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16Амурский зоологический журнал I(4), 2009. 283-295
УДК 595.768.1
Amurian zoological journal I(4), 2009. 283-295
CONTRIBUTION TO THE KNOWLEDGE OF THE MESOZOIC CURCULIONOIDEA (COLEOPTERA)
A.A. Legalov
[Легалов А.А. К познанию мезозойских жуков надсемейства Curculionoidea (Coleoptera)]
Institute of Animal Systematics and Ecology, Siberian Zoological Museum, Frunze street, 11, 630091, Novosibirsk, Russia Институт систематики и экологии животных СО РАН, Сибирский зоологический музей, Фрунзе, 11, 630091, Новосибирск, Россия. E-mail: legalov@ngs.ru Key words: Coleoptera, Curculionoidea, Mesozoic.
Ключевые слова: Coleoptera, Curculionoidea, Мезозой.
Summary. 111 species of the Curculionid-beetles are known from the Mesozoic (Obrieniidae - 7, Nemonychidae - 63, Ithyceridae -24, Belidae - 4, Anthribidae - 2, Rhynchitidae - 3, Brentidae - 2, Curculionidae - 5, Scolytidae - 1). New subfamily Metrioxenoidinae Legalov, subfam.n. (type genus: Metrioxenoides Gratshev, Zherikhin, Jarzembowski, 1998), new tribes Kenderlykanini Legalov, trib.n. (type genus: Kenderlyka Zherikhin & Gratshev, 1993), Probelini Legalov, trib.n. (type genus: Probelus L. Arnoldi, 1977), Karataucarini Legalov, trib.n. (type genus: Karataucar Legalov, gen.n.), Gobicarini Legalov, trib.n. (type genus: Gobicar Gratshev & Zherikhin, 1999), Cretanthribini Legalov, trib.n. (type genus: Cretanthribus Legalov, gen.n.), Axeirodiellini Legalov, trib.n. (type genus: Axeirodiellus Zhericin & Gratshev, 2004), Cretuliini Legalov, trib.n. (type genus: Cretulio Zherikhin, 1993), new subtribe Abrocarina Legalov, subtrib.n. (type genus: Abrocar Liu & Ren, 2006) of tribe Baissorhynchini Zherikhin, 1993, new genera Archaeorrhynchoides Legalov, gen.n. (type species: Archaeorrhynchus paradoxopus L. Arnoldi, 1977), Arnoldibelus Legalov, gen.n. (type species: Belonotaris karatavicus L. Arnoldi, 1977), Belonotaroides Legalov, gen.n. (type species: Belonotaris lineatipunctatus L. Arnoldi, 1977), Khetanamonyx Legalov, gen.n. (type species: Oxycorynoides crassirostris Zherikhin, 1993), Karataucar Legalov, gen.n. (type species: Oxycorynoides progressivus L. Arnoldi, 1977), Gratshevicar Legalov, gen.n. (type species: Scelocamptus dubius L. Arnoldi, 1977), Brasilnemonyx Legalov, gen.n. (type species: Brasilnemonyx zherichini Legalov, sp.n.), Chinabrenthorrhinus Legalov, gen.n. (type species: Brenthorrhinus longidigitatus Ren, 1995), Abrenthorrhinus Legalov, gen.n. (type species: Brenthorrhinus brevirostris Gratshev & Zherikhin, 1996), Cretanthribus Legalov, gen.n. (type species: Anthribites cretaceus Zherikhin, 1993), Orapauletes Legalov, gen.n. (type species: Orapauletes cretaceus Legalov, sp.n.), new subgenera Probeloides Legalov, subgen.n. (type species: Probelus tibialis L. Arnoldi, 1977) of the genus Probelus L. Arnoldi, 1977, Pseudoxycorynoides Legalov, subgen.n. (type species: Oxycorynoides brevipes L. Arnoldi, 1977) of the genus Oxycorynoides L. Arnoldi, 1977, Hispanocarodes Legalov, subgen.n. (type species: Gobicar hispanicus Gratshev & Zherikhin, 2000) and Rugosocar Legalov, subgen.n. (type species: Cretonanophyes rugosithorax Gratshev & Zherikhin, 2000) of the genus Montsecanomalus Soriano, Gratshev, Delclos, 2006 and new species Brasilnemonyx zherichini Legalov, sp.n. (Early Cretaceous, Brazil: Santana) and Orapauletes cretaceus Legalov, sp.n. (Late Cretaceous (Botswana: Orap) are described. New taxonomic name Montsecanomalus sorianoi Legalov, nom.n. for M. zherichini Soriano, Gratshev, Delclos, 2006 non Liu & Ren, 2006 is proposed. New synonyms Procurculionini L. Arnoldi, 1977, syn.n. for Eccoptarthrinae L. Arnoldi, 1977, Probelopsis L. Arnoldi, 1977, syn.n. (type species: Probelopsis acutiapex L. Arnoldi, 1977) for Probelus L. Arnoldi, 1977 (type species: Probelus curvispinus L. Arnoldi, 1977), Pseudobrenthorrhinus Gratshev & Zherikhin, 1996, syn.n. (type species: Pseudobrenthorrhinus crassicornis Gratshev & Zherikhin, 1996) for Eccoptarthrus L. Arnoldi, 1977 (type species: Eccoptarthrus crassipes L. Arnoldi, 1977) are presented. New systematic placements are proposed: Belonotaris L. Arnoldi, 1977, placem.n. (type species: Belonotaris punctatissimus L. Arnoldi, 1977) is transferred from tribe Eobelini to tribe Oxycorynoidini; Paleocartinae Legalov, 2003, placem.n., Nebrenthorrhinini Legalov, 2007, placem.n., Eccoptarthrinae L. Arnoldi, 1977, placem.n., Brenthorrhinoidini Legalov, 2003, placem.n. are transferred from family Ithyceridae to family Nemonychidae; Montsecanomalus hispanicus (Gratshev & Zherikhin, 2000), placem.n. is transferred from Gobicar Gratshev & Zherikhin, 1999 to Montsecanomalus Soriano, Gratshev, Delclos, 2006; Zherichiniletes Legalov, 2007, placem.n. is transferred from Auletini subtribe incertae sedis to tribe Sanyrevilleini Legalov, 2003. Paleocartinae Legalov, 2003, stat.n. and Distenorrhininae L. Arnoldi, 1977, stat.n. are upgraded from tribes to subfamilies. 15 new combinations are proposed: Archaeorrhynchoides latitarsis (L. Arnoldi, 1977), comb.n., A. paradoxopus (L. Arnoldi, 1977), comb.n., Probelus acutiapex (L. Arnoldi, 1977), comb.n., Arnoldibelus karatavicus (L. Arnoldi, 1977), comb.n., Belonotaroides lineatipunctatus (L. Arnoldi, 1977), comb.n., Khetanamonyx crassirostris (Zherikhin, 1993), comb.n., Karataucar progressivus (L. Arnoldi, 1977), comb.n., Gratshevicar dubius (L. Arnoldi, 1977), comb.n., Eccoptarthrus crassicornis (Gratshev & Zherikhin, 1996), comb.n., E. magnus (Gratshev & Zherikhin, 1996), comb.n., E. tenuicornis (Gratshev & Zherikhin, 1996), comb.n., Abrenthorrhinus brevirostris (Gratshev & Zherikhin, 1996), comb.n., Chinabrenthorrhinus longidigitatus (Ren, 1995), comb.n., Montsecanomalus hispanicus (Gratshev & Zherikhin, 2000), comb.n. and Cretanthribus cretaceus (Zherikhin, 1993), comb.n.
