d'Entomologie 3 (2). P. 107-123.
Coifaith H., 1974. Coléoptères Staphylinidae de la région paléarctique occidentale. II. Sous famille Staphilininae (sic) Tribus Philonthini et Staphylinini// Nouvelle Revue d'Entomologie, Supplément 4 (4). P. 1-593.
Coifaith H., 1976. Nouveaux Staphylinidae (Coléoptères) d'Iran et de Turquie// Nouvelle Revue d'Entomologie 6 (1). P. 61-69.
Coifaith H. 1977a. Staphylinides anophtalmes nouveaux du Muséum de Genéve// Revue Suisse de Zoologie 84 (3). P. 659-663.
Coifaith H., 1977b. Note sur quelques Quedius et Heterothops nouveaux ou mal connus (Col., Staphylinidae)// Nouvelle Revue d'Entomologie 7 (2). P. 133-143.
Coifaith H., 1978. Coléoptères staphylinides de la région paléarctique occidentale. III. Sous famille Staphylininae, Tribu Quediini. Sous famille Paederinae, Tribu Pinophilini// Nouvelle Revue d'Entomologie, Supplément 8 (4). P. 1-364.
Coifaith H., 1980. Staphylinides nouveaux de la région Méditerranéenne// Nouvelle Revue d'Entomologie 10 (3). P. 251-264.
Coifaith H., 1981a. Staphylinides nouveaux du Moyen Orient appartenant au Musée Hongrois d'Historie Naturelle (Coleoptera)// Folia Entomologica Hungarica 42 (1). P. 15-23.
Coifaith H., 1981b. Sur quelques staphylinides nouveaux ou mal connus// Nouvelle Revue d'Entomologie 11 (1). P. 51-57.
Coifaith H., 1982. Coléoptères Staphylinidae de la région paléarctique occidentale. IV. Sous famille Paederinae. Tribu Paederini 1 (Paederi, Lathrobii)// Nouvelle Revue d'Entomologie, Supplément 12 (4). P. 1-440.
Coifaith H., 1984. Coléoptères staphylinides de la région paléarctique occidentale V Sousfamille Paederinae, Tribu Paederini 2. Sous famille Euaesthetinae// Nouvelle Revue d'Entomologie, Supplément 13 (4). P. 1-424.
Davies A. 2004. Tribe Cyrtoscydmini (Scydmaenidae) L.W. Schaufuss, 1889. In: Löbl, I. & A. Smetana (eds), Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup. P. 206-223.
Eppelsheim E., 1889. Neue Staphylinen aus den Kaukasusländern, besonders aus Circassien// Wiener Entomologische Zeitung 8. P. 11-22.
Fagel G., 1968a. Contribution à la connaissance des Staphylinidae. CI. Remarques diverses sur quelques Quedius de Méditerranée orientale// Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 44 (12). P. 1-13.
Fagel G., 1968b. Contribution à la connaissance des Staphylinidae. CIII. Le complexe des Quedius oblitteratus ER. - nemoralis baudi// Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 44 (24). P. 1-14.
Fagel G., 1969. Contribution à la connaissance des Staphylinidae. CIX. Les Melagria Casey de la région paléarctique occidentale// Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 45 (34). P. 1-43.
Fagel G., 1970. Contribution a la connaissance des Staphylinidae. CXI. Sur quelques Bledius du sous-genre Elbidus Rey de la région paléarctique occidentale// Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 46 (21). P. 1-12.
Faldermann F., 1835. Additamenta entomologica ad faunam rossicam in itineribus jussu imperatoris augustissimi annis 1827-1831 a CI. Ménétriés et Szovitz susceptis collecta, in lucem edita// Nouvelle Mémories de la Société Impériale des Naturalistes de Moscou 4. P. 1-310.
Fauvel A., 1871. Faune Gallo-Rhénane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhénanes et la
Valais avec tableaux synoptiques et planches gravées. Bulletin de la Société Linnéenne de Normandie (2) 5: (1869-70). P. 27-192. (Note: The date for this article is usually cited as 1869 which is the year of the meeting at which the article was presented. The inside title page of the journal is dated 1870 but the outside one is 1871. The Zoological Record, 1872 indicates the article may have been published in 1872 (Herman 2001).
Fauvel A., 1872. Faune Gallo-Rhénane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la prusse Rhénane, la Nassau et la Valais avec tableaux synoptiques et planches gravées. Caen. 214 pp.
Fauvel A., 1873. Faune Gallo-Rhénane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhénanes et la Valais avec tableaux synoptiques et planches gravées. Bulletin de la Société Linnéenne de Normandie (2)
7. P. 8-131. (Note: This publication includes the 1er supplément (Herman (2001).
Fauvel A., 1874. Faune Gallo-Rhénane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhénanes et la Valais avec tableaux synoptiques et planches gravées. Bulletin de la Société Linnéenne de Normandie (2)
8. P. 167-340. (Note: This publication includes the 2e supplément (Hermann (2001).
Fauvel A., 1875. Faune Gallo-Rhénane Catalogue systématique des staphylinides de la Faune Gallo-Rhénane avec l'addition synonymique des espèces européennes, siberiénnes, caucasiques et Mediterranéennes et descriptions nouvelles. Tome 3. Livraison 6. Caen: Le Blanc-Hardel. P. 545-738.
Fauvel A., 1900. Staphylinides paléarctiques nouveaux// Revue d'Entomologie 19. P. 218-253.
Franz H., 1981. Eine neue Euconnus-Art (Col. Scydmaenidae) aus dem Iran// Entomologica Basiliensia 6. P. 317-318.
Frisch J., 1994. Neue Arten der Gattung Scopaeus Erichson aus Griechenland, Anatolien und dem Iran (Staphylinidae: Paederinae)// Coleoptera: Schwanfelder Coleopterologische Mitteilungen 2. P. 1-46.
