Contribution to the knowledge of the leaf-rolling weevils (Coleoptera, Rhynchitidae, Attelabidae)

Легалов А.А.

УДК 595.768.1

CONTRIBUTION TO THE KNOWLEDGE OF THE LEAF-ROLLING WEEVILS (COLEOPTERA, RHYNCHITIDAE, ATTELABIDAE)

A.A. Legalov

[Легалов А.А. К познанию жуков-трубковертов (Coleoptera, Rhynchitidae, Attelabidae)]

Institute of Animal Systematics and Ecology, Siberian Zoological Museum, Frunze street, 11, 630091, Novosibirsk, Russia Институт систематики и экологии животных СО РАН, Сибирский зоологический музей, Фрунзе, 11, 630091, Новосибирск, Россия. E-mail: legalov@ngs.ru

Key words: Coleoptera, Curculionoidea, Rhynchitidae, Attelabidae, new taxa, new status, new placement, new record, new combination. Ключевые слова: Coleoptera, Curculionoidea, Rhynchitidae, Attelabidae, новые таксоны, новые статусы, новые систематические положения, новые находки, новые комбинации.

Summary. New genus Tanzanauletes Legalov, gen.n. (type species: Auletobius hustachei Dalla Torre & Voss, 1937), new subgenera Tanzanominurus Legalov, subgen.n. (type species: Kuschelanthus tangensis Legalov, 2007) of the genus Pseudominurus Voss, 1956, Madauletes Legalov, subgen.n. (type species: Auletobius gibbipennis Hustache, 1955) and Rubrauletoides Legalov, subgen.n. (type species: Pseudomesauletes jizushanensis Legalov, sp.n.) of the genus Pseudomesauletes Legalov, 2001 and new species Auletobius iviei Legalov, sp.n. (Madagascar), A. baishuiensis Legalov, sp.n. (Yunnan), Pseudomesauletes collarti Legalov, sp.n. (Zaire), P. friedmani Legalov, sp.n. (Tanzania), P. jizushanensis Legalov, sp.n. (Yunnan), Proinvolvulus flandriensis Legalov, sp.n. (Zaire), Heterorhynchites korshunovi Legalov, sp.n. (Cambodia, Laos, Thailand, Vietnam) and Allapoderus bhutanensis Legalov, sp.n. (Bhutan) are described. New synonyms are established: Auletobius montanus Voss, 1922, syn.n. for Pseudominurus hortulanus (Faust, 1899), Alonsoiauletes Legalov, 2003, syn.n. for Pseudomesauletes s. str., Pseudomesauletes stanleyvillensis Legalov, 2007, syn.n. for P. subsignatus (Voss, 1922), Auletobius subtuberculatus Voss, 1921, syn.n., A. tuberculatus Voss, 1921, syn.n. and A. hirtellus Voss, 1941, syn.n. for Pseudomesauletes formosanus (Voss, 1921), Rhynchites pullus Voss, 1935, syn.n. for Metarhynchites longulus (Gyllenhal, 1833), Rhynchites homalinus Voss, 1930, syn.n. for Cyllorhynchites ursulus rostralis (Voss, 1930), Rhynchitesplatynotus Voss, 1938, syn.n. for Clinorhynchites despectus (Voss, 1938), Rhynchites collarti Voss, 1938, syn.n. and Rh. benitoensis Voss, 1938, syn.n. for Clinorhynchidius flexirostris (Voss, 1938), Rhynchites bipubescens Hustache, 1929, syn.n. for Afrorhynchites villosus (Boheman, 1845), Rhynchites semiopacus Voss, 1939, syn.n. for Proinvolvulus rugosipennis (Voss, 1938), Rhynchites argutus Faust, 1882, syn.n. for Heterorhynchites alcyoneus (Pascoe, 1875), Aspidobyctiscus giganteus Legalov, 2003, syn.n. for Aspidobyctiscus sculpturatus (Pascoe, 1875), Lagenoderes brevicollis Fairmaire, 1897, syn.n. and L. vadoni Voss, 1966, syn.n. for L. dentipennis (Gyllenhal, 1839). New systematic placements are made: Australetobius incostans (Lea, 1910), placem.n. and A. rubricollis (Voss, 1922), placem.n. from Auletobius Desbrochers des Loges, 1869 to Australetobius Legalov, 2007; subgenus Mandelschtamius Legalov, 2003, placem.n. from Kuschelanthus Alonso-Zarazaga & Lyal, 1999, stat.n. to Pseudominurus Voss, 1956; Yunnanuletesperturbatus (Voss, 1930), placem.n. from genus Pseudomesauletes Legalov, 2001 to genus Yunnanuletes Legalov, 2007; Pseudomesauletes podocarpi (Voss, 1933), placem.n. from genus Kuschelanthus Alonso-Zarazaga & Lyal, 1999 to genus Pseudomesauletes Legalov, 2001; Pseudomesauletes fuscofasciatus (Voss, 1933), placem.n. and Pseudomesauletes maculatus (Voss, 1933), placem.n. from subgenus Rubrauletes Legalov, 2003 to subgenus Fascauletes Legalov, 2007; Pseudodicranognathusfuliginosus (Voss, 1933), placem.n. from genus Pseudomesauletes Legalov, 2001 to genus Pseudodicranognathus Legalov, 2001; Caenorhinus rufiventris (Voss, 1921), placem.n. from genus Neoarodepus Legalov, 2003 to genus Caenorhinus C.G. Thomson, 1859; Cartorhynchites breviusculus (Voss, 1939), placem.n. from subgenus Hyperinvolvulus Legalov, 2003 to subgenus Cartorhynchoides Legalov, 2003; Cartorhynchites crassiusculus (Voss, 1938), placem.n. from subgenus Cartorhynchoides Legalov, 2003 to subgenus Hyperinvolvulus Legalov, 2003; Metarhynchites longulus (Gyllenhal, 1833), placem.n. from Rhynchitidae ins. sedis to genus Metarhynchites Voss, 1923; Afrorhynchites villosus (Boheman, 1845), placem.n. from genus Pararhynchites Legalov, 2003 to genus Afrorhynchites Legalov, 2003; Heterorhynchites pruinosus (Voss, 1938), placem.n. from subgenus Eosawadaia Legalov, 2004 to subgenus Eosawadaia s. str; Omolabus centomyrciae (Voss, 1925), placem.n. from subgenus Paralabus Legalov, 2004 to subgenus Pseudomolabus Legalov, 2004; Apleurolabus evanescens (Voss, 1928), placem.n. from genus Pleurolabus Jekel, 1860 to genus Apleurolabus Legalov, 2007; Metriotracheloides regularis (Ter-Minassian, 1986), placem.n. from genus Trachelophoridius Voss, 1929 to genus Metriotracheloides Legalov, 2008. Changes of status: Auletobius ebenus Hustache, 1955, stat.n., Clinorhynchites rufofemoratus (Voss, 1938), stat.n. and Trachelismusprolixus (Voss, 1929), stat.n. are upgraded from variation and form to species, and Kuschelanthus Alonso-Zarazaga & Lyal, 1999, stat.n. and Paralleuscelus Legalov, 2004, stat.n. are downgraded from genera to subgenera. Hamiltoniauletes trifasciatus (Suffrian, 1870) in fauna of Dominican Republic, Pseudomesauletes ueleanus (Voss, 1939) in fauna of Cameroon, P. gestroi (Faust, 1894) in fauna of Laos, Eumetopon flavomaculatus (Voss, 1922) in fauna of Zejiang, Clinorhynchidius flexirostris (Voss, 1938) in fauna of Gabon, Heterorhynchites wahnesi (Hartmann, 1899) in fauna of Sabah, H. elysius (Pascoe, 1875) in faunae of Malaysia, Kalimantan and Java, H. subdentatus (Voss, 1938) in fauna of Kalimantan, Suniops gorochovi Legalov, 2003 in fauna of Thailand, Kobusynaptops verrucosus Legalov & X. Zhang, 2007 in fauna of Sichuan, Riedeliops zolotarenkoi Legalov, 2003 in fauna of Thailand, Parasynaptopsis chinensis (Voss, 1922) in fauna of Anhui, Parasynaptopsis lespedezae koreanus (Voss, 1924) in fauna of Hubei and P. moanus Legalov, 2003 in fauna of Yunnan are recorded for the first time. 30 new combinations are established.

Резюме. В работе описаны новый род Tanzanauletes Legalov, gen.n. (типовой вид: Auletobius hustachei Dalla Torre & Voss, 1937), новые подроды Tanzanominurus Legalov, subgen.n. (типовой вид: Kuschelanthus tangensis Legalov, 2007) рода Pseudominurus Voss, 1956,Madauletes Legalov, subgen.n. (типовой вид: Auletobius gibbipennis Hustache, 1955) и Rubrauletoides Legalov, subgen.n. (типовой вид: Pseudomesauletes jizushanensis Legalov, sp.n.) рода Pseudomesauletes Legalov, 2001 и новые виды Auletobius iviei Legalov, sp.n. (Мадагаскар), A. baishuiensis Legalov, sp.n. (Юньнань), Pseudomesauletes collarti Legalov, sp.n. (Заир), P. friedmani Legalov, sp.n. (Танзания), P. jizushanensis Legalov, sp.n. (Юньнань), Proinvolvulusflandriensis Legalov, sp.n. (Заир), Heterorhynchites korshunovi Legalov, sp.n. (Камбоджа, Лаос, Таиланд и Вьетнам) и Allapoderus bhutanensis Legalov, sp.n. (Бутан). Установлены