Резюме. Из Мезозоя известно 111 видов долгоносикообразных жуков (Obrieniidae - 7, Nemonychidae - 63, Ithyceridae - 24, Belidae - 4, Anthribidae - 2, Rhynchitidae - 3, Brentidae - 2, Curculionidae - 5, Scolytidae - 1). В работе описаны новое подсемейство Metrioxenoidinae Legalov, subfam.n. (типовой род: Metrioxenoides Gratshev, Zherikhin, Jarzembowski, 1998), новые трибы Kenderlykanini Legalov, trib.n. (типовой род: Kenderlyka Zherikhin & Gratshev, 1993), Probelini Legalov, trib.n. (типовой род: Probelus L. Arnoldi, 1977), Karataucarini Legalov, trib.n. (типовой род: Karataucar Legalov, gen.n.), Gobicarini Legalov, trib.n. (типовой род: Gobicar Gratshev & Zherikhin, 1999), Cretanthribini Legalov, trib.n. (типовой род: Cretanthribus Legalov, gen.n.), Axeirodiellini Legalov, trib.n. (типовой род: Axeirodiellus Zhericin & Gratshev, 2004), Cretuliini Legalov, trib.n. (типовой род: Cretulio Zherikhin, 1993), новая подтриба Abrocarina Legalov, subtrib.n. (типовой род: Abrocar Liu & Ren, 2006) трибы Baissorhynchini Zherikhin, 1993, новые рода Archaeorrhynchoides Legalov, gen.n. (типовой вид: Archaeorrhynchus paradoxopus L. Arnoldi, 1977), Arnoldibelus Legalov, gen.n. (типовой вид: Belonotaris karatavicus L. Arnoldi, 1977), Belonotaroides Legalov, gen.n. (типовой вид: Belonotaris lineatipunctatus L. Arnoldi, 1977), Khetanamonyx Legalov, gen.n. (типовой вид: Oxycorynoides crassirostris Zherikhin, 1993), Karataucar Legalov, gen.n. (типовой вид: Oxycorynoides progressivus L. Arnoldi, 1977), Gratshevicar Legalov, gen.n. (ти-
повой вид: Scelocamptus dubius L. Arnoldi, 1977), Brasilnemonyx Legalov, gen.n. (типовой вид: Brasilnemonyx zherichini Legalov, sp.n.), Chinabrenthorrhinus Legalov, gen.n. (типовой вид: Brenthorrhinus longidigitatus Ren, 1995), Abrenthorrhinus Legalov, gen.n. (типовой вид: Brenthorrhinus brevirostris Gratshev & Zherikhin, 1996), Cretanthribus Legalov, gen.n. (типовой вид: Anthribites cretaceus Zherikhin, 1993), Orapauletes Legalov, gen.n. (типовой вид: Orapauletes cretaceus Legalov, sp.n.), новые подрода Probeloides Legalov, subgen.n. (типовой вид: Probelus tibialis L. Arnoldi, 1977) рода Probelus L. Arnoldi, 1977, Pseudoxycorynoides Legalov, subgen.n. (типовой вид: Oxycorynoides brevipes L. Arnoldi, 1977) рода Oxycorynoides L. Arnoldi, 1977, Hispanocarodes Legalov, subgen.n. (типовой вид: Gobicar hispanicus Gratshev & Zherikhin, 2000) и Rugosocar Legalov, subgen.n. (типовой вид: Cretonanophyes rugosithorax Gratshev & Zherikhin, 2000) рода Montsecanomalus Soriano, Gratshev, Delclos, 2006, новые виды Brasilnemonyx zherichini Legalov, sp.n. (нижний мел, Бразилия: Сантана) и Orapauletes cretaceus Legalov, sp.n. (верхний мел, Ботсвана: Орап). Предложено новое замещающее название Montsecanomalus sorianoi Legalov, nom.n. дляM. zherichini Soriano, Gratshev, Delclos, 2006 non Liu & Ren, 2006. Procurculionini L. Arnoldi, 1977, syn.n. сведена в синонимы к Eccoptarthrinae L. Arnoldi, 1977, Probelopsis L. Arnoldi, 1977, syn.n. (типовой вид: Probelopsis acutiapex L. Arnoldi, 1977) сведен в синонимы к Probelus L. Arnoldi, 1977 (типовой вид: Probelus curvispinus L. Arnoldi, 1977), Pseudobrenthorrhinus Gratshev & Zherikhin, 1996, syn.n. (типовой вид: Pseudobrenthorrhinus crassicornis Gratshev & Zherikhin, 1996) сведен в синонимы к Eccoptarthrus L. Arnoldi, 1977 (типовой вид: Eccoptarthrus crassipes L. Arnoldi, 1977). Изменено систематическое положение Belonotaris L. Arnoldi, 1977, placem.n. (типовой вид: Belonotarispunctatissimus L. Arnoldi, 1977) (из трибы Eobelini в трибу Oxycorynoidini), Paleocartinae Legalov, 2003, placem.n., Eccoptarthrinae L. Arnoldi, 1977, placem.n., Nebrenthorrhinini Legalov, 2007, placem.n., Brenthorrhinoidini Legalov, 2003, placem.n. (из семейства Ithyceridae в семейство Nemonychidae), Montsecanomalus hispanicus (Gratshev & Zherikhin, 2000), placem.n. (из рода Gobicar Gratshev & Zherikhin, 1999 в род Montsecanomalus Soriano, Gratshev, Delclos, 2006), Zherichiniletes Legalov, 2007, placem.n. (из Auletini subtribe incertae sedis в трибу Sanyrevilleini Legalov, 2003). Изменен статус Paleocartinae Legalov, 2003, stat.n. и Distenorrhininae L. Arnoldi, 1977, stat.n. (из триб до подсемейств). Установлено 15 новых комбинаций (Archaeorrhynchoides latitarsis (L. Arnoldi, 1977), comb.n., A. paradoxopus (L. Arnoldi, 1977), comb.n., Probelus acutiapex (L. Arnoldi, 1977), comb.n., Arnoldibelus karatavicus (L. Arnoldi, 1977), comb.n., Belonotaroides lineatipunctatus (L. Arnoldi, 1977), comb.n., Khetanamonyx crassirostris (Zherikhin, 1993), comb.n., Karataucar progressivus (L. Arnoldi, 1977), comb.n., Gratshevicar dubius (L. Arnoldi, 1977), comb.n., Eccoptarthrus crassicornis (Gratshev & Zherikhin, 1996), comb.n., E. magnus (Gratshev & Zherikhin, 1996), comb.n., E. tenuicornis (Gratshev & Zherikhin, 1996), comb.n., Abrenthorrhinus brevirostris (Gratshev & Zherikhin, 1996), comb.n., Chinabrenthorrhinus longidigitatus (Ren, 1995), comb.n., Montsecanomalus hispanicus (Gratshev & Zherikhin, 2000), comb.n. и Cretanthribus cretaceus (Zherikhin, 1993), comb.n.).
INTRODUCTION
Curculionid beetles play major role in ecosystems being phytophages of various plants. Transition from development on generative organs of plants to vegetative parts was a basis for differentiation of Curculionids in the Mesozoic. The basic groups of Curculionoidea were formed in Mesozoic at first on proangiosperms and then on angiosperms. Study of Mesozoic Curculionid beetles is necessary for the decision of problems of evolution and phylogeny of recent groups.
111 Curculionoidea species are hitherto known from the Mesozoic (6 species from the Triassic, 49 species from the Jurassic, 9 species from Jurassic or Cretaceous and 47 species from the Cretaceous (43 from Early and 4 from Late)) [Alonso-Zarazaga, Lyal, 1999; Arnoldi, 1977; Gratshev, 1998; Gratshev, Zherichin, 1995, 1996, 1997, 1999, 2000a, 2000b, 2003; Kirejtshuk, Azar, Beaver, Mandelshtam, Nel, 2009; Kuschel, 1959, 1983; Kuschel, Poinar, 1993; Kuschel, Oberprieler, Rayner, 1994; Legalov, 2003, 2007, 2009a, 2009b; Liu, Ren, 2006, 2007; Liu, Ren, Shih, 2006; Liu, Ren, Tan, 2006; Martynov, 1926; Poinar, 2006, 2008, 2009; Poinar, Brown, 2009; Ponomarenko, Zherikhin, Kirejtshuk, 2004; Ren, 1995; Rheinheimer, 2004; Soriano, Gratshev, Delclos, 2006; Soriano, 2009; Whalley, Jarzembowski, 1985; Zherichin, 1977, 1986, 1993; Zherichin, Gratshev, 1993, 1997, 2003, 2004].
This work is based on the materials of Paleontological Institute of the Russian Academy of Sciences (Moscow).
RESULTS
Key to Mesozoic weevil families (Curculionoidea)
1. Middle coxa impinging on mesepimerum and metepisternum. Body narrow, elongated, with distinct
striae......................................Obrieniidae
- middle coxa not impinging on mesepimerum and
metepisternum. Body wider, with weak striae or without striae...................................................2
2. 1st and 2nd ventrites elongated, much longer than 3rd and
4thventrites.............................................3
- ventrites homonomic, sometimes 1st ventrite a little
elongated................................................5
3. Antennae not geniculate. Rostrum without antennal stria Brentidae
- antennae geniculate. Rostrum with antennal stria......4
4. Rostrum very short...........................Scolytidae
- rostrum elongated........................Curculionidae
5. Antennae inserted dorsally....Ithyceridae (Ulyaninae)
- antennae inserted laterally or ventrally..............6
6. Rostrum sharply narrowed to apex........................
.............................Ithyceridae (Slonikinae)
- rostrum not or slightly narrowed to apex..............7
7. Rostrum widened and flattened, short. Pronotum usually
with basal carina...........................Anthribidae
- rostrum more or less elongated, narrow. Pronotum
without basal carina.....................................8
8. Antennae inserted near basis or in first third of rostrum 9
- antennae inserted submedially or subapically..........11
9. Body flattened. Elytra with carina along the margins ... Belidae (Oxycorininae)
- body convex. Elytra without carina along the margins ... ....................................................... 10
10. Body granulated, larger (5.2 mm).......................
.............................Belidae (Belinae, in part)
- body not granulated, smaller (1.1-3.4 mm)...............
.........................Ithyceridae (Carinae, in part)
11. Segments of tarsi widened............................12
- segments of tarsi not widened, if widened then body
large, elytra and pronotum not granulated.........13
12. Body granulated, larger (6.7 mm)...................
............................Belidae (Belinae, in part)
- body not granulated, smaller (2.5-6.0 mm)...........