Frisch J., 1997. A revision of some west Palaearctic species of Scopaeus Erichson (Staphylinidae: Paederinae)// Revue Suisse de Zoologie 104 (3). P. 523-557.
Frisch J., 1998. A revision of some west Palaearctic species of Scopaeus Erichson (Staphylinidae: Paederinae)// Revue Suisse de Zoologie 105 (1). P. 89-124.
Frisch J., 1999a. A revision of the Scopaeus debilis species group, with description of a new species from Madagascar (Staphylinidae: Paederinae)// Revue Suisse de Zoologie 106 (2). P. 361-383.
Frisch J., 1999b. Scopaeus korelli n.sp. From North Anatolia (Staphylinidae: Paederinae), with synonymical and distributional notes on the Scopaeus elegans species group// Entomologische Blätter für Biologie und Systematik der Käfer 95. P. 159-165.
Frisch J., 2003. A revision of the Scopaeus laevigatus species group, with descriptions of ten species from the east Palaearctic, the Oriental and the Australian regions (Coleoptera: Staphylinidae, Paederinae)// Memoirs on
Entomology, International 17. P. 650-725.
Frisch J., 2006a. The genus Scopaeus (Coleoptera: Staphylinidae, Paederinae) in Iran, with description of new species from Zagros Mountains// Deutsche Entomologische Zeitschrift 53. P. 5-22.
Frisch J., 2006b. Scopaeus hyrcanus sp.n. from Elburz and Talish Mountain in Iran (Coleoptera: Staphylinidae, Paederinae)// Deutsche Entomologische Zeitschrift 53. P. 264-270.
Frisch J., 2007a. Scopaeus farsensis sp.n. from South Iran, with remarks on S. chalcodactylus (Kolenati 1846) (Coleoptera, Staphylinidae, Paederinae)// Deutsche Entomologische Zeitschrift 54. P. 79-88.
Frisch J., 2007b. A review of the Scopaeus gracilis species group (Coleoptera, Staphylinidae, Paederinae), with description of new species from Sardinia, southern Africa and Middle East// Deutsche Entomologische Zeitschrift Z. 54. P. 195-218.
Frisch J., 2008. A new species of Scopaeus Erichson 1839 (Staphylinidae, Paederinae), from the Qohrud Mountains in Iran// Deutsche Entomologische Zeitschrift 55 (2). P. 285-289.
Frisch J., 2009. A revision of the Scopaeus mutatus species group (Staphylinidae, Paederinae), with description of new species from Anatolia and the Caucasus// Deutsche Entomologische Zeitschrift 56. P. 271-287.
Frisch J., Burchardt D., Wolters V., 2002. Rove beetles of the subtribe Scopaeina Mulsant & Rey (Coleoptera: Staphylinidae) in the West Palaearctic: phylogeny, biogeography and species catalogue. Appendix 2: Synonymic catalogue of West Palaearctic Scopaeina including biogeographic information// Organisms Diversity & Evolution 2, Electronic Supplement 2 (1). P. 1-16.
Gemminger M., Harold E., 1868. Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. 2. P. 425-752 + 6. Monachii: E.H. Gummi.
Ghahari H., Tabari M., Haji-Amiri M., Sakenin H., Ostovan H., 2009a. Population fluctuation of rice stem borer, Chilo suppressalis Walker (Lepidoptera: Pyralidae) in paddy fields of northern Amol in Mazandaran Province// Journal of Plant Protection 23 (1). P. 41-49 (In Persian with English Summary).
Ghahari H., Anlas S., Sakenin H., Ostovan H., M. Havaskary, 2009b. Biodiversity of rove beetles (Coleoptera: Staphylinoidea: Staphylinidae) from the Arasbaran biosphere reserve and vicinity, northwestern Iran// Linzer biologische Beiträge 41 (2). P. 1949-1958.
Ghahari H., Anlas S., Sakenin H., Ostovan H., Tabari M., 2009c. A contribution to the rove beetles (Coleoptera: Staphylinoidea: Staphylinidae) of Iranian rice fields and surrounding grasslands// Linzer biologische Beiträge 41 (2). P. 1959-1968.
Ghahari H., Havaskary M., Tabari M., Ostovan H., Sakenin H., Satar A., 2009d. An annotated catalogue of Orthoptera (Insecta) and their natural enemies from Iranian rice fields and surrounding grasslands// Linzer biol. Beitr. 41/1. P. 639-672.
Gildenkov M.Y., 2001a. The Palaearctic Carpelimus fauna (Coleoptera: Staphylinidae). The problems of species and the formation of species. The first part. The history
of studying. Morpho-ecological features. The system of genus. The description of species (in Russian, English abstract)// Izdatel'stvo SGPU (Publishing House SSPU), Smolensk. 304 p.
Gildenkov M.Y., 2001b. Phylogenetic relations in the Oxytelinae subfamily. The Palaearctic Thinodromus fauna (Coleoptera: Staphylinidae: Oxytelinae) (in Russian, English abstract)// Izdatel'stvo SGPU (Publishing House SSPU), Smolensk. 204 p.
Gildenkov M.Y., 2009a. Novye dannye o rasprostranenii v Palearktike vidov roda Thinodromus I vidov roda Carpelimus iz podrodov Carpelimus s.str., Paratrogophloeus, Bucephalinus (Coleoptera, Staphylinidae)// Izvestiia Smolenskogo
Gosudarstvennogo Universiteta 2 (6). P. 25-42.
Gildenkov M.Y., 2009b. Novye dannye o rasprostranenii v Palearktike vidov roda Carpelimus iz podrodov Myopinus, Trogophloeus, Troginus (Coleoptera, Staphylinidae)// Izvestiia Smolenskogo
Gosudarstvennogo Universiteta 3 (7). P. 237-255.