новые синонимы: Auletobius montanus Voss, 1922, syn.n. к Pseudominurus hortulanus (Faust, 1899), Alonsoiauletes Legalov, 2003, syn.n. к Pseudomesauletes s. str., Pseudomesauletes stanleyvillensis Legalov, 2007, syn.n. к P. subsignatus (Voss, 1922), Auletobius subtuberculatus Voss, 1921, syn.n., A. tuberculatus Voss, 1921, syn.n. и A. hirtellus Voss, 1941, syn.n. к Pseudomesauletes formosanus (Voss, 1921), Rhynchites pullus Voss, 1935, syn.n. к Metarhynchites longulus (Gyllenhal, 1833), Rhynchites homalinus Voss, 1930, syn.n. к Cyllorhynchites ursulus rostralis (Voss, 1930), Rhynchites platynotus Voss, 1938, syn.n. к Clinorhynchites despectus (Voss, 1938), Rhynchites collarti Voss, 1938, syn.n. и Rh. benitoensis Voss, 1938, syn.n. к Clinorhynchidius flexirostris (Voss, 1938), Rhynchites bipubescens Hustache, 1929, syn.n. к Afrorhynchites villosus (Boheman, 1845), Rhynchites semiopacus Voss, 1939, syn.n. к Proinvolvulus rugosipennis (Voss, 1938), Rhynchites argutus Faust, 1882, syn.n. к Heterorhynchites alcyoneus (Pascoe, 1875), Aspidobyctiscus giganteus Legalov, 2003, syn.n. к Aspidobyctiscus sculpturatus (Pascoe, 1875), Lagenoderes brevicollis Fairmaire, 1897, syn.n. и L. vadoni Voss, 1966, syn.n. к L. dentipennis (Gyllenhal, 1839). Australetobius incostans (Lea, 1910), placem.n. и A. rubricollis (Voss, 1922), placem.n. перемещены из рода Auletobius Desbrochers des Loges, 1869 в род Australetobius Legalov, 2007; подрод Mandelschtamius Legalov, 2003, placem.n. из Kuschelanthus Alonso-Zarazaga & Lyal, 1999, stat.n. в Pseudominurus Voss, 1956; Yunnanuletes perturbatus (Voss, 1930), placem.n. из рода Pseudomesauletes Legalov, 2001 в род Yunnanuletes Legalov, 2007; Pseudomesauletes podocarpi (Voss, 1933), placem.n. из рода Kuschelanthus Alonso-Zarazaga & Lyal, 1999 в род Pseudomesauletes Legalov, 2001; Pseudomesauletes fuscofasciatus (Voss, 1933), placem.n. и Pseudomesauletes maculatus (Voss, 1933), placem.n. из подрода Rubrauletes Legalov, 2003 в подрод Fascauletes Legalov, 2007; Pseudodicranognathus fuliginosus (Voss, 1933), placem.n. из рода Pseudomesauletes Legalov, 2001 в род Pseudodicranognathus Legalov, 2001; Caenorhinus rufiventris (Voss, 1921), placem.n. из рода Neoarodepus Legalov, 2003 в род Caenorhinus C.G. Thomson, 1859; Cartorhynchites breviusculus (Voss, 1939), placem.n. из подрода Hyperinvolvulus Legalov, 2003 в подрод Cartorhynchoides Legalov, 2003; Cartorhynchites crassiusculus (Voss, 1938), placem.n. из подрода Cartorhynchoides Legalov, 2003 в подрод Hyperinvolvulus Legalov, 2003; Metarhynchites longulus (Gyllenhal, 1833), placem.n. из Rhynchitidae ins. sedis в род Metarhynchites Voss, 1923; Afrorhynchites villosus (Boheman, 1845), placem.n. из рода Pararhynchites Legalov, 2003 в род Afrorhynchites Legalov, 2003; Heterorhynchites pruinosus (Voss, 1938), placem.n. из подрода Eosawadaia Legalov, 2004 в подрод Eosawadaia s. str; Omolabus centomyrciae (Voss, 1925), placem.n. из подрода Paralabus Legalov, 2004 в подрод Pseudomolabus Legalov, 2004; Apleurolabus evanescens (Voss, 1928), placem.n. из рода Pleurolabus Jekel, 1860 в род Apleurolabus Legalov, 2007; Metriotracheloides regularis (Ter-Minassian, 1986), placem.n. из рода Trachelophoridius Voss, 1929 в род Metriotracheloides Legalov, 2008. Изменен статус Auletobius ebenus Hustache, 1955, stat.n., Clinorhynchites rufofemoratus (Voss, 1938), stat.n. и Trachelismus prolixus (Voss, 1929) (из форм и вариаций до видов), а также Kuschelanthus Alonso-Zarazaga & Lyal, 1999, stat.n. и Paralleuscelus Legalov, 2004, stat.n. (из родов до подродов). Hamiltoniauletes trifasciatus (Suffrian, 1870) впервые указан для фауны Доминиканской республики, Pseudomesauletes ueleanus (Voss, 1939) для фауны Камеруна, P. gestroi (Faust, 1894) для фауны Лаоса, Eumetopon flavomaculatus (Voss, 1922) для фауны провинции Чжэцзян, Clinorhynchidius flexirostris (Voss, 1938) для фауны Габона, Heterorhynchites wahnesi (Hartmann, 1899) для фауны Сабаха, H. elysius (Pascoe, 1875) для фаун Малайзии, Калимантана и Явы, H. subdentatus (Voss, 1938) для фауны Калимантана, Suniops gorochovi Legalov, 2003 для фауны Таиланда, Kobusynaptops verrucosus Legalov & X. Zhang, 2007 для фауны провинции Сычуань, Riedeliops zolotarenkoi Legalov, 2003 для фауны Таиланда, Parasynaptopsis chinensis (Voss, 1922) для фауны провинции Аньхой, Parasynaptopsis lespedezae koreanus (Voss, 1924) для фауны провинции Хубэй и P. moanus Legalov, 2003 для фауны провинции Юньнань. Установлены 30 новых комбинаций названий.

INTRODUCTION

The families Rhynchitidae and Attelabidae have about 3000 species in sum, being small groups within the superfamily Curculionoidea. Both families are monophyletic [Legalov, 2007]. Rhynchitidae emerged in early Cretaceous [Legalov, 2009b, 2009c], Attelabidae are known from Paleogene [Legalov, 2007]. Family Rhynchitidae consists of species rolling leaves into tubes for the larvae development as well as of species using other substrata for oviposition. All representatives of Attelabidae roll leaves into tubes [Legalov, 2004].

Taxa within both families are usually difficult to distinguish. The armament of endophallus could be good diagnostic character for many genera. During 2005-2009 the author studied numerous materials on the leaf-rolling weevils from different museums, including type specimens. Results of this work are presented in this paper.

MATERIAL AND METHODS

Types and specimens are kept in the following collections and museums: ACD - A. Allen Collection (USA: Boise); APB - A. Podlussany Collection (Hungary: Budapest); BMNH - The Natural History Museum (United Kingdom: London); CBN - R. Borovec Collection (Czech

Republic: Nechanice); CJPM - J. Pelletier Collection (France: Monnaie); CKJU - P. Kresl Collection (Czech Republic: Janovice nad Uhlavou); DEI - Deutsches Entomologisches Institut (Germany: Muncheberg);

HNHM - Hungarian Natural History Museum (Hungary: Budapest); ISNB - Institut Royal des Sciences Naturelles de Belgique (Belgium: Brussels); MCSN - Museo Civico di Storia Naturale “Giacomo Doria” (Italy: Genova); MCZ - Harvard University, Museum of Comparative Zoology (USA: Cambridge); MNHN - Museum National d’Histoire Naturelle (France: Paris); MMUE - The Manchester Museum, The University of Manchester (UK: Manchester); MRAC - Musee Royal de l’Afrique Centrale (Belgium: Tervuren); MZLU - Lund University (Sweden: Lund); NHRS - Naturhistoriska riksmuseet (Swedish Museum of Natural History) (Sweden; Stockholm); NME

- Naturkundemuseum Erfurt (Germany: Erfurt); NMKE

- National Museum of Kenya (Kenya: Nairobi); NMPC

- National Museum of Natural History (Czech Republic: Prague); RDP - R. Dunda Collection (Czech Republic: Prague); SMTD - Staatliches Museum fur Tierkunde (Germany: Dresden); SMWN - National Museum of Namibia (Namibia: Windhoek); SZMN - Siberian Zoological Museum, Institute of Animal Systematics and Ecology (Russia: Novosibirsk); TAUI - Tel Aviv University (Israel: Tel Aviv); USNM - National Museum of Natural

History, [formerly, United States National Museum] (USA: Washington); VRP - V Ryjacek Collection (Czech Republic: Prague); VSM - V Savitsky Collection (Russia: Moscow); ZFMK - Zoologische Forschungsinstitut und Museum “Alexander Koenig” (Germany: Bonn); ZIN

- Zoological Institute of Russian Academy of Sciences (Russia: St. Petersburg); ZMAN - Zoologisch Museum, Instituut voor Taxonomische Zoologie, Universiteit van Amsterdam (Netherlands: Amsterdam); ZMHB - Museum fur Naturkunde der Humboldt-Universitat (Germany: Berlin); ZMUC - Zoological Museum, University of Copenhagen (Denmark: Copenhagen); ZMUM -Zoomuseum of Moscow State University (Russia: Moscow).

RESULTS

Family Rhynchitidae Gistel, 1848 Supertribe Rhinocartitae Voss, 1931 Tribe Rhinocartini Voss, 1931 Genus Rhinocartus Voss, 1922 Rhinocartus tessmanni Voss, 1922 (col. pl. I: a, IX: 1) Rhinocartus tessmanni Voss, 1922a: 18 Rhinocartus dahli Voss, 1956b: 1139 Remarks. Specimen studied: a male from the collection MRAC with labels “Musee du Congo, Uele: Bambesa, 20.09.1933, J. Leroy”, “R. Det. T. 3887”, “Rhinocartus tessmanni m., Det. E. Voss”.

Distribution. Cameroon, Guinea, Zaire.

Tribe Sanyrevilleini Legalov, 2003 Subtribe Parauletanina Legalov, 2007 Genus Australetobius Legalov, 2007

Australetobius incostans (Lea, 1910), comb.n., placem.n. (col. pl. I: b-d, IX: 2-3)

Auletes incostans Lea, 1910: 41

Remarks. The lectotype is designated by the author - a male from the collection DEI with labels “Tasmania”, “Coll. Hacker”, “Syntypus”, “Auletes incostans Lea, Tasmania, Cotype”, “Auletobius incostans Lea i. l.?”, “coll. DEI Muncheberg”, “Auletobius incostans Lea”, “Lectotype Auletes incostans Lea, 1910, A. Legalov design. 2009”. Previously, this species has been wrongly placed in the genus Auletes from the tribe Auletini.