.............................Nemonychidae (in part)
13. Antennae located subapically.......................
.............................Nemonychidae (in part)
- antennae located submedially......................14
14. Procoxa occupy almost all prothorax................
........................Ithyceridae (Carinae, in part)
- precoxal part of prothorax elongated..............15
15. Gular suture double......Nemonychidae (in part)
- gular suture single..Ithyceridae (Carinae, in part)
LIST OF THE FAMILY OBRIENIIDAE
2nd ventrite.
Genus Kenderlyka Zherikhin & Gratshev, 1993 Kenderlyka Zherikhin & Gratshev, 1993: 52 Type species: Kenderlyka consobrina Zherikhin & Gratshev, 1993
Kenderlyka consobrina Zherikhin & Gratshev, 1993 Kenderlyka consobrina Zherikhin & Gratshev, 1993: 60 Distribution. Late Triassic (East Kazakhstan: Kenderlyk).
Genus Madygenorhynchus Zherikhin & Gratshev, 1993 Madygenorhynchus Zherikhin & Gratshev, 1993: 52 Type species: Madygenorhynchus multifidus Zherikhin & Gratshev, 1993
Family Obrieniidae Zherikhin & Gratshev, 1993 Obrieniidae Zherikhin & Gratshev, 1993: 51 Type genus: Obrienia Zherikhin & Gratshev, 1993 Remarks. 6 species are described from the Triassic and 1 species from the Jurassic.
Madygenorhynchus multifidus Zherikhin & Gratshev, 1993
Madygenorhynchus multifidus Zherikhin & Gratshev, 1993:59
Distribution. Late Triassic (Kirghizia: Madygen).
Subfamily Obrieniinae Zherikhin & Gratshev, 1993 Obrieniinae Zherikhin & Gratshev, 1993: 51 Type genus: Obrienia Zherikhin & Gratshev, 1993
Tribe Kararhynchini Zherikhin & Gratshev, 1993 Kararhynchinae Zherikhin & Gratshev, 1993: 52 Type genus: Kararhynchus Zherikhin & Gratshev, 1993
Genus Obrienia Zherikhin & Gratshev, 1993 (Col. pl. II: a, b)
Obrienia Zherikhin & Gratshev, 1993: 51 Type species: Obrienia kuscheli Zherikhin & Gratshev, 1993
Obrienia illaetabilis Zherikhin & Gratshev, 1993 Obrienia illaetabilis Zherikhin & Gratshev, 1993: 56 Distribution. Late Triassic (Kirghizia: Madygen). Obrienia ingurgata Zherikhin & Gratshev, 1993 Obrienia ingurgata Zherikhin & Gratshev, 1993: 55 Distribution. Late Triassic (Kirghizia: Madygen). Obrienia kuscheli Zherikhin & Gratshev, 1993 Obrienia kuscheli Zherikhin & Gratshev, 1993: 53 Distribution. Late Triassic (Kirghizia: Madygen).
Genus Guillermia Zherikhin & Gratshev, 1993 Guillermia Zherikhin & Gratshev, 1993: 52 Type species: Guillermia lecticula Zherikhin & Gratshev, 1993
Guillermia lecticula Zherikhin & Gratshev, 1993 Guillermia lecticula Zherikhin & Gratshev, 1993: 57 Distribution. Late Triassic (Kirghizia: Madygen).
Subfamily Kararhynchinae Zherikhin & Gratshev, 1993 Kararhynchinae Zherikhin & Gratshev, 1993: 52 Type genus: Kararhynchus Zherikhin & Gratshev, 1993
Tribe Kenderlykanini Legalov, trib.n.
Type genus: Kenderlyka Zherikhin & Gratshev, 1993 Remarks. For the description, see that of Kenderlyka and Madygenorhynchus [Zherikhin & Gratshev, 1993: 58-60]. Diagnosis. The new tribe differs from tribe Kararhynchini by the subcontigous metacoxa and 1st ventrite shorter than
Genus Kararhynchus Zherikhin & Gratshev, 1993 Genus Kararhynchus Zherikhin & Gratshev, 1993: 52 Type species: Kararhynchus occiduus Zherikhin & Gratshev, 1993
Kararhynchus occiduus Zherikhin & Gratshev, 1993 Kararhynchus occiduus Zherikhin & Gratshev, 1993: 62 Distribution. Late Jurassic (Kazakhstan: Karatau).
FAMILY NEMONYCHIDAE Remarks. 49 species are described from the Jurassic, 11 species from the Cretaceous and 3 species from Jurassic or Cretaceous.
Key to Mesozoic supraspecific taxa of family Nemonychidae
1. Procoxa located in the middle or closer to apical margin
of prothorax.........................................2
- procoxa located closer to basal margin of prothorax...
..................................................... 12
2. Procoxa located closer to apical margin of prothorax. Mandibles narrow. (Eccoptarthrinae L. Arnoldi, 1977) ... 3
- procoxa located in the middle of prothorax. Mandibles
wide.................................................4
3. 1st-2nd segments of funicle widened. Femora narrow ... Eccoptarthrus L. Arnoldi, 1977
- 1st-2nd segments of funicle not widened. Femora
widened..................Procurculio L. Arnoldi, 1977
4. Antennae inserted subapically. Rostrum slightly
elongated. (Brenthorrhininae L. Arnoldi, 1977).......5
- Antennae inserted in the middle or beyond the middle
of rostrum. Rostrum long. (Distenorrhininae L. Arnoldi, 1977)................................................8
5. Mandibles with teeth at exterior margin.
(Brenthorrhinoidini Legalov, 2003) .....................
.......Brenthorrhinoides Gratshev & Zherikhin, 1996
- mandible without teeth at exterior margin. (Brenthorrhinini
L. Arnoldi, 1977).....................................б
6. Rostrum short. Antennae long, reaching humeri. The
widened in the basis (lateral view). Elytra behind almost
rectangular (lateral view)..................................22
- frons strongly convex. Rostrum almost cylindrical in the basis (lateral view). Elytra behind smoothed (lateral view)..........................................................24
greatest width of pronotum is in the basis......................... 22. Rostrum thick and short. Body short
....................Abrenthorrhinus Legalov, gen.n.
- rostrum longer. Antennae shorter, not reaching elytra. The greatest width of pronotum is in the middle......7
.....................Khetanamonyx Legalov, gen.n.
- rostrum longer and thin. Body elongated. (Oxycorynoides L. Arnoldi, 1977).................................23
7. Tarsi narrow. Femora slightly widened......................... 23. Frons flat. Rostrum strongly triangular widened in the
................. Chinabrenthorrhinus Legalov, gen.n.
- tarsi wide. Femora distinctly widened...................
.....................Brenthorrhinus L. Arnoldi, 1977
8. Body narrower. Pronotum with weak lateral carina.
(Distenorrhinus L. Arnoldi, 1977)......................9
- body wider. Pronotum with strong lateral carina......11
9. Protibiae widened, biconcave. Profemora strongly
basis (lateral view) ....................................
..............Pseudoxycorynoides Legalov, subgen.n.
- frons slightly convex. Rostrum slightly triangular,
widened in the basis (lateral view) .....................
.................................. Oxycorynoides s. str.
24. Profemora slightly widened............................25
- Profemora strongly widened..............................28
widened..................................................... 25. Tarsi not elongated. Intervals of elytra narrow
.....Parabrenthorrhinus Gratshev & Zherikhin, 1996
- tibiae narrow, straight. Profemora slightly widened or not
widened .............................................10
10. Larger (greater than 5 mm in length). Rostrum shorter Asternorrhinus Gratshev & Zherikhin, 1996
- smaller (less than 5 mm). Rostrum longer................
..................................Distenorrhinus s. str.
11. Apical margin of prothorax strongly curved. Labrum
.......................Belonotaroides Legalov, gen.n.
- Tarsi elongated. Intervals of elytra wide............26
27. Body short. Elytra with distinct rows of points Cratomacer Zherichin & Gratshev, 2004
- body elongated. Elytra without distinct rows of points.
Elytra flattened...Microprobelus Liu, Ren, Shih, 2006
28. Forehead very strongly convex. Rostrum separated
from frons..................Ampliceps L. Arnoldi, 1977
elongated.................................................. - frons strongly convex. Rostrum not separated from frons
.....Microbrenthorrhinus Gratshev & Zherikhin, 2000
- apical margin of prothorax straight. Labrum not elongated .......Megabrenthorrhinus Gratshev & Zherikhin, 1996
12. Eyes not convex or slightly convex..................13
- eyes strongly convex..................................35
13. Antennae inserted submedially. (Eobelinae L. Arnoldi,
1977)................................................. 14
- antennae inserted subapically.........................31
14. Metathorax short. (Nanophydini L. Arnoldi, 1977).......
..........................Nanophydes L. Arnoldi, 1977
- metathorax long.......................................15
15. Rostrum thin and long. Pronotum with weak lateral
carinae. Larger (7.0-10.2 mm).........................16
- rostrum thicker and short. Pronotum with sharper lateral
carinae. Smaller (2.1-6.2 mm).........................20
16. Tarsi strongly widened. (Eobelini L. Arnoldi, 1977) .... 17
- tarsi not widened or slightly widened. (Probelini Legalov,
trib.n.).............................................. 19
17. Protarsi strongly widened and large....................
..................Archaeorrhynchoides Legalov, gen.n.