Grebenikov V.V., Newton A.F., 2009. Good-bye Scydmaenidae, or why the ant-like stone beetles should become megadiverse Staphylinidae sensu latissimo (Coleoptera)// European Journal of Entomology 106
(2). P. 275-301.
Gridelli E., 1922a. Settimo contributo allo studio degli Staphylini. Note su alcuni Philonthus della Persia e delle regioni adiacenti// Annali del Museo Civico di Storia Naturale «Giacoma Doria» (3) 10. P. 175-176.
Gridelli E., 1922b. Descrizioni di alcuni nuovi Staphylinidae paleartici// Bollettino della Società Entomologica Italiana 54. P. 21-28.
Gridelli E., 1922c. Studi sul genre Quedius Steph. (Coleoptera, Staphylinidae). Primo contributo al subgen. Sauridus REY e Raphirus Steph// Atti dell'Accademia Scientifica Veneto-Trentino-Istriana
(3) 12/13. P. 123-140.
Gridelli E., 1924a. Settimo contributo allo studio degli Staphylini. Note su alcuni Philonthus della Persia e delle regioni adiacenti (continuation of 1922a)// Annali del Museo Civico di Storia Naturale «Giacoma Doria» (3) 10. P. 177-181.
Gridelli E., 1924b. Studi sul genre Quedius Steph. (Coleoptera, Staphylinidae). Secondo contributo. Specie della regione paleartica// Memorie della Società Entomologica Italiana 3 (1). P. 5-112.
Gridelli E., 1925. Studi sul genre Quedius Steph. (Coleoptera, Staphylinidae). Secondo contributo. Specie della regione paleartica (continuation of 1924b)// Memorie della Società Entomologica Italiana 3 (2). P. 113-180.
Gusarov V, 1994. New and little known Palaearctic Paederinea (Coleoptera, Staphylinidae)// Annales de la Société Entomologique de France (N.S.) 30 (4). P. 431-446.
Gusarov V., 1995. Novye i maloizvestnye Palearkticheskie stafilinidy (Coleoptera: Staphylinidae). Soobshchenie 6// Entomologicheskoi Obozrenie 74 (1). P. 81-96.
Hangay G., Nadal L., Szekel K., 2005. Report on Hungarian entomological expeditions to Iran// Folia Historico Naturalia Musei Matraensis 29. P. 7-18.
Hematyian R., Vafaei Shushtari R., Ebrahimi E., 2010.
Fauna and distribution of aquatic Coleoptera in parts of centeral Alborz// Proceedings of 19th Iranian Plant Protection Congress, Iranian Research Institute of Plant Protection, Tehran, 31 July - 3 August 2010, p. 137.
Herman L.H., 1970. Phylogeny and reclassification of the genera of the rove-beetle subfamily Oxytelinae of the World (Coleoptera, Staphylinidae)// Bulletin of the American Museum of Natural History 142 (5). P. 343-454.
Herman L.H., 1983. Eppelsheimius: Revision, distribution, sister group relationship (Staphylinidae, Oxytelinae)// American Museum Novitates 2764. P. 1-18.
Herman L.H. 1986. Revision of Bledius. Part IV Classification of species group, phylogeny, natural history, and catalogue (Coleoptera, Staphylinidae, Oxytelinae)// Bulletin of the American Museum of Natural History 184. P. 1-368.
Herman L.H., 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Volumes I-VII// Bulletin of the American Museum of Natural History 265. P. 1-4218.
Hlavac P., Jaszay T., 2009. A revision of the genus Zyras (Zyras) Stephens, 1835 (Coleoptera, Staphylinidae, Aleocharinae). I. Current classification status and the redefinition of the genus// ZooKeys 29. P. 49-71.
Horion A. 1963. Faunistik der mitteleuropäischen Käfer. Staphylinidae. 1. Micropeplinae bis Euaesthetinae. 9. P. xii + 1-412. Überlingen-Bodensee: A. Feyel.
Horion A., 1965. Faunistik der mitteleuropäischen Käfer. Staphylinidae. 2. Paederinae bis Staphylininae. 10. P. xv + 1-335. Überlingen-Bodensee: A. Feyel.
Hromadka L., 1979. Zwei neue paläarktische Stenus-Arten (Coleoptera, Staphylinidae)// Reichenbachia 17 (22). P. 183-187.
Hromadka L., 2001. Zwei neue Staphyliniden-Arten aus Iran (Insecta: Coleoptera: Staphylinidae: Tachyporinae et Staphylininae)// Reichenbachia 34 (16). P. 137-141.
Jarrige J., 1971. Contribution a la faune de l' Iran 21. Coléoptéres brachelytra// Annales de la Société Entomologique de France (N.S.) 7 (2). P. 483-502.
Karaman Z., 1957. Die balkanischen Bythininen (Col. Pselaphidae). Ihre Systematic, Zoogeographie und Phylogenie// Bioloski Glasnik 10. P. 161-208.
Karaman Z., 1961. Neue Gruppierung der Arten des Genus Brachygluta Thompson (Coleoptera Pselaphidae)// Acta Musei Macedonici Scientarium Naturalium-Skopje 7. P. 127-174.
Kashcheev VA., 1988. A new genus and new species of the staphylinid tribe Oxytelini (Coleoptera, Staphylinidae) from the USSR and Iran// Entomologicheskoe Obozrenie 67 (4). P. 780-789 (in Russian, English translation in Entomological Review 68 (6): 44-53 (1989).
Kirshenblat Y.D., 1951. Novye Palearkticheskie Staphylinidae (Coleoptera)// Entomologicheskoe Obozrenie 31. P. 541-545.
Kistner D.H., 1982. The social insects' bestiary. In: Hermann, H.R. (ed.), Social Insects. Vol. 3. Academic Press, New York. P. 1-244.