Distribution. Tasmania.

Australetobius rubricollis (Voss, 1922), comb.n., placem.n. (col. pl. II: a-c)

Auletobius rubricollis Voss, 1922a: 32 Remarks. The holotype was studied: a female from the collection DEI with labels “Australien”, “Holotypus”, “Auletobius rubricollis n. sp.”, “1029”, “coll. DEI Muncheberg”, “Auletobius rubricollis Voss”, “Holotype Auletobius rubricollis Voss, 1922, A. Legalov det. 2009”. Previously, this species has been wrongly placed in the genus Auletobius Desbrochers des Loges, 1869 of the tribe Auletini.

Distribution. Australia.

Supertribe Rhynchititae Gistel, 1848 Tribe Auletini Desbrochers des Loges, 1908 Subtribe Auletobiina Legalov, 2001

Genus Auletobius Desbrochers des Loges, 1869 Subgenus Auletobius s. str.

Auletobius (Auletobius) aeneus Voss, 1922 (col. pl. I: e-g)

Auletobius aeneus Voss, 1922a: 49 Remarks. The holotype was studied: a female from the collection DEI with labels “Australien”, “1035”, “Auletes densus Lea”, “Holotypus”, “Voss det.”, “Auletobius aeneus m., nov. spec.”, “coll. DEI Muncheberg”, “Auletobius aeneus Voss”, “Holotype Auletobius aeneus Voss, 1922, A. Legalov det. 2009”.

Distribution. Australia.

Auletobius (Auletobius) albipilosus Voss, 1922 (col. pl. I: h-i)

Auletobius albipilosus Voss, 1922a: 33 Remarks. The holotype was studied: a female from the collection DEI with labels “Australien”, “Lea has not”, “1419”, “Holotypus”, “Voss det.”, “Auletobius albipilosus m., nov. spec.”, “coll. DEI Muncheberg”, “Auletobius albipilosus Voss”, “Holotype Auletobius albipilosus Voss, 1922, A. Legalov det. 2009”.

Distribution. Australia.

Auletobius (Auletobius) imitator (Lea, 1910) (col. pl. II: d-f, IX: 4-5)

Auletes imitator Lea, 1910: 41

Remarks. The lectotype is designated by the author - a female from the collection DEI with labels “Hobart Tas: Lea”, “Coll. Hacker”, “1419”, “Voss det.”, “Syntypus”, “Auletes imitator Lea, Tasmania, Cotype”, “Auletobius imitator Lea i. l.?”, “coll. DEI Muncheberg”, “Auletobius imitator Lea”, “Lectotype Auletes imitator Lea, 1910, A. Legalov det. 2009”. The specimens studied by the author: a female from the collection DEI with labels “Australien”, “1030”, “Auletes imitator Lea”, “Voss det.”, “coll. DEI Muncheberg”; a female from the collection DEI with labels “Australien”, “1031”, “Voss det.”, “coll. DEI Muncheberg”; a male from the collection DEI with labels “Australien”, “1032”, “Voss det.”, “coll. DEI Muncheberg”.

The record [Voss, 1922a] of this species from New Caledonia belongs to Auletobius montrouzieri Voss, 1942. Distribution. Australia, Tasmania.

Auletobius (Auletobius) laterirostris (Lea, 1926) (col. pl. I: ig-h, IX: 6-7)

Auletes laterirostris Lea, 1926: 35 Remarks. The lectotype is designated by the author - a male from the collection NHRS with labels “Cedar creek”, “Queensl., Mioberg”, “Cotype”, “Auletes laterirostris Lea, Co-type”, “7210 E91”, “Lectotype Auletes laterirostris Lea, 1926, A. Legalov det. 2009”. Paralectotypes: a female with labels “Malanda”, “Queensl., Mioberg”, “Cotype”, “7211 E91”, “Paralectotype Auletes laterirostris Lea, 1926, A. Legalov det. 2009” and a female with labels “Blackal Range”, “Queensl., Mioberg”, “sept.”, “7212 E91” “Paralectotype Auletes laterirostris Lea, 1926, A. Legalov det. 2009”. Specimen with labels “Kimberley district”, “N.V Austr. Mioberg”, “Cotype”, “Paratypus”, “7213 E91”, “laterirostris Lea” belongs to family Curculionidae. Distribution. Australia.

Auletobius (Auletobius) melanocephalus (Erichson, 1842) (col. pl. II: l, o)

Rhynchites melanocephalus Erichson, 1842: 185 Material. 1 male (HNHM), Australia, Queensland,

09.1980, G. Hangay; 2 females (ZMUM), Australia, S of Canberra, Tidbinbilla, 27-28.II.1997, D. Shcherbakov. Remarks. The lectotype is designated by the author

- a female from the collection ZMHB with labels “34387”, “Type”, “melanocephalus Er., Vadem Schaya”, “HOLOTYPUS Rhynchites melanocephalus (Erichson, 1842), labelled by MNHUB 2009” “Lectotype Rhynchites melanocephalus Erichson, 1842, A. Legalov design. 2009”. The record [Voss, 1922, 1934; Legalov, 2003a, 2007] of this species from Tasmania belongs to other species, probably to Auletobius melanocephalus.

Distribution. Australia.

Auletobius (Auletobius) montrouzieri Voss, 1942 (col. pl. II: i-k, m, IX: 8-9)

Auletobius montrouzieri Voss, 1942a: 61 Remarks. The lectotype is designated by the author - a male from the collection DEI with labels “New Caledonien”, “Syntypus”, “Coll. Kraatz”, “Auletobius montrouzieri n. sp., det. E. Voss”, “Eugnamptus Montrouzieri Faust, N. Caled., Type”, “coll. DEI Muncheberg”, “Lectotype male Auletobius montrouzieri Voss, 1942, Kuschel 2005”, “Auletobius montrouzieri Voss”, “Lectotype Auletobius montrouzieri Voss, 1942, A. Legalov det. 2009”. Paralectotype - a female from the collection DEI with labels “New Caledonien”, “Coll. Kraatz”, “Voss det.”, “Syntypus”, “coll. DEI Muncheberg”, “Paralectotype Auletobius montrouzieri Voss, 1942, Kuschel 2005”, “Paralectotype Auletobius montrouzieri Voss, 1942, A. Legalov det. 2009”. I studied following specimens: a female

- “New Caledonia: Noumea: Anse Vita, 30 m, 6.08.1979”, “G.A. Samuelson”, “Auletobius montrouzieri Voss, 1942, Kuschel 2004” and a male - “N Caledon, Mortong, Marscal”, “Coll. Haag”, “Voss det.”, “Auletobius trossulus Schauf. i. l.”, “Auletobius imitator Lea, Voss 1922, 1933”, “Auletobius montrouzieri Voss, 1942, Kuschel 2005”. Distribution. New Caledonia.

Key to species of the genus Auletobius from Madagascar

1. Body larger (2.0-2.2 mm)................................2

- body smaller (1.3-1.6 mm)...............................3

2. Frons more densely punctate. Pronotum and elytra with

smaller points..................................A. fausti

- frons more sparsely punctate. Pronotum and elytra with

larger points...................................A. ebenus

3. Body paler. Frons with smaller points. Rostrum shorter.

Sides of pronotum stronger rounded. Armament of the endophallus (col. pl. IX: 10-11)........ ... A. pygmaeus

- body darker. Frons with larger points. Rostrum longer.

Sides of pronotum mildly rounded. Armament of the endophallus (col. pl. IX: 13).....................A. iviei

Auletobius (Auletobius) fausti Voss, 1922

Auletobius fausti Voss, 1922a: 32 Remarks. The holotype was studied: a female from the collection SMTD with labels “Madagascar, Sikora”, “Coll. J. Faust, Ankauf 1900”, “Staatl. Museum fur Tierkunde, Dresden”, “Typus”, “Metopon Fausti n. sp.”, “Holotype Auletobius fausti Voss, A. Legalov design. 2005”. Distribution. Madagascar.

Auletobius (Auletobius) ebenus Hustache, 1955, stat.n.

(col. pl. I: j, IX: 12)

Auletobiuspygmaeus v. ebenus Hustache, 1955: 191 Remarks. The holotype was studied: a male from the collection MNHN with labels “Nandihizina”, “Madagascar, Maroantesetra, 12.38, Vadon !”, “Type”, “pygmaeus v. ebenus m.”, “Museum Paris, 1919, Col. A. Hustache”, “Holotype Auletobius pygmaeus v ebenus Hustache, 1955, A. Legalov det. 2009”.

Distribution. Madagascar.

Auletobius (Auletobius) pygmaeus Hustache, 1955 (col. pl. II: n, p, IX: 10-11)

Auletobius pygmaeus Hustache, 1955: 190 Remarks. The lectotype is designated by the author - a male from the collection MNHN with labels “Nandihizina”, “Madagascar, Maroantesetra, 12.38, Vadon !”, “male”, “Type”, “Auletobius (Parauletes Voss) pygmaeus m.”, “Museum Paris, 1919, Col. A. Hustache”, “Lectotype Auletobius pygmaeus Hustache, 1955, A. Legalov design. 2009”.

Distribution. Madagascar.