- protarsi widened, but not large.......................18
18. Profemora narrow.............Eobelus L. Arnoldi, 1977
...................................................... 29
29. Rostrum long. Body larger (6.2 mm). Protibiae straight
.............................Belonotaris L. Arnoldi, 1977
- rostrum shorter. Body smaller (3.6-4.0 mm). Protibiae
slightly curved..............Scelocamptus L. Arnoldi, 1977
30. 1st segment of tarsi widened...........................
..............................Karataucar Legalov, gen.n.
- 1st-3rd segments of tarsi widened.........................
.......................... Gratshevicar Legalov, gen.n.
31. Elytra gently punctate, with weak striae or without
them. (Paleocartinae Legalov, 2003)................... 32
- elytra roughly punctate, with distinct striae.
(Metrioxenoidinae Legalov, subfam.n.).................33
32. Mandible with tooth at exterior margin. Pronotum sharply transversal. Profemora strongly widened.
(Nebrenthorrhinini Legalov, 2007) .......................
.....................Nebrenthorrhinus Legalov, 2003
- mandible without tooth at exterior margin. Pronotum slightly transversal. Profemora slightly widened.
(Paleocartini Legalov, 2003) ............................
.............................Paleocartus Legalov, 2003
33. Sizes of body larger and metathorax convex.............
..........................Brasilnemonyx Legalov, gen.n.
....... 34
- Profemora widened....................................... - sizes of body smaller and metathorax flat .
....................Archaeorrhynchus Martynov, 1926
19. Tibiae with mucro and sizes of the body smaller (7.08.2 mm)........................Probelus L. Arnoldi, 1977
- tibiae without mucro and sizes of the body larger (7.5-8.5
mm).......................Arnoldibelus Legalov, gen.n.
20. Segments of tarsi not widened, narrower than tibiae.
(Oxycorynoidini L. Arnoldi, 1977)....................21
- segments of tarsi widened, wider than tibiae.
(Karataucarini Legalov, trib.n.).....................30
21. Frons flat or slightly convex. Rostrum triangular
34. Rostrum long. Eyes smaller. Head and pronotum small
punctate.........Libanorhinus Kuschel & Poinar, 1993
- rostrum short. Eyes larger. Head and pronotum
largely punctate .......................................
Metrioxenoides Gratshev, Zherikhin, Jarzembowski, 1997
35. Elytra with distinct striae. (Rhinorhynchinae Voss,
1922)......................Renicimberis Legalov, 2009
- elytra without striae. (Cimberindinae des Gozis, 1882) ...........................Chinocimberis Legalov, 2009
Subfamily Eobelinae L. Arnoldi, 1977 Tribe Eobelini L. Arnoldi, 1977 Genus Archaeorrhynchoides Legalov, gen.n. (col. pl. II, e) Type species: Archaeorrhynchus paradoxopus L. Arnoldi, 1977
Remarks. For the description, see that ofArchaeorrhynchus paradoxopus [Arnoldi, 1977: 149-150].
Diagnosis. The new genus is close to genus Archaeorrhynchus but differs by the strongly widened and larger protarsi.
Etymology. The name is formed by addition of the ending “-ides” to “Archaeorrhynchus”.
Archaeorrhynchoides latitarsis (L. Arnoldi, 1977), comb.n.
Archaeorrhynchus latitarsis L. Arnoldi, 1977: 150 Distribution. Late Jurassic (Kazakhstan: Karatau). Archaeorrhynchoides paradoxopus (L. Arnoldi, 1977), comb.n.
Archaeorrhynchus paradoxopus L. Arnoldi, 1977: 149 Distribution. Late Jurassic (Kazakhstan: Karatau).
Tribe Probelini Legalov, trib.n.
Type genus: Probelus L. Arnoldi, 1977 Description. Body brown. Rostrum long, straight or curved. Antennae inserted submedially. Frons convex. Eyes large. Temples short. Antennae long, reaching the middle of pronotum. Pronotum not elongated, punctate, with lateral carina. Elytra elongated, probable with weak rows of points. Procoxa located near basal margin of prothorax. Precoxal part of pothorax elongated. Metathorax elongated. Abdomen convex. 1st ventrite of almost equal length. 5th ventrite sometimes elongated. Legs long. Procoxa large. Femora slightly widened. Tibiae slightly curved. Metatibiae with mucro. Tarsi more or less long, not widened. Clausal segment elongated with 2 claws. Claws with teeth. Length of body: 7.0-8.5 mm.
Diagnosis. The new tribe differs from tribe Eobelini by not widened segments of tarsi.
Genus Probelus L. Arnoldi, 1977 (Col. pl. I, b, c) Probelus L. Arnoldi, 1977: 151 Type species: Probelus curvispinus L. Arnoldi, 1977 Probelopsis L. Arnoldi, 1977: 153, syn.n.; type species: Probelopsis acutiapex L. Arnoldi, 1977 Remarks. A study of type species of this genera has shown that Probelopsis L. Arnoldi, 1977, syn.n. is synonym to Probelus L. Arnoldi, 1977.
Subgenus Probelus s. str.
Probelus (Probelus) acutiapex (L. Arnoldi, 1977), comb.n. (Col. pl. I, c)
Probelopsis acutiapex L. Arnoldi, 1977: 154 Distribution. Late Jurassic (Kazakhstan: Karatau). Probelus (Probelus) curvispinus L. Arnoldi, 1977 Probelus curvispinus L. Arnoldi, 1977: 151 Distribution. Late Jurassic (Kazakhstan: Karatau). Probelus (Probelus) longitarsus L. Arnoldi, 1977 Probelus longitarsus L. Arnoldi, 1977: 152 Distribution. Late Jurassic (Kazakhstan: Karatau).
Subgenus Probeloides Legalov, subgen.n.
Type species: Probelus tibialis L. Arnoldi, 1977 Remarks. For the description, see that of Probelus tibialis [Arnoldi, 1977: 152-153].
Diagnosis. This new subgenus differs from nominative subgenus by the antennae located behind the middle of rostrum and not widened 5th ventrite.
Etymology. The name is formed by addition of the ending “-ides” to “Probelus”.
Probelus (Probeloides) tibialis L. Arnoldi, 1977 Probelus tibialis L. Arnoldi, 1977: 152 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Arnoldibelus Legalov, gen.n. (Col. pl. I, a) Type species: Belonotaris karatavicus L. Arnoldi, 1977 Remarks. For the description, see that of Belonotaris karatavicus [Arnoldi, 1977: 156].
Diagnosis. This new genus resembles the genus Probelus but differs by the apex of tibiae without mucro and larger sizes of the body.
Etymology. The new genus is named in honour of L. Arnoldi.
Arnoldibelus karatavicus (L. Arnoldi, 1977), comb.n. (Col. pl. I, a)
Belonotaris karatavicus L. Arnoldi, 1977: 156 Distribution. Late Jurassic (Kazakhstan: Karatau).
Tribe Oxycorynoidini L. Arnoldi, 1977 Genus Belonotaris L. Arnoldi, 1977, placem.n.
Type species: Belonotaris punctatissimus L. Arnoldi, 1977
Belonotaris punctatissimus L. Arnoldi, 1977 Belonotaris punctatissimus L. Arnoldi, 1977: 155 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Belonotaroides Legalov, gen.n.
Type species: Belonotaris lineatipunctatus L. Arnoldi, 1977 Remarks. For the description, see that of Belonotaris lineatipunctatus [Arnoldi, 1977: 160-161].
Diagnosis. This new genus resembles the genera Cratomacer and Microprobelus but differs by not elongated tarsi and narrow intervals of elytra.
Etymology. The name is formed by addition of the ending “-ides” to “Belonotaris”.
Belonotaroides lineatipunctatus (L. Arnoldi, 1977), comb.n.
Belonotaris lineatipunctatus L. Arnoldi, 1977: 155 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Oxycorynoides L. Arnoldi, 1977 Subgenus Pseudoxycorynoides Legalov, subgen.n. Type species: Oxycorynoides brevipes L. Arnoldi, 1977 Remarks. For the description, see that of Oxycorynoides brevipes [Arnoldi, 1977: 160-161].
Diagnosis. This new subgenus differs from nominative subgenus by flat forehead and rostrum strongly triangular and widened in the basis (lateral view).
Etymology. The name is formed by addition of the prefix
287
“pseudo-” to “oxycorynoides”.
Oxycorynoides (Pseudoxycorynoides) brevipes L. Arnoldi, 1977
Oxycorynoides brevipes L. Arnoldi, 1977: 160 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Khetanamonyx Legalov, gen.n.
Type species: Oxycorynoides crassirostris Zherikhin, 1993 Remarks. For the description, see that of Oxycorynoides crassirostris [Zherikhin, 1993: 21, 23].
Diagnosis. This new genus differs from genus Belonotaroides by small points in rows on elytra, longer rostrum and larger sizes of body.
Etymology. The name is formed from the words “Khetana” (toponym) and “monyx”.