Koch H., 1937. Beitrag zur Systematik und geograpischen Verbreitung der Achenium-Arten (Coleoptera, Stapylinidae)// Pubblicazioni del Museo Entomologico «Pietro Rossi» (Duino) 2. P. 51-187.
Koch H., 1938. Über neue und wenig bekannte paläarktische Paederinae// Entomologische Blätter 34 (3). P. 103-116.
Koch H., 1939. Über neue und wenig bekannte paläarktische Paederinae (Col., Staphylinidae). III. Entomologische Blätter 35 (3). P. 156-172.
Kolenati F.A., 1846. Meletemata entomologica. Fasc. III. Brachelytra caucasi cum distributione geographica adnexis pselaphinis, scydmaenis, notoxidibus et xylophagis. 44 pp., 3 pls. Peyropolis: Imperialis Academiae Scientiarum.
Korge H., 1971. Über einige Quedius-Arten aus dem Iran (Col. Staphylinidae)// Entomologische Blätter für Biologie und Systematik der Käfer 67 (1). P. 9-20.
Korge H., 1973. Studien über westpalaearktische Arten der Gattung Xantholinus Serville (Coleoptera: Staphylinidae)// Mitteilungen der Deutschen Entomologischen Gesellschaft 31 (1972). P. 4-11.
Lawrence J., Newton A., 1982. Evolution and Classification of Beetles// Annual Review of Ecology and Systematics 13. P. 261-290.
Li X.-Y., Zhou H.-Z., 2008. Taxonomy and biogeography of the subgenus Eopaederus of the genus Paederus Fabricius (Coleoptera: Staphylinidae: Paderinae)// Annales de la Société Entomologique de France (N.S.) 44 (3). P. 327-344.
Likovsky Z., 1981. Ergebnisse der tschechoslowakischiranischen entomologischen Expeditionen nach dem Iran. Coleoptera: Staphylinidae, Subtribus Aleocharinae// Acta Entomologica Musei Nationalis Pragae 40. P. 359-370.
Löbl I., 1970. Revision derpaläarktischenArten der Gattungen Scaphisoma Leach und Caryoscapha Ganglbauer der Tribus Scaphisomini (Col. Scaphidiidae)// Revue Suisse de Zoologie 77 (4). P. 727-799.
Löbl I., 2004. Subfamily Scaphidiinae Latreille, 1807. In: Löbl I. & A. Smetana (eds), Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea - Histeroidea -Staphylinoidea. Apollo Books, Stenstrup. P. 495-504.
Löbl I., Besuchet C., 2004. Subfamily Pselaphinae Latreille, 1802. In: Löbl I. & A. Smetana (eds), Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea -Histeroidea - Staphylinoidea. Apollo Books, Stenstrup. P. 272-329.
Löbl I., Smetana A. (eds.), 2004. Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea - Histeroidea -Staphylinoidea. Apollo Books, Stenstrup. 942 p.
Luze G., 1906. Revision der paläarktischen Arten der Staphyliniden-Genera: Xylodromus, Omalium, Phyllodrepa, Hypopycna, Dialycera, Pycnoglypta und Phloeonomus// Verhandlungen der KaiserlichKöniglichen Zoologisch-Botanischen Gesellschaft in Wien 56. P. 485-602.
Makranczy G., 2006. Systematics andphylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae)// Annales Historico-Naturales Musei Nationalis Hungarici 98. P. 29-120.
Maus C.H., 1998. Taxonomical contributions to the subgenus Coprochara Mulsant & Rey, 1874 of the genus Aleochara Gravenhorst, 1802 (Coleoptera: Staphylinidae)// Koleopterologische Rundschau 68. P. 81-100.
Meggiolaro G., 1961. Descrizione di un nuovo Pselaphidae della Persia// Bollettino del Museo Cvico di Storia Naturale di Venezia 14. P. 53-56.
Menetrises E., 1832. Catalogue raisonné des objects de zoologie recueillis dans un voyage au Caucase et jusqu'aux frontières actuelles de la Perse entrepris par ordre de S.M. L'Empereur. 271 + xxxii + iv pp. St. Petersbourg: L'Académie Impériale des Sciences.
Müller G. (also J.), 1932. Settimo contributo alla conoscenza del genre Staphylinus L.// Bollettino della Società Entomologica Italiana 64 (5). P. 75-88.
Müller G. (also J.), 1943. Ottavo contributo alla conoscenza del genere Staphylinus (1)// Atti del Museo Civico di Storia Naturale Trieste 15 (5). P. 95-109.
Newton A.F., 2007. Documenting biodiversity: how well are we doing in Staphyliniformia (Coleoptera)? Entomological Society of America poster presentation D0471 (available (ESA members only) at http://esa. confex.com/esa/2007/techprogram/paper_32168.htm).
Newton A.F., Chandler D.S., 1989. World catalog of the genera of Pselaphidae (Coleoptera)// Fieldiana Zoology (N.S.) 53. P. 1-93.
Newton A.F. and Thayer M.K. 1992. Current classification and family-group names in Staphyliniformia (Coleoptera)// Fieldiana: Zoology (N.S.), 67. P. 1-92.
Newton A.F., Thayer M.K., 2005. Catalog of higher taxa, genera and subgenera of Staphyliniformia. Field Museum of Natural History, Chicago (last updated 27 August 2005; available from URL: http://www. fieldmuseum.org/peet_staph/db_1a.html).
Nikbakhtzadeh M., Sadeghiani C., 1999. Dermatite due à deux espèces de Paederus au sud de l'Iran// Bulletin de la Société de Pathologie Exotique 92 (1). P. 56.
Nikbakhtzadeh M., Tirgari S., 1999. Two medically important beetles of Fars province, Iran: Paederus spp. (Col: Staphylinidae) & its linear dermatitis (in Persian)// Iranian Journal of Public Health 28 (1-4). P. 73-86.