Auletobius iviei Legalov, sp.n. (col. pl. III: a-b, IX: 13) Material. Holotype - male (SZMN), “Madagascar, 54 km E Antananarivo, N. Rt 2, 6 km E Manjakantriana, sweeping, 12.11.1994, M.A. Ivie & D.A. Pollock”. Description. Body brown. Prolegs paler. Body with almost adpressed, pale, short setae. Rostrum long, 8.33 times longer than wide, 1.56 times longer than pronotum, very weakly curved, widened to apex, very sparsely and finely punctate. Antennae located near the rostrum basis. Eyes not large, strongly convex. Frons wide, convex, densely punctate. Temples straight, short. Antennae long, reaching apical margin of pronotum. Scapus and 1st segment of funicle oval. 2nd-6th segments elongated-oval, narrower. 2nd segment longer than 1st segment. 3rd segment shorter than 2nd segment. 4th segment of almost equal length to 3rd segment. 5th and 6th segments of almost equal length, shorter. 7th segment almost trapezoid, shorter than 6th segment. Clava wide, almost compact, pointed. 1st and 2nd segments almost square. 3rd segment tear-shaped, longer than 2nd segment. Pronotum almost campaniform, of almost equal length and width, with weakly rounded sides, slightly narrowed to basis and apex. Disk convex, finely and densely punctate. Greatest width before the middle. Scutellum trapezoid. Elytra almost rectangular, elongated, 1.33 times longer than wide. Greatest width behind the middle. Humeri weakly smoothed. Striae reduced. Points small and sparse. Intervals weakly convex. Thorax finely and sparsely punctate. Metepisternum very narrow. Abdomen convex. 1st and 2nd ventrites wide. 2nd ventrite hardly wider than 1st ventrite. 3rd and 4th ventrites narrower, narrower than 2nd ventrite. 5th ventrite narrow, narrower than 4th ventrite. Pygidium convex, punctate. Legs long. Femora widened. Tibiae almost straight. Tarsi long. 1st segment elongated-triangular. 2nd segment wide triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 1.4 mm. Diagnosis. See key to species of the genus Auletobius from Madagascar.

Etymology. The new species is named in honour of M.A. Ivie.

Distribution. Madagascar.

Auletobius (Auletobius) calvus (Sharp, 1889)

Auletes calvus Sharp, 1889a: 72 Auletobius irkutensis ssp. japonicus Voss, 1922a: 31 Remarks. The specimens studied by the author - a male with labels “Japan, Hillef’, “male”, “Coll. Schilsky”, “puberulus Faust” and a female with labels “Japan, Hiller”, “female”, “Coll. Schilsky” from the collection ZMHB. Distribution. Japan.

Auletobius baishuiensis Legalov, sp.n.

Material. Holotype - female (NMPC), China, Yunnan, Lijiang, Yulongshan, Bai Shui, 2900-3300 m, 7-18.07.1994,

C. Holzschuh.

Description. Female. Body black, with thin pale semierect setae. Elytra (without spot near scutellum, suture and margins) red-brown. Tibiae, tarsi, scapus and funicle of antennae brownish. Rostrum long, 7.2 times longer than wide, 1.44 times longer than pronotum, almost straight, weakly widened to apex, almost smooth. Antennae located on the basis of rostrum. Eyes not large, strongly convex. Frons wide, strongly convex, densely and largely punctate. Temples straight, weakly elongated. Antennae long, reaching humeri. Scapus and 1st segment of funicle elongated-oval. 2nd - 4th segments elongated, narrow. 2nd segment longer than 1st segment. 3rd segment shorter than 2nd segment. 4th segment shorter than 3rd segment. 5th and 6th segments oval, approximately equal in length. 5th segment much shorter than 4th segment. 7th segment short, oval. Clava wide, almost compact, pointed, much shorter than funicle. 1st and 2nd segments short, weakly transversal. 3rd segment tear-shaped, slightly shorter than the previous segments taken together. Pronotum almost campaniform, 1.08 times wider than long, with weakly rounded sides, weakly narrowed to the basis and apex. Disk convex, largely and densely punctate. Greatest width on the middle. Scutellum trapezoid. Elytra almost obovate, elongated, 1.33 times longer than wide. Greatest width behind the middle. Humeri weakly smoothed. Striae reduced. Points small. Intervals narrow, flat. Thorax finely and sparsely punctate. Metepisternum narrow. Abdomen convex, finely rugosely punctate. 1st and 2nd ventrites wide, approximately equal in length. 3rd and 4th ventrites narrower, narrower than 2nd ventrite. 5th ventrite narrow, narrower than 4th ventrite. Pygidium convex, punctate. Legs long. Femora widened. Tibiae almost straight, weakly widened to apex. Protibiae narrow and long. Tarsi long, weakly flattened. 1st segment elongated. 2nd segment wide triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 2.7 mm. Diagnosis. This new species is close to Auletobius rufus Legalov, 2007 but differs with narrower body, shorter rostrum, weakly rounded sides of pronotum, dark suture of elytra and black legs.

Etymology. The name is derived from the location “Bai Shui” - “baishuiensis”.

Distribution. China (Yunnan).

Subtribe Mandelschtamiina Legalov, 2003 Key to genera of the subtribe Mandelschtamiina

1. Antennae located in the middle or in the first third of rostrum, but not near the basis. Africa, Madagascar.......

............................Pseudominurus Voss, 1956

- antennae located almost at the basis of rostrum. Frons

weakly convex........................................2

2. Elytra with distinct punctate striae in the first third. Body with relatively sparse thin semierect setae. Eyes strongly

convex. Tarsi elongated. Rwanda........................

............................Pilosauletes Legalov, 2009

- elytra without punctate striae. Body with dense long appressed pale setae. Eyes weakly convex. Tarsi not elongated. Cabo Verde ... Caboverdeletus Legalov, 2007

Genus Pseudominurus Voss, 1956

Auletobius subgenus Pseudominurus Voss, 1956a: 601 Type species: Auletobius discedens Voss, 1956

Key to subgenera of genus Pseudominurus

1. Antennae located before the basis of rostrum. Tanzania .... ........................................ Tanzanominurus

- antennae located further from the basis of rostrum..2

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2. Rostrum short. Eyes strongly convex. Antennae located

almost in the middle of rostrum or in the first third closer to the middle. Elytra with punctate striae, usually distinct in the first third. Africa........Pseudominurus s. str.

- rostrum long. Eyes often weaker convex . 3

3. 1st segment of protarsi not flattened and widened in

females. Africa......................Mandelschtamius

- 1st segment of protarsi flattened and widened in females.

Madagascar..............................Kuschelanthus

Subgenus Pseudominurus s. str. Pseudominurus (Pseudominurus) hortulanus (Faust, 1899) (col. pl. III: c-d, IX: 14)

Minurus hortulanus Faust, 1899a: 343 Auletobius montanus Voss, 1922a: 36, syn.n.

Remarks. The holotype was studied: a male from the collection ZMHB with labels “A. d. Sammlung Dr. Chr. Schroder's, Parek, 2000 m” (view from above) “15/1” (view from below), “Auletobius montanus m.”, “HOLOTYPUS Auletobius montanus Voss, 1922, labelled by MNHUB 2009”, “Holotype Auletobius montanus Voss, 1922, A. Legalov det. 2009”.

Study of the types showed that Auletobius montanus Voss, 1922, syn.n. is synonym to Pseudominurus hortulanus (Faust, 1899).

Subgenus Kuschelanthus Alonso-Zarazaga & Lyal,

1999, stat.n.

Kuschelanthus Alonso-Zarazaga & Lyal, 1999: 42 Type species: Salacus perrieri Fairnaire, 1899 Salacus Fairmaire, 1899: 545 [non Gistel, 1848]; type species: Salacus perrieri Fairnaire, 1899 Remarks. Pseudominurus (Kuschelanthus) hustachei (Voss, 1932), comb.n., P (K.) madagasus (Legalov, 2007), comb.n., P (K.) perrieri (Fairmaire, 1899), comb.n. and P (K.) vadoni (Hustache, 1955), comb.n. are assigned to this subgenus.

Pseudominurus (Kuschelanthus) madagasus (Legalov, 2007), comb.n.

Kuschelanthus madagasus Legalov, 2007: 48 Material. 1 female (ISNB), “Madagascar, 1898, Ex.

Oberthur”; female (MRAC), “Madagascar: Fempanambo,

07.1960, J. Vadon”, “Salacus perrieri Fairm., Ferragu det.”.

Distribution. Madagascar.

Subgenus Mandelschtamius Legalov, 2003, placem.n.

Mandelschtamius Legalov, 2003a: 107 Type species: Mandelschtamius turneroides Legalov, 2003 Remarks. Pseudominurus (Mandelschtamius) nigrolimbatus (Peringuey, 1888), comb.n., P. (M.)

tanganyikus (Legalov, 2003), comb.n., P. (M.) turneri (Voss, 1933), comb.n. and P. (M.) turneroides (Legalov, 2003), comb.n. are assigned to this subgenus.

Pseudominurus (Mandelschtamius) tanganyikus (Legalov, 2003), comb.n. (col. pl. III: e-f)

Mandelschtamius tanganyikus Legalov, 2003a: 107 The specimen studied by the author - a female from the collection MRAC with labels “Coll. Mus. Congo, Tanganyika Terr.: Mt. Meru, Olkokola, versant N. O., 2800 m, 24.06. - 1.08.57”, “Mission Zoolog. I.R.S.A.C., en Afrique orientale (P. Basilewsky et N. Leleup)”, “Auletobius hortulanus Fst., female, E. Voss det., 1961”. Distribution. Tanzania.

Subgenus Tanzanominurus Legalov, subgen.n.

Type species: Kuschelanthus tangensis Legalov, 2007 Description. Body yellowish-brown or brown. Rostrum and clava usually darker. Head and rostrum usually red-brown. Scapus, funicle, abdomen and legs usually yellowish-brown. Body with dense pale semierect setae. Rostrum long, weakly curved, slightly widened to apex, punctate. Antennae located before the rostrum basis. Eyes large, strongly convex. Frons wide, convex, punctate. Temples short. Antennae long, reaching middle of pronotum or the first margin of elytra. Pronotum almost campaniform. Sides weakly rounded, narrowed to the basis and apex. Disk flattened, punctate. Greatest width in the middle or in basal third of pronotum. Scutellum small, triangular. Elytra elongated. Greatest width in or near the middle. Humeri smoothed. Punctate striae almost distinct. Points in striae large, dense. Intervals weakly convex, punctate. Metepisternum narrow. Abdomen convex, punctate. Legs long. Femora widened. Tibiae narrow. Tarsi long. 1st segment of tarsi elongated-triangular. 2nd segment triangular. 3rd segment bilobed. Clausal segment elongated. Claws with teeth. Length of body: 1.9-3.1 mm. Diagnosis. See key to subgenera of genus Pseudominurus. Remarks. Pseudominurus (Tanzanominurus) amaniensis (Legalov, 2007), comb.n. and P. (T.) tangensis (Legalov, 2007), comb.n. from Tanzania are assigned to this subgenus. Etymology. The name is formed from the words “Tanzania” and “Minurus”

Subtribe Pseudauletina Voss, 1933 Genus Pseudauletes Voss, 1922 Subgenus Eopseudauletes Legalov, 2007 Pseudauletes (Eopseudauletes) luceus (Gyllenhal, 1839) (col. pl. III: g-h, IX: 15)

Rhynchites luceus Gyllenhal, 1839: 322

Material. 1 ex. (SMTD), “Brasilien, Dohrn”, “Coll. J.