Khetanamonyx crassirostris (Zherikhin, 1993), comb.n. Oxycorynoides crassirostris Zherikhin, 1993: 21 Distribution. Early Cretaceous (Khabarovskii krai: Khetana).
Tribe Karataucarini Legalov, trib.n.
Type genus: Karataucar Legalov, gen.n. Description. Body brown. Head not constricted behind eyes. Rostrum long, slightly curved. Antennae inserted submedially. Frons more or less wide, convex. Eyes large. Temples short. Antennae long, reaching the middle of pronotum. Funicle segments almost trapezoid. Clava wide. Pronotum wide. Sides almost direct. Elytra slightly elongated, with weak striae. Humeri slightly smoothed. Prothorax long. Procoxa located near basal margin of prothorax. Precoxal part of prothorax elongated. Metathorax elongated. Legs long. Femora slightly widened. Tibiae almost straight. Protarsi long, of equal length to tibiae. 1st segment wide. 2nd segment narrow- or wide-triangular. 3rd segment bilobed. Clausal segment elongated. Length of body: 3.4-4.6 mm.
Diagnosis. The new subtribe differs from tribe Oxycorynoidini by widened segments of tarsi which are wider than tibiae.
Genus Karataucar Legalov, gen.n.
Type species: Oxycorynoides progressivus L. Arnoldi, 1977
Remarks. For the description, see that of Oxycorynoides progressivus [Arnoldi, 1977: 163].
Diagnosis. The new genus is very close to genus Gratshevicar but differs by only 1st segment of tarsi being widened.
Etymology. The name is formed from the words “Karatau” (toponym) and “car”.
Karataucar progressivus (L. Arnoldi, 1977), comb.n. (Col. pl. II, g)
Oxycorynoides progressivus L. Arnoldi, 1977: 163 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Gratshevicar Legalov, gen.n.
Type species: Scelocamptus dubius L. Arnoldi, 1977 Remarks. For the description, see that of Scelocamptus
dubius [Arnoldi, 1977: 165-166].
Diagnosis. The new genus is close to genus Karataucar but differs by widened 1st-3rd segments of tarsi. Etymology. The new genus is named in honour of VG. Gratshev.
Gratshevicar dubius (L. Arnoldi, 1977), comb.n. (Col. pl.
II, f)
Scelocamptus dubius L. Arnoldi, 1977: 165 Distribution. Late Jurassic (Kazakhstan: Karatau).
Subfamily Paleocartinae Legalov, 2003, stat.n., placem.n. Paleocartini Legalov, 2003: 78 Type genus: Paleocartus Legalov, 2003
Tribe Paleocartini Legalov, 2003 Paleocartini Legalov, 2003: 78 Type genus: Paleocartus Legalov, 2003
Genus Paleocartus Legalov, 2003 Paleocartus Legalov, 2003: 78 Type species: Brenthorrhinoidespubescens Gratshev & Zherikhin, 1996
Paleocartus pubescens (Gratshev & Zherikhin, 1996) (Col. pl. III, a)
Brenthorrhinoides pubescens Gratshev & Zherikhin, 1996: 115
Distribution. Late Jurassic (Kazakhstan: Karatau).
Tribe Nebrenthorrhinini Legalov, 2007, placem.n. Nebrenthorrhinina Legalov, 2007: 34 Type genus: Nebrenthorrhinus Legalov, 2003
Genus Nebrenthorrhinus Legalov, 2003 Nebrenthorrhinus Legalov, 2003f: 89 Type species: Nebrenthorrhinus lacasai Gratshev & Zherikhin, 2000
Nebrenthorrhinus lacasai (Gratshev & Zherikhin, 2000) Brenthorrhinoides lacasai Gratshev & Zherikhin, 2000b: 41 Distribution. Early Cretaceous (Spain: Montsec Range).
Subfamily Metrioxenoidinae Legalov, subfam.n.
Type genus: Metrioxenoides Gratshev, Zherikhin, Jarzembowski, 1998 Description. Body brown, with rare semierect setae. Rostrum short or long, slightly curved. Antennae inserted subapically. Frons convex or flattened. Eyes large, oval. Temples short. Antennae long, reaching the middle of pronotum. Pronotum not elongated, largely or small punctate, with lateral carina. Elytra elongated, with rows of large points. Procoxa located near basal margin of prothorax. Metathorax elongated, sometimes convex. Abdomen more or less largely punctate. 1st ventrite little longer than 2nd ventrite. 3rd-5th ventrites of almost equal length. Legs long. Procoxa large. Femora slightly widened. Tibiae almost direct. Protarsi long, little longer than tibiae. Clausal segment elongated with 2 claws. Length of body: 2.0-4.5 mm.
Diagnosis. The new subfamily differs from subfamily
288
Rhinorhynchinae by the rough and large punctuate elytra, distinct lateral carina on pronotum and not convex eyes.
Genus Metrioxenoides Gratshev, Zherikhin, Jarzembowski, 1998 Metrioxenoides Gratshev, Zherikhin, Jarzembowski, 1998: 323
Type species: Metrioxenoidespusillus Gratshev, Zherikhin, Jarzembowski, 1998
Metrioxenoides pusillus Gratshev, Zherikhin,
Jarzembowski, 1998
Metrioxenoides pusillus Gratshev, Zherikhin,
Jarzembowski, 1998: 324
Distribution. Early Cretaceous (England).
Genus Libanorhinus Kuschel & Poinar, 1993 Libanorhinus Kuschel & Poinar, 1993: 144 Type species: Libanorhinus succinus Kuschel & Poinar, 1993
Libanorhinus succinus Kuschel & Poinar, 1993 Libanorhinus succinus Kuschel & Poinar, 1993: 144 Distribution. Early Cretaceous (Lebanon, Amber).
Genus Brasilnemonyx Legalov, gen.n.
Type species: Brasilnemonyx zherichini Legalov, sp.n. Description. Rostrum short. Frons flattened. Eyes large. Pronotum with lateral carina. Elytra elongated, with rows of large points. Metathorax elongated and convex. Abdomen slightly convex. 1st ventrite little longer than 2nd ventrite. 3rd-5th ventrites of almost equal length. Legs long. Length of body: 4.5 mm.
Diagnosis. The new genus differs from other genera of the subfamily Metrioxenoidinae by the larger sizes of body and convex metathorax.
Etymology. The name is formed from the words “Brazil” and “nemonyx”.
Genus Eccoptarthrus L. Arnoldi, 1977 Eccoptarthrus L. Arnoldi, 1977: 169 Type species: Eccoptarthrus crassipes L. Arnoldi, 1977 Pseudobrenthorrhinus Gratshev & Zherikhin, 1996: 114, syn.n.
Type species: Pseudobrenthorrhinus crassicornis Gratshev & Zherikhin, 1996
Remarks. A study of type species of this genera has shown that Pseudobrenthorrhinus Gratshev & Zherikhin, 1996, syn.n. is synonym to Eccoptarthrus L. Arnoldi, 1977.
Eccoptarthrus crassicornis (Gratshev & Zherikhin, 1996), comb.n.
Pseudobrenthorrhinus crassicornis Gratshev & Zherikhin, 1996: 114
Distribution. Late Jurassic (Kazakhstan: Karatau). Eccoptarthrus crassipes L. Arnoldi, 1977 (Col. pl. II, d) Eccoptarthrus crassipes L. Arnoldi, 1977: 169 Distribution. Late Jurassic (Kazakhstan: Karatau). Eccoptarthrus magnus (Gratshev & Zherikhin, 1996), comb.n.
Pseudobrenthorrhinus magnus Gratshev & Zherikhin, 1996: 114
Distribution. Late Jurassic (Kazakhstan: Karatau). Eccoptarthrus tenuicornis (Gratshev & Zherikhin, 1996), comb.n.
Pseudobrenthorrhinus tenuicornis Gratshev & Zherikhin, 1996: 114
Distribution. Late Jurassic (Kazakhstan: Karatau).
Subfamily Brenthorrhininae L. Arnoldi, 1977 Brenthorrhininae L. Arnoldi, 1977: 171 Type genus: Brenthorrhinus L. Arnoldi, 1977
Tribe Brenthorrhinoidini Legalov, 2003, placem.n.
Brenthorrhinoidini Legalov, 2003f: 88 Type genus: Brenthorrhinoides Gratshev & Zherikhin, 1996
Brasilnemonyx zherichini Legalov, sp.n.
Material. Holotype - 1 ex. (American Museum of Natural History) № 43316.
Description. Rostrum short, little longer than pronotum, slightly curved. Frons flattened. Eyes large, oval. Temples short. Pronotum not elongated, with lateral carina. Elytra elongated, with rows of large points. Metathorax elongated and convex. Abdomen slightly convex. 1st ventrite little longer than 2nd ventrite. 3rd-5th ventrites of almost equal length. Legs long. Femora slightly widened. Tibiae almost direct. Length of body: 4.5 mm.
Distribution. Early Cretaceous (Brazil: Santana).
Etymology. The new species is named in honour of V.V Zherichin.
Remarks. See figure 4 in Zherichin & Gratshev [2004: 65].