Nikbakhtzadeh M., Tirgari S., 2008. Medically important beetles (Insecta: Coleoptera) of Iran// Journal of Venomous Animals and Toxins including Tropical Diseases 14 (4). P. 597-618.
Orousset J., 2008. Révision des espèces paléarctiques du genre Chevrolatia Jacquelin du Val, 1850 (Coleoptera, Scydmaenidae)// Bulletin de la Société Entomologique de France 113 (3). P. 313-329.
Pace R., 1983. Specie del genere Geostiba Thompson raccolte dal Dr. C. Besuchet e collaborati in Marocco, nella Penisola Iberica e Balcanica, e nel Medio Oriente (Coleoptera: Staphylinidae)// Revue Suisse de Zoologie 90 (1). P. 3-46.
Pace R., 1984. Nuove Aleocharinae microftalme mediterranee e dell'Iran, del Muséum d'Histoire naturelle di Ginevra (Coleoptera, Staphylinidae)// Archives des Sciences (Genève) 37. P. 211-219.
Pace R., 1989. Monografia del genere Leptusa Kraatz: Supplemento IV (Coleoptera, Staphylinidae)// Memorie del Museo Civico di Storia Naturale di Verona (IIa Serie), Sezione Scienze della Vita (A: Biologica) (2) 8. P. 1-307.
Pasnik G., 2006. A revision of the World species of the genus Tachyusa Erichson 1837 (Coleoptera, Staphylinidae:
Aleocharinae)// Zootaxa 1146. P. 1-152.
Puthz V., 1970. Stenus (Paratenus) heinzianus n.sp. aus dem Iran (Coleoptera, Staphylinidae)// Reichenbachia 13 (23). P. 223-226.
Puthz V., 1972a. Das subgenus «Hemistenus» (Coleoptera, Staphylinidae)// Annales Entomologici Fennici 38 (2). P. 75-92.
Puthz V., 1972b. Ein neuer Parastenus aus dem Iran sowie Bemerkungen über zwei alte kaukasische Parastenen (Coleoptera, Staphylinidae)// Mitteilungen der Schweizerischen Entomologischen Gesellschaft 45. P. 123-126.
Puthz V., 1973. Über einige alte und neue Steninen aus der Sowjetunion (Coleoptera, Staphylinidae)// Reichenbachia 14 (35). P. 291-296.
Puthz V., 1977. Neue westpaläarktische Stenus, vorwiegend aus dem Genfer Museum Coleoptera, Staphylinidae)// Revue Suisse de Zoologie 84 (1). P. 109-117.
Puthz V, 1979a. Drei neue Stenus-Arten aus dem nörddlichen Iran (Coleoptera, Staphylinidae)// Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen 31. P. 54-58.
Puthz V, 1979b. Wissenschaftliches Ergebnis der zoologischen Expedition des Nationalmuseums in Prag nach der Türkei. Coleoptera-Staphylinidae, Subfam. Steninae// Acta Entomologica Musei Nationalis Pragae 39 (1977). P. 319327. (Note: The date of publication is December 30, 1977 as stated in a notation on the back of the title page. According to the author of this article, this issue of the journal was published in 1979 (Herman (2001).
Puthz V., 1980a. Die Stenus-Arten (Stenus s.str. und Nestus Rey) der Orientalis: Bestimmungstabelle und Neubeschreibungen (Coleoptera, Staphylinidae)// Reichenbachia 18 (3). P. 23-41.
Puthz V, 1980b. Zur Synonymie und Stellung einiger Steninen V (Coleoptera, Staphylinidae// Entomologische Blätter für Biologie und Systematik der Käfer 76 (1). P. 33-43.
Puthz V, 1981. Neue westpaläarktische Stenus, vorwiegend aus dem Genfer Museum Coleoptera, Staphylinidae)// Revue Suisse de Zoologie 88 (3). P. 693-706.
Puthz V. 1983. Einige Stenus-Arten aus dem Kaukasus und Elburs (Insecta: Coleoptera: Staphylinidae)// Senckenbergiana Biologica 63 (1982). P. 347-362.
Puthz V., 2001. Beiträge zur Kenntnis der Steninen CCLXX. Neue und alte paläarktische Arten der Gattung Stenus Latreille (Staphylinidae, Coleoptera)// Philippia 10 (1). P. 53-64.
Puthz V., 2003. Stenus alpicola Fauvel und seine Nahverwandten (Coleoptera, Staphylinidae). 274. Beitrag zur Kenntnis der Steninen// Entomologische Blätter für Biologie und Systematik der Käfer 98 (2) (2002). P. 89-110.
Puthz V, 2006. On the genus Stenus Latreille, mostly from southern Africa, with a revision of the Stenus mendicus species-group (Coleoptera: Staphylinidae)// Annals of the Transvaal Museum 43. P. 29-67.
Puthz V., 2008. Stenus Latreille und die segenreiche Himmelstochter (Coleoptera, Staphylinidae) 1// Linzer biologische Beiträge 40 (1). P. 137-230.
Puthz V. 2009. Neue und alte paläarktische Stenus-Arten
(Coleoptera, Staphylinidae) 305. Beitrag zur Kenntnis der Steninen// Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen 61. P. 29-50.
Reitter E., 1882. Bestimmungs-Tabellen der europäischen Coleopteren. V. Paussidae, Clavigeridae, Pselaphidae und Scydmaenidae// Verhandlungen der KaiserlichKöniglichen Zoologisch-Botanischen Gesellschaft in Wien 31 (1881). P. 443-593.
Reitter E., 1898. Siebzehnter Beitrag zur Coleopteren-Fauna des russischen Reiches// Wiener Entomologische Zeitung 17. P. 109-127.
Rougemont G.M., 1988. Notes on some palearctic Stilicus species with special reference to Turkey (Col. Staphylinidae, Paederinae)// Revue Suisse de Zoologie 95 (2). P. 513-520.