Faust, Ankauf 1900”, “Staatl. Museum fur Tierkunde, Dresden”; 1 ex. (SMTD), “Petropolis, Dr. Ohaus”, “24, 1906”, “Staatl. Museum fur Tierkunde, Dresden”. Remarks. The lectotype is designated by the author - a female from the collection NHRS with labels “Typus”, “Rh. atropurpureus Schh., Brazil, Fald.”, “Rh. luceus”, “411, 65”, “Lectotype Rhynchites luceus Gyllenhal, 1839, A. Legalov design. 2009”. One more specimen studied - a male with labels “Rh. luceus, Brazil, Germar”, “411, 65”. Distribution. Brazil.

Subtribe Pseudomesauletina Legalov, 2003 Genus Aletinus Desbrochers des Loges, 1908 Subgenus Aletinus s. str.

Aletinus (Aletinus) maculipennis (Jacquelin du Val, 1854) (col. pl. III: i)

Auletes maculipennis Jacquelin du Val, 1854: 8 Auletobius maculipennis var. concolor Desbrochers des Loges, 1869: 406

Auletobius maculipennis var. lepigrei Hoffmann, 1958: 1732

Material. 1 ex. (RDP), Morocco, SW Tiznit Oued, Massa, 8.05.2003, M. Snizek; 1 ex. (RDP), Tunis, Kairuen,

11.06.1982, A. Olexa; 1 ex. (RDP), Morocco, Asm env., 50 rm S of Merksech, 1150 m, 26.06.1990, Z. Kejval; 1 ex. (RDP), Africa sept., Algeria, Hamman, Salihine, 2425.05.1971, A. Hoffer & J. Horak; 1 ex. (RDP), Africa sept., Algeria, Gr. Kabylia, Tiziozuzou, 15.06.1971, A. Hoffer & J. Horak; 1 ex. (HNHM), “Ins. Elba, 1908, Paganetti”; 2 ex. (MZLU), Tunisia, Sousse, 17-29.05.1969, Thure Palm; 1 ex. (MZLU), Sardinien, S. Glorgie, 19-27.06.1967, Thure Palm; 4 ex. (MZLU), Sardinien, Alghero, 1528.06.1967, Thure Palm; 15 ex. (MZLU), Tun., Gabes, 30.05.-5.06.1969, Thure Palm; 9 ex. (NHRS), Tunisia, “Tu. Sousse, 20-26.05.1969, T-E Leiler”; 16 ex. (NHRS), Tunisia, “Tu. Gabes, 31.05.1969, T-E Leiler”.

Remarks. The lectotype of A. m. var. concolor is designated by the author - a female from the collection HNHM with labels “Algir, Edough”, “Paratypus 1869 Auletobius maculipennis Jacq. var. concolor Desbrochers des Loges”, “v. concolor m.”, “Coll. Reitter”, “Lectotype Auletobius maculipennis var. concolor Desbrochers des Loges, 1969, A. Legalov design. 2009”.

Distribution. Algeria, Morocco, France, Italy.

Genus Hamiltoniauletes Legalov, 2001 Hamiltoniauletes subseriepunctatus (Voss, 1922)

Auletobius subseriepunctatus Voss, 1922a: 34 Remarks. The holotype was studied: a female from the collection ZMHB with labels “Chico”, “Mexico, J. Flohr

G.”, “Auletobius subseriepunctatus m.”, “HOLOTYPUS Auletobius subseriepunctatus Voss, 1922, labelled by MNHUB 2009”, “Holotype Auletobius subseriepunctatus Voss, 1922, A. Legalov det. 2009”.

Distribution. Mexico.

Hamiltoniauletes trifasciatus (Suffrian, 1870) (col. pl.

III: j-k)

Rhynchites trifasciatus Suffrian, 1870: 229 Auletobius cubanus Voss, 1922a: 34 Remarks. The lectotype of Auletobius cubanus is designated by the author - a female (left) from the collection

ZMHB with labels “Cuba”, “Cuba”, “255”, “Coll. L.W. Schufuss”, “Auletobius cubanus m.”, “SYNTYPUS Auletobius cubanus Voss, 1922, labelled by MNHUB 2009”, “Lectotype Auletobius cubanus Voss, 1922, A. Legalov design. 2009”. Paralectotype - a female (right). Specimens with label “Republica Dominicana: La Altagrada, Boca de Yuma, P.N. del Este (18.21.35 N, 68.37.10 W), 80 m, 26.03.2002, canopy fogging, coll. B. Farell & K. Guerrero” from the collection MCZ (http:// insects.oeb.harvard.edu) belong to this species. Possibly, Auletobius sp. from Andros [Turnbow, Thomas, 2008] should be assigned to this species. This species is reported for the first time for the fauna of Dominican Republic. Distribution. Cuba, Dominican Republic, Bahamas: New Providence [Turnbow, Thomas, 2008; as Auletobius cubanus Voss].

Genus Yunnanuletes Legalov, 2007

Yunnanuletes perturbatus (Voss, 1930), comb.n., placem.n.

Auletobius perturbatus Voss, 1930: 65 Remarks. Previously, this species has been wrongly placed in the subgenus Rubrauletes Legalov, 2003 of the genus Pseudomesauletes Legalov, 2001.

Distribution. China (Sichuan).

Genus Tanzanauletes Legalov, gen.n. (col. pl. IV: a-b, IX: 16) Type species: Auletobius hustachei Dalla Torre & Voss,

1937

Description. Body black-brown. Antennae and legs brown. Body with long, erect, thin setae. Rostrum long, 5.63 times longer than wide, 1.40 times longer than pronotum, weakly curved, widened to the apex, densely punctate. Antennae located in the middle of rostrum. Eyes large, strongly convex. Frons wide, convex, densely punctate. Temples short. Antennae long, reaching beyond the front of pronotum. Scapus and 1st segment of funicle oval. 2nd-3rd segments long-oval, narrower. 2nd segment longer than 1st segment. 4th segment shorter and wider than 3rd segment. 5th and 6th segments trapezoid, shorter. 6th segment wider than 5th segment. 7th segment rounded, shorter than 6th segment. Clava wide, almost compact, pointed, shorter than funicle. 1st and 2nd segments transversal. 3rd segment stilliform, slightly shorter than previous segments. Pronotum almost campaniform, of equal length and width, weakly narrowed to basis and apex. Disk convex, finely and densely punctate. Greatest width in the middle. Scutellum trapezoid. Elytra almost rectangular, elongated, 1.39 times longer than wide. Greatest width near the middle. Humeri weakly smoothed. Striae reduced. Points large and deep. Intervals convex. Apex of elytra in males with sex patches. Thorax finely and sparsely punctate. Metepisternum narrow. Abdomen convex. 1st and 2nd ventrites wide; 2nd ventrite slightly wider than 1st. 3rd and 4th ventrites narrower than 2nd segment. 5th ventrite narrow, narrower than 4th segment. Pygidium convex, punctate. Legs long. Femora widened. Tibiae almost straight, slightly widened to apex. Tarsi long. 1st tarsal segment long-triangular. 2nd segment wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 2.9 mm.

Diagnosis. The new genus is close to the genus Hustacheletes Legalov, 2007 but differs by the protarsi not elongated and not flattened, body black, elytra with the irregular rows of points and antennae located near the middle of rostrum.

Etymology. The name is formed from the words “Tanzania” and “Auletes”.

Tanzanauletes hustachei (Dalla Torre & Voss, 1937), comb.n. (col. pl. IV: a-b, IX: 16)

Auletobius hustachei Dalla Torre & Voss, 1937: 26 [RN] Auletobius orientalis Hustache, 1929: 501 [non Lea, 1926] Remarks. The lectotype is designated by the author - a male from the collection MNHN with labels “Afrique or. allemande KILIMANDJARO, versant Sud-Est, Alluard & Jeannel”, “Type”, “Zone Inferieure, Neu-Moschi, 800 m, Avrill 1912 St. 72”, “Auletobius orientalis Hust.”, “Museum Paris”, “Lectotype Auletobius orientalis Hustache, 1929, A. Legalov design. 2009”.

Distribution. Tanzania.

Genus Pseudomesauletes Legalov, 2001 Key to subgenera of the genus Pseudomesauletes

1. Elytra obovate. Rows of points on elytra almost regular.

Southeast Asia....................Ovauletes Legalov, 2007

- elytra almost rectangular, usually with greatest width in the middle, not ovoid. Elytra without rows of points ... 2

2. Body metallic green or dark blue, almost naked. Western,

Central Africa.................Metallauletes Legalov, 2007

- body black, brown, and red, with setae..................3

3. Body black. Setae on elytra with metallic sheen.........4

- body red. Setae on elytra without metallic sheen........6

4. Armament of endophallus reduced. Antennae located behind middle of rostrum in males, and in middle of

rostrum in females. Eyes weaker convex. Africa............

............................Afromesauletes Legalov, 2003

- armament of endophallus distinct. Antennae located in

middle or before middle in males, and before middle of rostrum in females. Eyes stronger convex.................5

5. Antennae located in middle of rostrum in males, and

before middle in females. East, Southeast, Southern Asia, Northern America ............. Pseudomesauletes s. str.

- antennae located before middle of rostrum in males.