Subfamily Eccoptarthrinae L. Arnoldi, 1977, placem.n. Eccoptarthrini L. Arnoldi, 1977: 169 Type genus: Eccoptarthrus L. Arnoldi, 1977 Procurculionini L. Arnoldi, 1977: 157, syn.n. Eccoptothoracini L. Arnoldi, 1977: 158
Genus Brenthorrhinoides Gratshev & Zherikhin, 1996 Brenthorrhinoides Gratshev & Zherikhin, 1996: 119 Type species: Brenthorrhinoides mandibulatus Gratshev & Zherikhin, 1996
Brenthorrhinoides mandibulatus Gratshev & Zherikhin, 1996 (Col. pl. III, b)
Brenthorrhinoides mandibulatus Gratshev & Zherikhin, 1996: 115
Distribution. Late Jurassic (Kazakhstan: Karatau). Brenthorrhinoides robustus Gratshev & Zherikhin, 1996 (Col. pl. III, c)
Brenthorrhinoides robustus Gratshev & Zherikhin, 1996: 115
Distribution. Late Jurassic (Kazakhstan: Karatau). Remarks. A study of holotype has shown that antennae located submedially, mandible possibly rhynchitoid type: so this beetle belongs to genus Brenthorrhinoides.
Tribe Brenthorrhinini L. Arnoldi, 1977 Brenthorrhininae L. Arnoldi, 1977: 171 Type genus: Brenthorrhinus L. Arnoldi, 1977
Genus Abrenthorrhinus Legalov, gen.n.
Type species: Brenthorrhinus brevirostris Gratshev & Zherikhin, 1996 Remarks. For the description, see that of Brenthorrhinus brevirostris [Gratshev & Zherikhin, 1996: 126].
Diagnosis. The new genus differs from other genera of this tribe by the short rostrum, long antennae reaching humeri and the greatest width of the pronotum being on the basis. Etymology. The name is formed by addition of the prefix “a-” to “brenthorrhinus”.
Abrenthorrhinus brevirostris (Gratshev & Zherikhin, 1996), comb.n.
Brenthorrhinus brevirostris Gratshev & Zherikhin, 1996: 115 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Brenthorrhinus L. Arnoldi, 1977 Brenthorrhinus L. Arnoldi, 1977: 172 Type species: Brenthorrhinus mirabilis L. Arnoldi, 1977
Brenthorrhinus mirabilis L. Arnoldi, 1977
Brenthorrhinus mirabilis L. Arnoldi, 1977: 172 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Chinabrenthorrhinus Legalov, gen.n.
Type species: Brenthorrhinus longidigitatus Ren, 1995 Remarks. For the description, see that of Brenthorrhinus longidigitatus [Ren, 1995: 90].
Diagnosis. This new genus resembles genus Brenthorrhinus but differs by the narrow tarsi and slightly widened femora. Etymology. The name is formed from the words “Chinese” and “brenthorrhinus”.
Chinabrenthorrhinus longidigitatus (Ren, 1995), comb.n. Brenthorrhinus longidigitatus Ren, 1995: 90.
Distribution. Early Cretaceous (China: Xishan).
Subfamily Distenorrhininae L. Arnoldi, 1977, stat.n. Distenorrhinini L. Arnoldi, 1977: 170
Genus Distenorrhinus L. Arnoldi, 1977 Subgenus Distenorrhinus s. str.
Subgenus Parabrenthorrhinus Gratshev & Zherikhin, 1996
Distenorrhinus (Parabrenthorrhinus) xavieri (Zherichin & Gratshev, 2003)
Distenorrhinus xavieri Zherichin & Gratshev, 2003: 70 Distribution. Early Cretaceous (Spain: Las Hoyas).
Subfamily Cimberindinae des Gozis, 1882 Remarks. The genus Chinocimberis Legalov, 2009 with 2 species (Ch. angustipecteris (Liu, Ren, Tan, 2006) and Ch. magnoculi (Liu, Ren, Tan, 2006)) from Late Jurassic or Early Cretaceous of China has been assigned to this subfamily.
Subfamily Rhinorhynchinae Voss, 1922 Remarks. The genus Renicimberis Legalov, 2009 with species R. latipeeteris (Liu, Ren, Tan, 2006) from Late Jurassic or Early Cretaceous of China has been assigned to the tribe Mecomacerini Kuschel, 1994 of this subfamily.
FAMILY ITHYCERIDAE
Remarks. 1 species are described from the Jurassic,
13 species from the Cretaceous (Col. pl. IV, b-d) and 5
species from Jurassic or Cretaceous.
Key to supraspecific taxa of the supertribe Baissorhynchitae
1. Procoxa occupy almost all prothorax. (Baissorhynchini
Zherikhin, 1993).........................................3
- precoxal part of prothorax the elongated.............2
2. Body wider. Precoxal part of prothorax weaker elongated, 1.3 times longer than diameter of procoxa. Elytra with rows of points. (Gobicarini Legalov, trib.n.) Gobicar Gratshev & Zherikhin, 1999
- body elongated. Precoxal part of prothorax stronger
elongated, 1.5 times longer than diameter of procoxa . Elytra without rows of points. (Mesophyletini Poinar,
2006)........................Mesophyletis Poinar, 2006
3. Antennae inserted submedially. 1st segment of tarsi
not widened. Body elongated. (Abrocarina Legalov, subtrib.n.)....................................Abrocar
- antennae inserted in first third of rostrum. 1st segment
of tarsi usually widened. Body short or elongated. (Baissorhynchina Zherikhin, 1993).......................4
4. 2nd and 3rd segments of the tarsi bilobed.............
........Baissorhynchus Zherikhin, 1977 (Col. pl. IV, a)
- 2nd segment triangular...............................5
5. Frons strongly convex .... Gratshevibelus Soriano, 2009
- frons slightly convex................................б
6. Tibiae curved..............Emanrhynchus Zherikhin, 1993
- tibiae straight......................................7
7. Clausal segment short.................................
........Hispanocar Soriano, Gratshev, Delclos, 2006
- clausal segment long.................................8
8. Protibiae granulated .................................
................Martinsnetoa Zherichin & Gratshev, 2004
- protibiae not granulated.............................9
9. Tibiae wider.........................................10
- tibiae narrower......................................11
10. Pronotum without lateral carina. Rostrum curved,
shorter...............Cretocar Gratshev & Zherikhin, 2000
- pronotum with weak lateral carina. Rostrum almost
direct, longer ..........................................
...........Jarzembowskia Zherikhin & Gratshev, 1997
11. Body shorter (Col. pl. III, d). Rostrum longer.........
.....................Cretonanophyes Zherikhin, 1977
- body more elongated. Rostrum less long. (Montsecanomalus Soriano, Gratshev, Delclos, 2006) .. 12
12. Pronotum rugosely punctate. Elytra forming single arc
(lateral view)...............Rugosocar Legalov, subgen.n.
- pronotum weak punctate or nearly smooth. Elytra not
forming single arc (lateral view)....................13
13. 1st ventrite much longer than 2nd ventrite ...........
..................Hispanocarodes Legalov, subgen.n.
- 1st and 2nd ventrites of almost equal length.........14
14. Rostrum longer. Apex of elytra almost rectangular
(lateral view)...............Leptocar Liu & Ren, 2007
- rostrum less long. Apex of elytra smoothed (lateral view) .................................Montsecanomalus s. str.
Tribe Gobicarini Legalov, trib.n. (Col. pl. III, e)
Type genus: Gobicar Gratshev & Zherikhin, 1999 Remarks. For the description, see that of Gobicar [Gratshev & Zherikhin, 1999: 40-41].
Diagnosis. The new tribe differs from tribe Mesophyletini by the wider body, weaker elongated precoxal part of prothorax that is 1.3 times longer than diameter of procoxa, and elytra with rows of points. From Baissorhynchini, it differs by elongated precoxal part of prothorax.
Genus Gobicar Gratshev & Zherikhin, 1999 (Col. pl. III, e) Gobicar Gratshev & Zherikhin, 1999: 40 Type species: Gobicar ponomarenkoi Gratshev & Zherikhin, 1999
Gobicar ponomarenkoi Gratshev & Zherikhin, 1999 Gobicar ponomarenkoi Gratshev & Zherikhin, 1999: 41 Distribution. Late Jurassic (Mongolia: Gobi-Altai aimak).
Tribe Baissorhynchini Zherikhin, 1993 Baissorhynchini Zherikhin, 1993: 30 Type genus: Baissorhynchus Zherikhin, 1977
Subtribe Abrocarina Legalov, subtrib.n.
Type genus: Abrocar Liu & Ren, 2006 Remarks. For the description, see that of Abrocar [Liu & Ren, 2006: 62-64, 66; 2007: 644-645].
Diagnosis. The new subtribe differs from nominative subtribe by the antennae inserted submedially, not widened 1st segment of tarsi and elongated body.
Genus Abrocar Liu & Ren, 2006 Abrocar Liu & Ren, 2006: 62 Type species: Abrocar brachyorhinos Liu & Ren, 2006
Abrocar brachyorhinos Liu & Ren, 2006 Abrocar brachyorhinos Liu & Ren, 2006: 64 Distribution. Late Jurassic or Early Cretaceous (China: Liaoning Prov.).