Ryvkin A.B., 1990. Stafilinidy podsemeistva Steninae (Coleoptera, Staphylinidae) Kavkaza i sopredel'nykh territorii. pp. 137-234. In: Striganova, B. (ed.), Fauna nazemnykh bespozvonochnykh Kavkaza. Sbornik Nauchnykh Trudov. Moskva: Akademiia Nauka SSSR.
Sabella G., Kurbatov S.A., 2003. Contribution to the knowledge of the genus Tychus Leach, 1817 (Coleoptera: Staphylinidae: Pselaphinae)// Annales de la Société Entomologique de France (N.S.) 38 (3) (2002). P. 299-317.
Sabella G., Bückle C., Brachat V., Besuchet C., 2004. Revision der Paläarktischen arten der gattung Brachygluta Thomson 1859 (Coleoptera, Staphylinidae) 1. Teil// Muséum d'Histoire Naturelle, Genéve, 283 pp.
Sakenin H., Raheb J., Imani S., Havaskary M., Shirdel F., Mohseni H., 2008a. A preliminary survey on dipteran predators and parasitoids and Odonata in Iranian rice fields// Proceedings of National Conference of Agronomical Rice Breeding, Young Research Club Islamic Azad University of Ghaemshahr, 26-27 November, Abstract p. 80 (Full paper in CD Rom, 10 pp) (In Persian with English summary).
Sakenin H., Imani S., Shirdel F., Samin N., Havaskary M., 2008b. Identification of Pentatomidae (Heteroptera) and their host plants in central and eastern Mazandaran province and introducing of many dominant natural enemies// Plant and Ecosystem 15. P. 37-51 (in Persian, English abstract).
Sakenin H., Eslami B., Samin N., Imani S., Shirdel F., Havaskary M., 2008c. A contribution to the most important trees and shrubs as the hosts of woodboring beetles in different regions of Iran and identification of many natural enemies// Plant and Ecosystem 16. P. 2746 (in Persian, English abstract).
Sakenin H., Ghahari H., Tabari M., Monem R., Havaskary M., Rashidi A., 2008d. A preliminary survey on the fauna of ants (Hymenoptera: Formicidae, Mutillidae) and rove beetles (Coleoptera: Staphylinidae) in Iranian rice fields and surrounding grasslands// Proceedings of National Conference of Agronomical Rice Breeding, Young Research Club Islamic Azad University of Ghaemshahr, 26-27 November, Abstract p. 80 (Full paper in CD Rom, 10 pp) (In Persian with English Summary).
Sakenin H., Ghahari H., Ostovan H., Imani S., Samin N., 2010. Faunistic note on rove beetles (Coleoptera: Staphylinidae) from Semnan Province, Iran//
International Society for Southeast Asian Agricultural Sciences, 2009 International Congress, January 11-15, 2010, Pattaya, Thailand, Abstract p. 188.
Saulcy F. de, 1876. Species des Paussides, Clavigérides, Psélaphides & Scydménides de l'Europe et des pays circonvoisins (continued)// Bulletin de la Société d'Histoire Naturelle de Metz 14. P. 25-100.
Scheerpeltz O., 1929. Monographie der Gattung Olophrum ER. (Coleoptera: Staphylinidae)// Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 79 (1). P. 1-257.
Scheerpeltz O., 1933, 1934. Staphylinidae VII: Supplementum I, II. In: Schenkling, S. (ed.), Coleopterorum Catalogus. Pars 129, 130. W. Junk, Berlin. P. 989-1881.
Scheerpeltz O., 1955. Eine neue Art der Gattung Platystethus Mannh., mit einer Bestimmungstabelle der westpaläarktischen Arten und Formen dieser Gattung (Col. Staphylinidae)// Koleopterologische Rundschau 33. P. 78-88.
Scheerpeltz O., 1957. Vorläufige Diagnosen einiger neuer paläarktischer Arten und Formen der Gattungen Paederidus Muls. Rey, Paederus Fabr. (mit den neuen Untergattungen Eopaederus, Paederus s.str. nov., Heteropaederus, Dioncopaederus und Oedopaederus), Parameropaederus nov.gen., Lobopaederus nov.gen. und Megalopaederus nov.gen.// Memorie del Museo di Storia Naturale della Venezia Tridentina 11. P. 447-475.
Scheerpeltz O., 1958a. Wissenschaftliche Ergebnisse der von Herrn Dr. K. Lindberg, Lund, im Jahre 1956 nach der Türkei und Armenien unternommenen Reise. Coleoptera-Staphylinidae// Entomologisk Tidskrift (Supplementum) 78 (1957). P. 3-37.
Scheerpeltz O., 1958b. Eine neue Art der Gattung Eppelsheimius Bernh (Col. Staphylinidae) nebst einer Bestimmungstabelle der bis heute bekannt gewordenen Arten dieser Gattung (89. Beitrag zur Kenntnis der paläarktischen Staphyliniden)// Koleopterologische Rundschau 35 (1957). P. 14-17.
Scheerpeltz O., 1960a. Die von Prof. Dr. A. Gilli während seines Aufenthaltes in Afghanistan 1949/1951 aufgesammelten Staphyliniden (Coleoptera, Staphylinidae)// Koleopterologische Rundschau 37/38. P. 67-81.
Scheerpeltz O., 1960b. Die von Dr. Chr. Lindemann gelegentlich ihrer Reise 1955/56 in Westpakistan aufgesammelten Staphyliden (Coleoptera)// Opuscula Zoologica, München 51. P. 1-7.
Scheerpeltz O., 1961a. Die von den Herren W. Richter und Dr. F. Schäuffele in den Jahren 1954 umd 1956 im Sudiran aufgefundenen Staphyliniden (Col.)// Stuttgarter Beiträge zur Naturkunde 50. P. 1-31.