Columbia.........................Colombletes Legalov, 2007

6. Rostrum usually longer and thinner. Elytra usual with

almost regular rows of points...........................7

- rostrum usually shorter and thicker. Elytra without almost

regular rows of points..................................8

7. Top with pattern of pale setae. Body externally similar

to Hamiltoniauletes. Disk of pronotum near basis granulated. Apex of elytra weakly pressed (almost as at genus Pseudodicranognathus), yellow. Southeast Asia .... ...............................Fascauletes Legalov, 2007

- top without pattern of pale setae. Disk of pronotum

near the basis punctate. Apex of elytra not pressed. Madagascar...............Madauletes Legalov, subgen.n.

8. Basal margin of elytra much wider than pronotum. Eyes

weaker convex. Rostrum longer. Yunnan...................

......................Rubrauletoides Legalov, subgen.n.

- basal margin of elytra slightly wider than pronotum. Eyes

strongly convex. Rostrum shorter........................9

9. Body red. Rostrum, antennae, sides and apex of elytra, legs black-brown. Humeri smoothed. Body narrower and

long. Southeast Asia, Java.............................

...........................Faustiauletes Legalov, 2003

- body red or red-brown; rostrum, antennae, legs and thorax often dark, but sides and apex of elytra always pale. Humeri weaker smoothed. Body wider. East, Southeast, Southern Asia.....Rubrauletes Legalov, 2003

Subgenus Pseudomesauletes s. str.

Alonsoiauletes Legalov, 2003a: 128, syn.n.; type species: Auletobius simillimus Voss, 1933 Remarks. Comparative study of African and Asian species showed that Alonsoiauletes Legalov, 2003, syn.n. is synonym to Pseudomesauletes s. str.

Pseudomesauletes (Pseudomesauletes) podocarpi (Voss, 1933), comb.n., placem.n. (col. pl. III: l, IX: 19-20) Auletobius podocarpi Voss, 1933: 128 Remarks. The lectotype is designated by the author - a male from the collection NHRS with labels “Uitenhage, Cape Col., J. Oneie”, “Auletobius podocarpi m.”, “Paratypus”, “Auletobiuspodocarpi det Voss”, “7180 E91”, “podocarpi Voss”, “Lectotype Auletobius podocarpi Voss, 1933, A. Legalov design. 2009”.

Previously, this species has been placed wrongly in the genus Kuschelanthus Alonso-Zarazaga & Lyal, 1999. Distribution. S-Africa.

Pseudomesauletes (Pseudomesauletes) ueleanus (Voss, 1939)

Auletobius ueleanus Voss, 1939c: 46

Material. 2 ex. (ISNB), 1 ex. (SZMN), “Cameroun,

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Faro Game Reserve N 8'23'6'2', E 12'50'7'1, Wooded

Savanna Pinata, 25.05.2007, Jocque, Loosveld, Baert &

Alderweireldt”

Remarks. This species is reported for the first time for the fauna of Cameroon.

Distribution. Cameroon, Guinea, Zaire. Pseudomesauletes (Pseudomesauletes) subsignatus (Voss, 1922), comb.n. (col. pl. IV: c-d, f, IX: 17-18)

Auletobius subsignatus Voss, 1922a: 36 Pseudomesauletes stanleyvillensis Legalov, 2007: 62, syn.n.

Material. 1 ex. (ISNB), Ghana, Kakum Nat. Parc, Primary rain forest, Fogging, 5.20'55 N, 1.23 E, 159 m, 17.11.2005,

D. Debakker.

Remarks. The lectotype is designated by the author -a male from the collection ZMHB with labels “Span. Guinea, Nkolentagan, 11.07 - 05.08., G. Tessmann S.G.”, “Auletobius subsignatus m.”, “SYNTYPUS Auletobius subsignatus Voss, 1922, labelled by MNHUB 2009”, “Lectotype Auletobius subsignatus Voss, 1922, A. Legalov design. 2009”. Paralectotypes: 3 males and 4 females labeled as lectotypes. Study of the type specimens proved that Pseudomesauletes stanleyvillensis Legalov, 2007, syn.n. is synonym to P subsignatus (Voss, 1922). Distribution. Guinea, Zaire.

Pseudomesauletes (Pseudomesauletes) ? simillimus (Voss, 1933), comb.n.

Auletobius simillimus Voss, 1933: 126

Remarks. Specimen studied: a male from the collection

MRAC with labels “Musee du Congo, Inongo, 1 - III -1015, R. Mayne”, “R. Det. N 919”, “Auletobius subsignatus Voss, Hustache det.”.

Distribution. Congo, Zaire.

Pseudomesauletes (Pseudomesauletes) ater (LeConte, 1876)

Auletes ater LeConte, 1876: 4

Material. 2 ex. (USNM), USA, NC: Moore Co., Southern Pines, 10.05.1964, G. Vogt.

Distribution. North America.

Pseudomesauletes (Pseudomesauletes) formosanus (Voss, 1921) (col. pl. I: vg, IX: 21-22)

Auletobius uniformis ssp. formosanus Voss, 1921: 277 Auletobius subtuberculatus Voss, 1921: 278, syn.n. Auletobius tuberculatus Voss, 1921: 278, syn.n.

Auletobius hirtellus Voss, 1941a: 240, syn.n.

Remarks. The lectotype for Auletobius uniformis ssp. formosanus is designated by the author - a male from the collection DEI with labels “Kankau (Koshun), Formosa, H. Sauter, V.1912”, “Voss det.”, “Syntypus”, “Auletobius uniformis Roel. ssp. formosanus m.”, “Coll. DEI Muncheberg”, “Auletobius uniformis ssp. formosanus Voss”, “Lectotype Auletobius uniformis formosanus Voss, A. Legalov des. 2007”. The holotype for Auletobius subtuberculatus was studied: a male from the collection DEI with labels “Holotypus”, “Holotypus”, “Kankau (Koshun), Formosa, H. Sauter, 05.1912”, “Auletobius subtuberculatus m., male”, “Voss det.”, “Auletobius subtuberculatus Voss”, “Pseudomesauletes subtuberculatus Voss, A. Legalov det.”, “coll. DEI Eberswalde”. The syntype for Auletobius tuberculatus was studied: a female from the collection DEI with labels “Syntypus”, “Syntypus”, “Hokuto, Formosa,

H. Sauter, 05.1912”, “Hokuto, auf Rubus Art, III.12”, “7.III.”, “male”, “Auletobius tuberculatus m.”, “Voss det.”, “Auletobius tuberculatus Voss”, “Pseudomesauletes tuberculatus Voss,A. Legalov det.”, “coll. DEI Eberswalde”. The lectotype is designated by the author - a male from the collection ZFMK with labels Kwangtseh - Fukien, J. Klapperich, 4.9.1937”, “Paratypoid Auletobius hirtellus Voss”, “Lectotype Auletobius hirtellus Voss, 1941, A. Legalov design. 2009”. Paralectotypes: male (ZFMK) with labels “Kwangtseh - Fukien, J. Klapperich, 10.9.1937”, “Paratypoid Auletobius hirtellus Voss”, “Paralectotype Auletobius hirtellus Voss, 1941, A. Legalov design. 2009”; female (with labels) with labels “Kwangtseh - Fukien, J. Klapperich, 21.09.1937”, “Paratypoid Auletobius hirtellus Voss”, “Paralectotype Auletobius hirtellus Voss, 1941, A. Legalov design. 2009”. Study of the type specimens and large materials from China and Vietnam proved that Auletobius subtuberculatus Voss, 1921, syn.n., A. tuberculatus Voss, 1921, syn.n. and A. hirtellus Voss, 1941, syn.n. are synonyms to Pseudomesauletes formosanus (Voss, 1921).

Distribution. South-eastern China, Taiwan, Vietnam. Pseudomesauletes (Pseudomesauletes) collarti Legalov, sp.n. (col. pl. IV: e, IX: 27)

Material. Holotype - male (ISNB), “Stanleyville,

1-10.11.1929, A. Collart”.

Description. Male. Body brown-black. Legs and antennae paler. Body with short almost adpressed pale setae.

Rostrum long, 5.0 times longer than wide, 1.18 times longer than pronotum, weakly curved, widened to the apex, punctate. Antennae located in the middle of rostrum. Eyes large, strongly convex. Frons wide, strongly convex, densely punctate. Temples short and straight. Antennae long, reaching apical margin of pronotum. Scapus and 1st segment of funicle oval. 2nd-4th segments long-oval, narrower. 2nd segment longer than 1st segment. 3rd segment shorter than 2nd segment. 4th segment of almost equal length to 3rd segment. 5th and 6th segments trapezoid. 7th segment transversal, wider than 6th segment. Clava wide, almost compact, pointed, slightly shorter than funicle. 1st and 2nd segments transversal. 3rd segment stilliform, slightly shorter than previous segments. Pronotum almost campaniform, 1.1 times longer than wide, very weakly narrowed to basis and apex. Disk convex, finely and densely punctate. Greatest width in the middle. Scutellum trapezoid. Elytra almost rectangular, elongated, 1.29 times longer than wide. Greatest width behind the middle. Humeri weakly smoothed. Striae reduced. Points large and deep. Intervals weakly convex. Apex of elytra with sex patches. Thorax finely and sparsely punctate. Metepisternum narrow. Abdomen convex. 1st and 2nd ventrites wide, approximately equal in length. 3rd and 4th ventrites narrower than 2nd segment. 5th ventrite narrow, narrower than 4th segment. Pygidium convex, punctate. Legs long. Femora widened. Tibiae almost straight, weakly widened to apex. Tarsi long. 1st tarsal segment long-triangular. 2nd segment wide-triangular, flattened. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 2.9 mm.

Diagnosis. The new species is close to P. subsignatus but differs with smaller body, weaker rounded sides of pronotum and with armament of endophallus.

Etymology. New species is named in honour of A. Collart. Distribution. Zaire.

Pseudomesauletes (Pseudomesauletes) friedmani Legalov, sp.n. (col. pl. IV: k-l, IX: 28)

Material. Holotype - male (NMKE), “Toug. Terr., Ukerewe, Father Conrads”, “III.2812”. Paratype - female (NMKE), idem.