Abrocar macilentus Liu & Ren, 2007 Abrocar macilentus Liu & Ren, 2007: 644 Distribution. Late Jurassic or Early Cretaceous (China: Liaoning Prov.).
Subtribe Baissorhynchina Zherikhin, 1993 Baissorhynchini Zherikhin, 1993: 30 Type genus: Baissorhynchus Zherikhin, 1977
Genus Montsecanomalus Soriano, Gratshev, Delclos, 2006 Montsecanomalus Soriano, Gratshev, Delclos, 2006: 558 Type species: Montsecanomalus zherichini Soriano, Gratshev, Delclos, 2006
Subgenus Montsecanomalus s. str.
Montsecanomalus (Montsecanomalus) sorianoi Legalov, nom.n.
Montsecanomalus zherichini Soriano, Gratshev, Delclos, 2006: 559 non Liu & Ren, 2006 Distribution. Early Cretaceous (Spain: Montsec Range). Montsecanomalus (Montsecanomalus) zherichini (Liu &
Ren, 2006)
Cretonanophyes zherichini Liu & Ren, 2006: 61 Distribution. Late Jurassic or Early Cretaceous (China: Liaoning Prov.).
Montsecanomalus (Montsecanomalus) punctatus (Liu & Ren, 2007)
Cretonanophyes punctatus Liu & Ren, 2007: 645 Distribution. Late Jurassic or Early Cretaceous (China: Liaoning Prov.).
Subgenus Leptocar Liu & Ren, 2007 Leptocar Liu & Ren, 2007: 642 Type species: Leptocar polychaetus Liu & Ren, 2007
Montsecanomalus (Leptocar) polychaetus (Liu & Ren,
2007)
Leptocar polychaetus Liu & Ren, 2007: 642 Distribution. Late Jurassic or Early Cretaceous (China: Liaoning Prov.).
Subgenus Hispanocarodes Legalov, subgen.n.
Type species: Gobicar hispanicus Gratshev & Zherikhin, 2000 Remarks. For the description, see that of Gobicar hispanicus [Gratshev & Zherikhin, 2000b: 42-43]. Diagnosis. The new subgenus differs from subgenera Leptocar and Montsecanomalus s. str. by the 1st ventrite much longer than 2nd ventrite.
Etymology. The name is formed from the words “Spanish” and “carodes”.
Montsecanomalus (Hispanocarodes) hispanicus (Gratshev & Zherikhin, 2000), comb.n., placem.n.
Gobicar hispanicus Gratshev & Zherikhin, 2000b: 42 Distribution. Early Cretaceous (Spain: Montsec Range).
Subgenus Rugosocar Legalov, subgen.n.
Type species: Cretonanophyes rugosithorax Gratshev & Zherikhin, 2000 Remarks. For the description, see that of Cretonanophyes rugosithorax [Gratshev & Zherikhin, 2000b: 43]. Diagnosis. The new subtribe differs from other subgenera of this genus by the rugosely punctate pronotum and elytra forming single arc (lateral view).
Montsecanomalus (Rugosocar) rugosithorax (Gratshev & Zherikhin, 2000)
Cretonanophyes rugosithorax Gratshev & Zherikhin, 2000b:43
Distribution. Early Cretaceous (Spain: Montsec Range).
FAMILY BELIDAE Remarks. 4 species are described from Early Cretaceous.
LIST OF THE FAMILY ANTHRIBIDAE
Family Anthribidae Billberg, 1820 (Col. pl. II, c) Subfamily Anthribinae Billberg, 1820 Tribe Cretanthribini Legalov, trib.n. (Col. pl. II, c)
Type genus: Cretanthribus Legalov, gen.n. Remarks. For the description, see that of Anthribites cretaceus [Zherikhin, 1993: 29-30].
Diagnosis. The new tribe differs from other tribes of the Anthribinae by the pronotum without basal carina and sagittate rostrum at apex.
Genus Cretanthribus Legalov, gen.n. (Col. pl. II, c) Type species: Anthribites cretaceus Zherikhin, 1993 Remarks. For the description, see that of Anthribites cretaceus [Zherikhin, 1993: 29-30]. Diagnosis as for Cretanthribini.
Etymology. The name is formed from the words “Cretaceous” and “anthribus”.
Cretanthribus cretaceus (Zherikhin, 1993), comb.n. (Col. pl. II, c)
Anthribites cretaceus Zherikhin, 1993: 29
Distribution. Early Cretaceous (Khabarovskii krai:
Khetana).
Orapauletes cretaceus Legalov, sp.n.
Material. Holotype - 1 ex. (Bernard Price Institute, University of the Witwatersrand, Johannesburg) № BP/2/26974.
Description. Rostrum short, 0.8 times shorter than head, not widened to apex, with mandibles at apex. Antennae located near the rostrum basis. Eyes large, slightly convex. Frons narrower than eye. Temples short. Antennae thin, reaching the middle of pronotum. Pronotum slightly transversal, 1.3 times wider than length, 1.25 times longer than rostrum. Sides very slightly rounded. Elytra almost rectangular, 1.79 times longer than wide. Humeri slightly smoothed. Elytra, possibly, without striae. Pygidium exposed. Femora widened. Length of body: 4.2 mm. Distribution. Late Cretaceous (Botswana: Orap). Remarks. See figure 12 in Kuschel, Oberprieler & Rayner [1994: 143].
Subfamily Choraginae Kirby, 1819 Genus Cretochoragus Soriano, Gratshev, Delclos, 2006 Cretochoragus Soriano, Gratshev, Delclos, 2006: 557 Type species: Cretochoragus pygmaeus Soriano, Gratshev, Delclos, 2006
Cretochoraguspygmaeus Soriano, Gratshev, Delclos, 2006 Cretochoragus pygmaeus Soriano, Gratshev, Delclos, 2006: 557
Distribution. Early Cretaceous (Spain: Montsec Range).
LIST OF THE FAMILY RHYNCHITIDAE Family Rhynchitidae Gistel, 1848 Tribe Sanyrevilleini Legalov, 2003 Subtribe Sanyrevilleina Legalov, 2003
Key to Mesozoic genera of the subtribe Sanyrevilleina
Genus Sanyrevilleus Gratshev & Zherikhin, 2000 Sanyrevilleus Gratshev & Zherikhin, 2000a: 244 Type species: Sanyrevilleus grimaldii Gratshev & Zherikhin, 2000
Sanyrevilleus grimaldii Gratshev & Zherikhin, 2000 Sanyrevilleus grimaldii Gratshev & Zherikhin, 2000a: 245 Distribution. Late Cretaceous (USA: New Jersey, Amber).
Genus Zherichiniletes Legalov, 2007, placem.n. Zherichiniletes Legalov, 2007: 69 Type species: Zherichiniletes khetanus Legalov, 2007
Zherichiniletes khetanus Legalov, 2007(Col. pl. IV, e)
Zherichiniletes khetanus Legalov, 2007: 69 Distribution. Early Cretaceous (Khabarovskii krai: Khetana).
1. Rostrum longer, curved. Body larger (5.0-6.0 mm)..........
......................................... Zherichiniletes
- rostrum shorter, almost direct. Body smaller (2.2-4.2
mm)......................................................2
2. Rostrum longer than pronotum. Body smaller (2.2 mm) Sanyrevilleus
- rostrum shorter than pronotum. Body larger (4.2 mm) .. ............................................... Orapauletes
Genus Orapauletes Legalov, gen.n.
Type species: Orapauletes cretaceus Legalov, sp.n.
LIST OF THE FAMILY BRENTIDAE Family Brentidae Billberg, 1820 Subfamily Eurhynchinae Lacordaire, 1863 Tribe Axeirodiellini Legalov, trib.n.
Type genus: Axeirodiellus Zhericin & Gratshev, 2004 Remarks. For the description, see that of Axeirodiellus [Zhericin & Gratshev, 2004: 66-67].
Diagnosis. The new tribe differs from the nominative tribe by the separated procoxal cavities, postmedial position of the procoxa and smaller and elongate 1st and 2nd segment of antennal club.
Description. Rostrum short, shorter than head, with mandibles at apex. Antennae inserted near the rostrum basis. Eyes large, slightly convex. Forehead narrower than eye. Antennae thin. Pronotum slightly transversal. Elytra almost rectangular, elongated. Elytra, possibly, without striae. Pygidium exposed. Femora widened. Length of body: 4.2 mm.
Diagnosis. The new genus resembles the genus Sanyrevilleus but differs by the rostrum shorter than pronotum and larger body (4.2 mm).
Etymology. The name is formed from the words “Orapa” and “auletes”.
Genus Axeirodiellus Zhericin & Gratshev, 2004 Axeirodiellus Zhericin & Gratshev, 2004: 66 Type species: Axeirodiellus ruptus Zhericin & Gratshev, 2004
Axeirodiellus ruptus Zhericin & Gratshev, 2004 Axeirodiellus ruptus Zhericin & Gratshev, 2004: 67 Distribution. Early Cretaceous (Brazil: Santana).