Scheerpeltz O., 1961b. Ergebnisse der Deutschen Afghanistan-Expedition 1956 der Landessammlungen für Naturkunde Karlsruhe. Staphylinidae (Coleoptera)// Beiträge Naturkundlichen Forschung in Südwestdeutschland 19 (3). P. 245-253.
Scheerpeltz O., 1961c. The 3rd Danish expedition to Central Asia. Zoological results 26. Staphylinidae (Insecta) aus Afghanistan// Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kbenhavn 123. P. 33-50.
Scheerpeltz O., 1961d. Die von Prof. Dr. Hakan Lindberg während seiner 1959 nach Portugal unternommenen Studienreise aufgesammelten Staphyliniden (Col.)// Notulae Entomologicae 40 (4) (1960). P. 132-139.
Scheerpeltz O., 1962. Staphyliniden aus dem Deutschen Entomologischen Institut seinerzeit von Bernhauer als neu erkannt, aber nicht mehr beschreiben. I. Teil: Arten aus der paläarktischen Faunenregion (Coleoptera: Staphylinidae)// Beiträge zur Entomologie 12. P. 565-606.
Scheerpeltz O., 1963a. Coleoptera, Staphylinidae. 116 Beitrag zur Kenntnis der Paläarktischen Staphyliniden// Lunds Universitets Ârsskrift (N.F.) Avd. 2. 58 (11). P. 1-38.
Scheerpeltz O., 1963b. Ergebnisse der Zoologischen Nubien-Expedition 1962. Teil VIII. Coleoptera - Staphylinidae (118. Beitrag zur Kenntnis der paläarktischen Staphyliniden)// Annalen des Naturhistorischen Museums in Wien 66. P. 425-450.
Scheerpeltz O., 1966. Eine neue Art der Grossgattung Staphylinus L., neue Subgenera und Bemerkungen über einige bereits bekannte paläarktische Arten dieser Grossgattung (Col.)// Nachrichtenblatt der Bayerischen Entomologen 15. P. 105-117.
Scheerpeltz O., 1969. Eine neue Art der Gattung Rheochara Mulsant & Rey, nebst einer Bestimmungstabelle der bis heute aus dieser Gattung bekannt gewordenen Arten (Col. Staphyl.)// Koleopterologische Rundschau 46/47 (1968/1969). P. 119-126.
Scheerpeltz O., 1970. Zwei neue Arten der Gattung Scopaeus Erichson vom Südufer des Kaspischen Meeres (Col. Staphylinidae) (142. Beitrag zur Kenntnis der paläarktischen Staphyliniden)// Entomologische Blätter für Biologie und Systematik der Käfer 66 (2). P. 73-79.
Schülke M., 1989a. Eine neue mittelasiatische Art der Gattung Medon Stephens (Coleoptera, Staphylinidae, Paederinae)// Entomologische Nachrichten und Berichte 33 (2). P. 83-87.
Schülke M., 1989b. Bemerkungen zur Verbreitung und Synonymie einiger Arten der Gattung Tachinus Gravenhorst (Coleoptera, Staphylinidae: Tachyporinae)// Entomologische Nachrichten und Berichte 33 (5). P. 230-232.
Schülke M., 1994. Eine neue Art der Gattung Carphacis Des Gozis aus dem Iran (Col., Staphylinidae)//Entomologische Nachrichten und Berichte 38 (1). P. 35-36.
Schülke M., 1998. Zwei bisher verkannte westpaläarktische Sepedophilus-Arten (Coleoptera: Staphylinidae)// Entomologische Blätter für Biologie und Systematik der Käfer 94 (1-2). P. 57-63.
Schülke M., 2000. Eine neue Art der Gattung Carphacis Des Gozis aus der chinesischen Provinz Sichuan (Coleoptera, Staphylinidae, Tachyporinae)// Linzer biologische Beiträge 32 (2). P. 891-895.
Schülke M., 2007. Drei neue Adventivarten der europäischen Staphyliniden-Fauna, mit Bemerkungen zu Coproporus colchicus Kraatz (Coleoptera, Staphylinidae, Tachyporinae)// Entomologische Blätter für Biologie und Systematik der Käfer 102 (1-3) (2006). P. 173-201.
Schülke M., 2008. Neumeldungen von paläarktischen Tachyporinen und Ergänzungen zum «Catalogue of Palaearctic Coleoptera, Volume 2» (Coleoptera:
Staphylinidae: Tachyporinae)// Linzer biologische Beiträge 40 (1). P. 927-942.
Schülke M., 2009. Zur Taxonomie und Faunistik westpaläarktischer Staphylinidae (Coleoptera: Staphylinidae: Omaliinae, Oxytelinae et Tachyporinae)// Linzer biologische Beiträge 41 (1). P. 803-844.
Semenov A.P., 1909. Coleoptera nova faunae turanicae// Russkoe Entomologicheskoe Obozrenie 8 (3-4) (1908). P. 295-304.
Smetana A., 1959. Palaearctic, Oriental and Nearctic species of the genus Ontholestes Gglb. (Col., Staphylinidae)// Acta Entomologica Musei Nationalis Pragae 33. P. 393-412.
Smetana A., 1962. Bestimmungstabelle der mitteleuropäischen Arten der Gattung Quedius Steph. (Col., Staphylinidae)// Entomologische Blätter für Biologie und Systematik der Käfer 58 (3). P. 133-155.
Smetana A., 1965. Zur Kenntnis einiger Philonthus und Gabrius Arten aus der Reitter Sammlung (Coleoptera, Staphylinidae)// Annales Historico-Naturales Musei Nationalis Hungarici 57. P. 253-258.