Description. Body brownish black, with short semierect pale setae. Male. Rostrum long, 4.38 times longer than wide, 1.4 times longer than pronotum, very weakly curved, widened to the apex, densely punctate. Antennae located in the middle of rostrum. Eyes large, strongly convex. Frons wide, strongly convex, densely punctate. Temples short. Antennae long, reaching beyond the apical margin of pronotum. Scapus and 1st segment of funicle oval. 2nd segment long-oval, narrower, longer than 1st segment. 3rd-6th segments trapezoid. 7th segment transversal. Clava wide, almost compact, pointed, shorter than funicle. 1st and 2nd segments transversal. 3rd segment stilliform, slightly shorter than previous segments. Pronotum almost campaniform, of equal length and width, weakly narrowed to basis and apex. Disk convex, densely punctate, with greatest width near the middle. Scutellum trapezoid. Elytra almost rectangular, elongated, 1.32 times longer than wide. Greatest width behind the middle. Humeri weakly smoothed. Striae reduced. Points large and deep. Intervals

weakly convex. Apex of elytra with sex patches. Thorax finely and sparsely punctate. Metepisternum narrow. Abdomen convex. 1st and 2nd ventrites wide. 2nd ventrite slightly wider than 1st segment. 3rd and 4th ventrites narrower than 2nd segment. 5th ventrite narrow, narrower than 4th segment. Pygidium convex, punctate. Legs long. Femora widened. Tibiae almost straight, weakly widened to apex. Tarsi long. 1st tarsal segment long-triangular. 2nd segment wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 2.8 mm. Female. Rostrum shorter, 3.7 times longer than wide, 1.23 times longer than pronotum. Antennae attached before the middle of rostrum. Pronotum 1.13 times wider than long. Elytra 1.25 times longer than wide, without sex patches. Length of body: 2.3 mm.

Diagnosis. The new species is close to P ueleanus differing with the shorter and thicker rostrum, more densely punctate disk of pronotum and with the armament of endophallus. Etymology. New species is named in honour of A.-L.-L. Friedman.

Distribution. Tanzania.

Pseudomesauletes (Pseudomesauletes) culex (Scudder,

1893)

Docirhynchus culex Scudder, 1893: 24 Distribution. Lower Oligocene (USA: Colorado). Pseudomesauletes (Pseudomesauletes) ibis (Wickham, 1912), comb.n.

Docirhynchus ibis Wickham, 1912: 34 Remarks. This species is very close to P. culex, hence it is transferred from genus Docirhynchus Scudder, 1893 to genus Pseudomesauletes.

Distribution. Lower Oligocene (USA: Colorado).

Subgenus Afromesauletes Legalov, 2003

Pseudomesauletes (Afromesauletes) punctipennis (Hustache, 1923) (col. pl. IV: h-j, IX: 23, 25)

Rhynchites punctipennis Hustache, 1923: 153 Remarks. The lectotype is designated by the author - a male from the collection MRAC with labels “Holotypus”, “Musee du Congo, Wombali, 07 - 1913, P. Vanderijst”, “R. Det. MM 920”, “Rhynchites punctipennis Type, Hust.”, “Lectotype Rhynchites punctipennis Hust., A. Legalov design. 2009”. Paralectotype - a male from the collection MRAC with labels “Paratypus”, “Musee du Congo, Wombali, 07 - 1913, P. Vanderijst”, “R. Det. M 920”, “R. Det. G 3415”, “Paralectotype Rhynchites punctipennis Hust., A. Legalov design. 2009”. Specimens studied: a female from the collection MRAC with labels “Musee du Congo, Haut-Uele: Moto, n. Dau, 01.1925, L. Burgeon”, “R. Det. 2321 C”, “Auletobius callosus Voss female, Hustache det.”, “R. Det. H 3415” and a female from the collection MRAC with labels “Musee du Congo, Wombali, 07.1913, P. Vanderijst”, “R. Det. F. 3415”s.

Distribution. Burkina Faso, Cameroon, Guinea, Nigeria, Zaire.

Subgenus Madauletes Legalov, subgen.n.

(col. pl. IV: m-n, V: a-b)

Type species: Auletobius gibbipennis Hustache, 1955 Description. Body yellowish brown. Pale short adpressed setae concentrated near eyes, near scutellum, on thorax,

form 3 longitudinal strips on pronotum or concentrated on head, legs, thorax, form not distinct spots on pronotum and elytra. Rostrum long, 6.11-6.28 times longer than wide, 1.52-1.57 times longer than pronotum, barely curved, widened to the apex, almost smooth or finely and sparsely punctate, sometimes flattened. Antennae located before the middle of rostrum. Eyes large, strongly convex. Frons wide, strongly convex, sparsely punctate. Temples short and straight. Antennae long, reaching the middle of pronotum. Scapus and 1st segment of funicle long-oval. 1st segment flattened, longer than scapus. 2nd segment long-oval, narrower and shorter than 1st segment. 3rd-4th segments trapezoid, shorter, approximately equal in length. 5th-6th segments oval, approximately equal in length, shorter than 3rd-4th. 7th segment almost roundish, short. Clava wide, almost compact, pointed, shorter than funicle. 1st and 2nd segments transversal. 1st segment narrower than 2nd segment. 3rd segment stilliform, slightly shorter than previous segments. Pronotum almost campaniform, slightly elongated or of equal length and width, weakly narrowed to basis and apex. Sides very weakly rounded. Disk convex, finely and densely punctate. Greatest width in the middle. Scutellum trapezoid. Elytra almost rectangular, elongated, 1.38-1.48 times longer than wide. Greatest width behind the middle. Humeri weakly smoothed. Striae reduced, points do not form rows or points form almost distinct rows in first half of elytra. Points large and deep. Intervals almost flat, wide or narrow. Apex of elytra with sex patches in males. Thorax finely and sparsely punctate. Metepisternum narrow. Abdomen convex. 1st and 2nd ventrites wide. 2nd ventrite slightly wider than 1st ventrite. 3rd and 4th ventrites narrower than 2nd ventrite, of almost equal length. 5th ventrite narrow, slightly narrower than 4th. ventrite Pygidium convex, punctate. Legs long. Femora widened. Tibiae almost straight, weakly widened to apex. Protibiae more narrow and long. Tarsi long. 1st tarsal segment long-triangular. 2nd segment more wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 2.6-2.9 mm. Diagnosis. See key to subgenera of the genus Pseudomesauletes.

Etymology. The name is formed from the words “Madagascar” and “auletes”.

Pseudomesauletes (Madauletes) ankaratraensis (Hustache, 1955), comb.n. (col. pl. IV: m-n)

Auletobius ankaratraensis Hustache, 1955: 192 Remarks. The holotype was studied: a female from the collection MNHN with labels “Madagascar, Ankarastra”, “male”, “Type”, “Auletobius (Aletinus) ankaratraensi m.”, “Museum Paris, 1919, Col. A. Hustache”, “Holotype Auletobius ankaratraensi Hustache, 19, A. Legalov det. 2009”.

Distribution. Madagascar.

Pseudomesauletes (Madauletes) gibbipennis (Hustache, 1955), comb.n. (col. pl. V: a-b)

Auletobius gibbipennis Hustache, 1955: 191 Remarks. The holotype was studied: a male from the collection MNHN with labels “Madagascar, La Mandraka, Vadon !”, “male”, “Type”, “Auletobius (Aletinus) gibbipennis m.”, “Museum Paris, 1919, Col. A. Hustache”,

“Holotype Auletobius gibbipennis Hustache, 19, A. Legalov det. 2009”.

Distribution. Madagascar.

Subgenus Rubrauletoides Legalov, subgen.n.

(col. pl. V: c, IX: 24, 26)

Type species: Pseudomesauletes jizushanensis Legalov, sp.n.

Description. Body black-brown, with adpressed pale setae. Rostrum long, weakly curved, punctate. Antennae in males located in the middle of rostrum. Eyes not large, strongly convex. Frons wide, punctate. Temples short. Pronotum almost campaniform. Disk convex, punctate. Elytra almost rectangular, elongated, with carinae. Striae reduced. Intervals weakly convex. Apex of elytra in males with sex patches. Thorax punctate. Abdomen convex. 1st and 2nd ventrites wide. 3rd and 4th ventrites narrower. 5th ventrite narrow. Pygidium convex. Legs long. Femora widened. Tibiae almost straight. Tarsi long. Claws with long teeth. Length of body: 3.1 mm.

Diagnosis. See key to subgenera of the genus Pseudomesauletes.

Etymology. The name is formed by addition of the ending “-ides” to “Rubrauletes”.

Pseudomesauletes (Rubrauletoides) jizushanensis Legalov, sp.n. (col. pl. V: c, IX: 24, 26)

Material. Holotype - male (NMPC), China, Yunnan prov., Jizushan, S slope, 1600-2300 m, 23.07.1995, Bolm. Description. Male. Body black-brown. Elytra reddish brown. Head and rostrum black. Body with short adpressed pale setae. Rostrum long, 7.29 times longer than wide, 1.65 times longer than pronotum, weakly curved, widened to the apex, densely punctate. Antennae located in the middle of rostrum. Eyes not large, strongly convex. Frons wide, convex, finely punctate. Temples short and straight. Pronotum almost campaniform, of almost equal length and width, weakly narrowed to basis and apex. Disk convex, finely and densely punctate. Greatest width in the middle. Scutellum trapezoid. Elytra almost rectangular, elongated, 1.39 times longer than wide, with weak carinae. Greatest width behind the middle. Humeri weakly smoothed. Striae reduced. Points large and deep. Intervals weakly convex. Apex of elytra with sex patches. Thorax finely and sparsely punctate. Metepisternum narrow. Abdomen convex. 1st and 2nd ventrites wide. 2nd ventrite slightly wider than 1st ventrite. 3rd and 4th ventrites narrower than 2nd ventrite. 4th ventrite slightly wider than 3rd ventrite. 5 th ventrite narrow, narrower than 4th ventrite. Pygidium convex, punctate. Legs long. Femora widened. Tibiae almost straight, slightly widened to apex. Tarsi long. 1st tarsal segment long-triangular. 2nd segment wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 3.1 mm.