Tribe Eurhynchini Lacordaire, 1863 Genus Orapaeus Kuschel & Oberprieler, 1994 Orapaeus Kuschel & Oberprieler, 1994: 139 Type species: Orapaeus cretaceus Kuschel & Oberprieler, 1994 292
Orapaeus cretaceus Kuschel & Oberprieler, 1994 Orapaeus cretaceus Kuschel & Oberprieler, 1994: 140 Distribution. Late Cretaceous (Botswana: Orap).
LIST OF THE FAMILY CURCULIONIDAE Family Curculionidae Latreille, 1802 (Col. pl. IV, f, g) Subfamily Erirhininae Schoenherr, 1825 Tribe Cretuliini Legalov, trib.n. (Col. pl. IV, f)
Type genus: Cretulio Zherikhin, 1993 Remarks. For the description, see that of Cretulio [Zherikhin, 1993: 33, 35].
Diagnosis. The new tribe differs from other tribes of the subfamily Erirhininae by the almost roundish body, V-shaped groove on first line of prothorax and truncated last stria of elytra.
Genus Cretulio Zherikhin, 1993 Cretulio Zherikhin, 1993: 33 Type species: Cretulio nucula Zherikhin, 1993
Cretulio nucula Zherikhin, 1993 (Col. pl. IV, f)
Cretulio nucula Zherikhin, 1993: 35
Distribution. Early Cretaceous (Khabarovskii krai:
Khetana).
Subfamily Molytinae Schoenherr, 1823 Genus Gyrbykana Zherikhin, 1993 Gyrbykana Zherikhin, 1993: 36 Type species: Gyrbykana curvipes Zherikhin, 1993
Gyrbykana curvipes Zherikhin, 1993 (Col. pl. IV, g)
Gyrbykana curvipes Zherikhin, 1993: 36
Distribution. Early Cretaceous (Khabarovskii krai:
Khetana).
Subfamily Cryptorhynchinae Schoenherr, 1825 Genus Paleocryptorhynchus Poinar, 2009 Type species: Paleocryptorhynchus burmanus Poinar, 2009
Paleocryptorhynchus burmanus Poinar, 2009 Paleocryptorhynchus burmanus Poinar, 2009: 587 Distribution. Early Cretaceous (Myanmar, Amber).
Subfamily Curculioninae Latreille, 1802 Genus Anchineus Poinar & Brown, 2009 Anchineus Poinar & Brown, 2009: 264 Type species: Anchineus dolichobothris Poinar & Brown, 2009
Anchineus dolichobothris Poinar & Brown, 2009 Anchineus dolichobothris Poinar & Brown, 2009: 266 Distribution. Early Cretaceous (Myanmar, Amber).
Subfamily Entiminae Schoenherr, 1823 Tribe Cylydrorhinini Lacordaire, 1863 Genus Dorotheus Kuschel, 1959 Dorotheus Kuschel, 1959: 50 Type species: Dorotheus guidensis Kuschel, 1959
Dorotheus guidensis Kuschel, 1959 Dorotheus guidensis Kuschel, 1959: 50 Distribution. Late Cretaceous (Chile: Magallanes).
LIST OF THE FAMILY SCOLYTIDAE Family Scolytidae Latreile, 1807 Subfamily Scolytinae Latreile, 1807 Tribe Cylindrobrotini Kirejtshuk, Azar, Beaver, Mandelshtam & Nel, 2009 Cylindrobrotini Kirejtshuk, Azar, Beaver, Mandelshtam & Nel, 2009: 102
Genus Cylindrobrotus Kirejtshuk, Azar, Beaver, Mandelshtam & Nel, 2009 Cylindrobrotus Kirejtshuk, Azar, Beaver, Mandelshtam & Nel, 2009: 105 Type species: Cylindrobrotus pectinatus Kirejtshuk, Azar, Beaver, Mandelshtam & Nel, 2009
Cylindrobrotus pectinatus Kirejtshuk, Azar, Beaver, Mandelshtam & Nel, 2009
Cylindrobrotus pectinatus Kirejtshuk, Azar, Beaver, Mandelshtam & Nel, 2009: 105 Distribution. Early Cretaceous (Lebanon, Amber).
ACKNOWLEDGEMENTS
The author is thankful to his colleagues for the help with the work, especially Prof. A.G. Ponomarenko (Moscow).
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Family Nemonychidae gen. spp.: a - Arnoldibelus karatavicus (L. Amoldi, 1977) (holotype, № 2066/2552), b -Probelus curvispinus L. Amoldi, 1977 (holotype, № 2554/709), с - Probelus acutiapex (L. Amoldi, 1977) (holotype, №2239/1554).
Представители Nemonychidae gen. spp.: a - Arnoldibelus karatavicus (L. Amoldi, 1977) (голотип, № 2066/2552), b - Probelus curvispinus L. Amoldi, 1977 (голотип, № 2554/709), с-Probelus acutiapex (L. Amoldi, 1977) (голотип, № 2239/1554).
Families Obrieniidae, Nemonychidae and Anthribidae gen. spp.: a - Obrienia kuscheli Zherikhin & Gratshev, 1993 (holotype, № 2971/435), b - Obrienia ingurgata Zherikhin & Gratshev, 1993 (holotype, № 2971/612), с - Cretanthribus cretaceus (Zherikhin, 1993) (holotype, № 3800/1209), d - Eccoptarthrus crassipes L. Amoldi, 1977 (holotype, № 2239/1507), e -Archaeorrhynchoides latitarsis (L. Amoldi, 1977) (holotype, № 2239/1531), f- Gratshevicar dubius (L. Amoldi, 1977) (holotype, № 2066/3037), g-Karataucar progressivus (L. Amoldi, 1977) (holotype, № 2066/2317). Представители Obrieniidae, Nemonychidae и Anthribidae gen. spp.: a. - Obrienia kuscheli Zherikhin & Gratshev, 1993 (голотип, № 2971/435), b - Obrienia ingurgata Zherikhin & Gratshev, 1993 (голотип, № 2971/612), с - Cretanthribus cretaceus (Zherikhin, 1993) (голотип, № 3800/1209), d - Eccoptarthrus crassipes L. Amoldi, 1977 (голотип, № 2239/1507), e - Archaeorrhynchoides latitarsis (L. Amoldi, 1977) (голотип, № 2239/1531), f- Gratshevicar dubius (L. Amoldi, 1977) (голотип, № 2066/3037), g - Karataucar progressivus (L. Amoldi, 1977) (голотип, № 2066/2317).
Nemonychidae and Ithyceridae gen. spp.: a - Paleocartus pubescens (Gratshev & Zherikhin, 1996) (holotype, № 2784/1464), b - Brenthorrhinoides mandibulatus Gratshev & Zherikhin, 1996 (holotype, № 2239/1508), с -Brenthorrhinoides robustus Gratshev & Zherikhin, 1996 (holotype, № 2066/2861), d - Cretonanophyes longirostris Zherikhin, 1977 (holotype, № 1668/1772), e - Gobicar ponomarenkoi Gratshev & Zherikhin, 1999 (holotype, № 4270/1125).
Представители Nemonychidae and Ithyceridae gen. spp.: a- Paleocartus pubescens (Gratshev & Zherikhin, 1996) (голотип, № 2784/1464), b - Brenthorrhinoides mandibulatus Gratshev & Zherikhin, 1996 (голотип, № 2239/1508), с - Brenthorrhinoides robustus Gratshev & Zherikhin, 1996 (голотип, № 2066/2861), d - Cretonanophyes longirostris Zherikhin, 1977 (голотип, № 1668/1772), e - Gobicar ponomarenkoi Gratshev & Zherikhin, 1999 (голотип, № 4270/1125).
Ithyceridae, Rhynchitidae, Curculionidae gen. spp.: a - Baissorhynchus tarsalis Zherikhin, 1977 (holotype, № 1989/3010), b - Slonik sibiricus Zherichin, 1977 (holotype, № 1989/2938), с - Ulyaniana excellens Gratshev, 1998 (holotype, № 3559/6465), d - Ulyanisca dentipes Gratshev, 1998 (holotype, № 3559/6466), e - Zherichiniletes khetanus Legalov, 2007 (holotype, № 3800/1208), f- Cretulio nucula Zherikhin, 1993 (holotype, № 3800/1200), g - Gyrbykana curvipes Zherikhin, 1993 (holotype, №4009/1).
Представители Ithyceridae, Rhynchitidae, Curculionidae gen. spp.: a - Baissorhynchus tarsalis Zherikhin, 1977 (голотип, № 1989/3010), b - Slonik sibiricus Zherichin, 1977 (голотип, № 1989/2938), с - Ulyaniana excellens Gratshev, 1998 (голотип, № 3559/6465), d - Ulyanisca dentipes Gratshev, 1998 (голотип, № 3559/6466), e - Zherichiniletes khetanus Legalov, 2007 (голотип, № 3800/1208), f- Cretulio nucula Zherikhin, 1993 (голотип, № 3800/1200), g -Gyrbykana curvipes Zherikhin, 1993 (голотип, №4009/1).