Smetana A., 1967. Wissenschaftliches Ergebnis der zoologischen Expedition des Nationalmuseum in Prag nach der Türkei. Coleoptera-Staphylinidae, Subfam. Oxytelinae// Acta Entomologica Musei Nationalis Pragae 37. P. 297-324.
Smetana A., 1971. Zur Kenntnis der Staphylinus- und Ocypus-Arten Irans (Coleoptera, Staphylinidae)// Acta Entomologica Bohemoslovaca 68 (4). P. 263-269.
SmetanaA., 1995. Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae). Classification, phylogeny and taxonomic revision// Memoirs on Entomology, International 3. P. I-X, 1-946.
Smetana A., 2004. Family Staphylinidae (except subfamilies Pselaphinae and Scaphidiinae). In: Löbl I. & A. Smetana (eds), Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea, Histeroidea, Staphylinoidea. Apollo Books, Stenstrup. P. 237-698.
Smetana A., Davies A., 2000. Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae)// American Museum Novitates 3287. P. 1-88.
Solodovnikov A.Y., 2002. Taxonomy and faunistics of some species of Quedius Stephens, 1829 from the Caucasus and Asia Minor (Coleoptera, Staphylinidae)// Koleopterologische Rundschau 72. P. 137-158.
Solodovnikov A.Y., 2004. Taxonomy and faunistics of some West Palearctic Quedius Stephens subgenus Raphirus Stephens (Coleoptera, Staphylinidae, Staphylininae)// Koleopterologische Rundschau 74. P. 221-243.
Solodovnikov A.Y., 2005. New and little known species of Quedius from West Palaearctic (Coleoptera, Staphylinidae, Staphylininae)// Zootaxa 902. P. 1-13.
Solodovnikov A.Y., Grebennikov K.A., 2005. Revision of Physetops Mannerheim 1830, a monotypic genus with the polymorphic species P tataricus (Pallas 1773) (Coleoptera: Staphylinidae, Staphylininae)// Mitteilungen aus dem Museum für Naturkunde in Berlin (Zoologische Reihe) 81 (1). P. 67-80.
Stourac P., 2000. Drei neue paläarktische Arten der
Gattung Heterothops (Coleoptera: Staphylinidae)// Folia Heyrovskyana 8 (1). P. 67-72.
Stourac P., 2002. Heterothopsbesucheti sp.n. und H. orientalis sp.n. aus der Türkei (Coleoptera : Staphylinidae)// Revue Suisse de Zoologie 109 (4). P. 735-739.
Thayer M.K., 1993. Osellia Zanetti a junior synonym of Orochares Kraatz, with a checklist of Orochares species (Coleoptera: Staphylinidae: Omaliinae)// The Coleopterists Bulletin 47 (3). P. 285-287.
Thayer M.K., 2005. Staphylinidae Latreille, 1802. In: Beutel, R.G. & R.A.B. Leschen, (eds), Coleoptera, Beetles. Vol. 1: Morphology and systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). Handbuch der Zoologie/Handbook of Zoology. Band/Vol. IV: Arthropoda: Insecta (Kristensen, N.P. & R.G. Beutel (eds), Teilband/Part 38. Walter de Gruyter, Berlin & New York. P. 296-344.
Tirgari S. and Nikbakhtzadeh M.R. 2002. Paederus beetles (Coleoptera: Staphylinidae): an urban problem in Iran// Proceedings of the 4th International Conference on Urban Pests, Charleston, South Carolina. P. 401-407.
Tronquet M., 2009. Révision des Aleochara (Heterochara) d'Europe, du basin méditerranéen et des îles Atlantiques (Coleoptera: Staphylinidae, Aleocharinae, Aleocharini)// Revue de l'Association Roussillonnaise d'Entomologie (R.A.R.E) 18 (3). P. 93-125.
Ullrich W.G., 1975. Monographie der Gattung Tachinus Gravenhorst (Coleoptera: Staphylinidae), mit Bemerkungen zur Phylogenie und Verbreitung der Arten. Dissertation zur Erlangung des Doktorgrades der Mathematisch Naturwissenschaftlichen Fakultät der Christian-Albrschts-Universität zu Kiel. 365 pp, 61 pls, Kiel.
Vit S. 2004. Subfamily Mastiginae (Scydmaenidae)
Fleming, 1821. In: Löbl I. & A. Smetana (eds), Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup. P. 227-228.
White RE., 1983. A field guide to the beetles of North America. (Peterson Field Guide Series). Boston, MA: Houghton Mifflin, XII + 368 p.
Winkler A. 1925. Catalogus Coleopterorum regionis Palaearcticae 1 (3). P. 241-368. Albert Winkler, Wien.
Zanetti A., 1993. Contribute alla conoscenza degli Eusphalerum del Caucaso, dell'Anatolia e delle regioni vicine. (Coleoptera, Staphylinidae: Omaliinae)// Bollettino del Museo Civico di Storia Naturale di Verona 17 (1990). P. 213-263.
Zargari O., Kimyai-Asadi A., Fathalikhani F., Panahi M., 2003. Paederus dermatitis in northern Iran: a report of 156 cases// International Journal of Dermatology 42 (8). P. 608-612.
Zehzad B., Kiabi B.H., Madjoonian H., 2002. The natural areas and landscape of Iran: an overview// Zoology in the Middle East 26. P. 7-10.
Zerche L., 1990. Monographie der paläarktischen Coryphiini (Coleoptera, Staphylinidae, Omaliinae)// Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik. 413 pp.
Zerche L., 1991. Beitrag zur Taxonomie und Verbreitung der Gattung Deliphrosoma Reitter 1909 (Coleoptera, Staphylinidae, Omaliinae)// Beiträge zur Entomologie 41 (2). P. 313-332.
Zerche L., 1993. Monographie der paläarktischen Coryphiini (Coleoptera, Staphylinidae, Omaliinae). Supplementum 1// Beiträge zur Entomologie 43 (2). P. 319-374.