Distribution. China (Yunnan).

Subgenus Fascauletes Legalov, 2007

Pseudomesauletes (Fascauletes) fuscofasciatus (Voss, 1933), placem.n.

Auletobius fuscofasciatus Voss, 1933: 127

Remarks. Previously, this species has been wrongly placed

in the subgenus Rubrauletes Legalov, 2003.

Distribution. India (South).

Pseudomesauletes (Fascauletes) maculatus (Voss, 1933), placem.n.

Auletobius maculatus Voss, 1933: 127

Remarks. Previously, this species has been wrongly placed

in Rubrauletes Legalov, 2003.

Distribution. Sri Lanka.

Subgenus Rubrauletes Legalov, 2003

Pseudomesauletes (Rubrauletes) chinensis (Voss, 1933)

Auletobius chinensis Voss, 1933: 128 Material. 1 ex. (RDP), China, Yunnan centr. occ., Dali, 19-21.05.1993, R. Cervenka; 1 ex. (RDP), Yunnan, Dali W env., 20-25.08.1998, O. Safranek, M. Tryzna. Distribution. China (Fujian, Yunnan).

Pseudomesauletes (Rubrauletes) poirasi Legalov, 2009 Pseudomesauletes poirasi Legalov, 2009: 64 Material. 5 ex. (NME), 2 ex. (SZMN), Nepal, N Kathmandu, Shivapuri, 1800-2500 m, 24.06.1980, C. Holzschuh; 1 ex. (NME), Nepal, Helambu upp., Chipling, 2200-2400 m, 29-30.08.1997, S. Fabrizi, D. Ahrens. Distribution. North India, Nepal.

Pseudomesauletes (Rubrauletes) binbyanicus Legalov, 2007

Pseudomesauletes binbyanicus Legalov, 2007: 65 Material. 1 ex. (RDP), China, Yunnan, Maguan, 23.04 N, 104.25 E, 1500-1600 m, 25-26.06.1994, Z. Cernin; 1 ex. (RDP), 2 ex. (SZMN), China, Yunnan, Malipo, 23.09 N, 104.41 E, 1250-1350 m, 23.06.1994, V. Kuban. Distribution. China (Yunnan).

Pseudomesauletes (Rubrauletes) nepalensis (Voss, 1974)

Auletobius nepalensis Voss, 1974: 44 Material. 1 ex. (NME), Nepal, N Kathmandu, Shivapuri, 1800-2500 m, 24.06.1980, C. Holzschuh; 1 ex. (VSM), Nepal, prov. Koshi, vill. Hilles, 06.1999, V. Patrikeev; 1 ex. (ZIN), Nepal, env. Of Kathmandu, Royal Forest-Shivapuri, 27.44’66 N, 85.17’76 E, 1730 m, 16.05.2000, Konstantinov, Lingafelter, Volkovitsh; 1 ex. (RDP), E Nepal, Kangchenjunga Himal Mts., Chiruwa vill., 27.29 N, 87.45 E, 1260 m, 30.06.-1.07.2000, J. Schneider; 1 ex. (RDP), India, Sikkim east., Gantok env., Fambong-Lho forest., 2000-2500 m, 8-15.07.1997, J. Schneider; 2 ex. (RDP), 1 ex. (SZMN), N India, Sikkim, Mangan vill. env., 3500 m, 24-27.05.2002, M. Tryzna & P. Benda. Distribution. East India, Nepal.

Subgenus Faustiauletes Legalov, 2003

Pseudomesauletes (Faustiauletes) gestroi (Faust, 1894)

Auletes gestroi Faust, 1894a: 169 Auletobius gestroi f. dispar Voss, 1935: 99 Material. 1 ex. (ZIN), Vietnam, Nghetinh Prov., mountains SW of Qui Chau, 200 m, 14.12.1963, O. Kabakov; 1 ex. (ZIN), Vietnam, Con Dao, Van Kan Is., 3.04.1987,

I. Darevsky; 1 ex. (SZMN), Vietnam, Hashonbin prov, Tuli, 16. 10.1990, A. Gorokhov; 1 ex. (APB), Laos centr., Vientiane env., Mekong river bank, 150 m, 102.37’3 E,

26.04.1997, M. Sirba & R. Hergovits.

Remarks. This species is reported for the first time for the fauna of Laos.

Distribution. Cambodia, India, Myanmar, Thailand,

Vietnam, Laos, Indonesia (Java).

Genus Cylauletes Legalov, 2007

Cylauletes combreti (Voss, 1933)

Auletobius combreti Voss, 1933: 129

Material. 2 ex. (RDP), 1 ex. (SZMN), S Africa, 25 km W

Pretoria, Saartjiesnek, 25.46 S, 27.54 E, 16-28.12.1997, S.

Bily.

Distribution. S-Africa.

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Genus Dicranognathus Redtenbacher, 1844

Dicranognathus nebulosus Redtenbacher, 1844

Dicranognathus nebulosus Redtenbacher, 1844: 538 Material. 2 ex. (RDP), 2 ex. (CKJU), 2 ex. (SZMN), N India, Uttaranchal state, 30 km N of Rishikesh, NW of Chamba, Arakot vill. env., 1500 m, 29-31.07.2003, Z. Kejval & M. Tryzna.

Distribution. North India.

Genus Pseudodicranognathus Legalov, 2001

Pseudodicranognathus fuliginosus (Voss, 1933), comb.n., placem.n.

Auletobiusfuliginosus Voss, 1933: 129

Remarks. Previously, this species has been wrongly placed

in the Rubrauletes Legalov, 2003.

Distribution. East India.

Genus Eumetopon Voss, 1922 Subgenus Eumetopon s. str.

Eumetopon (Eumetopon) flavomaculatus (Voss, 1922)

Auletobius flavomaculatus Voss, 1922a: 37 Auletobius flavomaculatus f. chinensis Voss, 1939a: 608 [non Voss, 1933]

Eumetopon flavomaculatus ssp. eduardi Legalov, 2003b: 13 [RN]

Material. 1 ex. (SZMN), China, Zejiang, Tianmu Shan, near Dequing, 5.10.1997.

Remarks. This species is reported for the first time for the fauna of Zejiang.

Distribution. South-eastern China, Southern India.

Tribe Isotheini Scudder, 1893 Subtribe Deporaina Voss, 1929 Genus Capylarodepopsis Legalov, 2003

Key to species similar to Capylarodepopsis nigrilineatus

1. Frons convex. Sides of pronotum stronger rounded. Basal sclerite as in fig. 215 in Legalov [2007]. Sumatra .. C. nigrilineatoides Legalov, 2007

- frons flattened. Sides of pronotum weaker rounded. Basal

sclerite as in col. pl. IX: 29.........................2

2. Frons more densely punctate with deep striae. Kalimantan .............................C. nigrilineatus (Voss, 1922)

- frons more sparsely punctate with weak striae. Kalimantan ..................................C. confinis (Voss, 1938)

Capylarodepopsis confinis (Voss, 1938) (col. pl. V: f, IX: 29)

Deporaus confinis Voss, 1938a: 91 Remarks. The lectotype is designated by the author - a male from the collection DEI with labels “Wahnes, Borneo”, “Coll. Kraatz”, “Syntypus”, “Deporaus (Capylarodepus) confinis n. sp., Det. E. Voss”, “coll. DEI Muncheberg”,

“Deporaus confinis Voss”, “Lectotype Deporaus confinis Voss, 1938, A. Legalov design. 2009”.

Distribution. Kalimantan.

Genus Biblarodepus Voss, 1924

Biblarodepus solitarius (Voss, 1938) (col. pl. V: d, g, IX: 30)

Deporaus solitarius Voss, 1938a: 93 Remarks. The holotype was studied: a male from the collection DEI with labels “Kina Balu”, “Coll. Kraatz”, “Holotypus”, “Deporaus (Arodepus) solitarius n. sp., Det. E. Voss”, “Dep. solitarius Voss”, “coll. DEI Muncheberg”, “Deporaus solitarius Voss”, “Holotype Deporaus solitarius Voss, 1938, A. Legalov det. 2009”.

Distribution. Malaysia (Sabah).

Biblarodepus solutus (Voss, 1938) (col. pl. V: h, j) Deporaus solutus Voss, 1938a: 94 Remarks. The lectotype is designated by the author - a female from the collection DEI with labels “Philippinen, Mindanao, Mangarin”, “Boettcher S. 11.1917”, “Deporaus solutus m.”, “Syntypus”, “coll. DEI Muncheberg”, “Deporaus solutus Voss”, “Lectotype Deporaus solutus Voss, 1938, A. Legalov design. 2009”.

Distribution. Philippines.

Genus Caenorhinus C.G. Thomson, 1859 Subgenus Metallarodepus Legalov, 2003

Caenorhinus (Metallarodepus) rufiventris (Voss, 1921), comb.n., placem.n. (col. pl. V: e, m, IX: 31)

Deporaus rufiventris Voss, 1921: 280 Remarks. The lectotype is designated by the author - a male from the collection ZMHB with labels “Formosa, Taihorinho, XI.09., H. Sauter S. G.”, “Deporaus rufiventris n. sp.”, “SYNTYPUS Deporaus rufiventris Voss, 1921 labelled by MNHUB 2009”, “Lectotype Deporaus rufiventris Voss, 1921, A. Legalov design. 2009”. Previously, this species has been wrongly placed in the genus Neoarodepus Legalov, 2003.

Distribution. China (Taiwan).

Subgenus Flavodeporaus Legalov, 2007

Caenorhinus (Flavodeporaus) fukienensis (Voss, 1941)

Deporaus fukienensis Voss, 1941a: 245 Remarks. Specimen studied: a female from the collection DEI with labels “China”, “Kuatun (2300 m), 27.40 n, Br., 117.406 L., J. Klapperich. 8.4., 1938 (Fukien)”, “coll. DEI Muncheberg.

Distribution. China (Fujian).

Tribe Rhynchitini Gistel, 1848 Subtribe Lasiorhynchitina Legalov, 2003 Genus Nelasiorhynchites Legalov, 2003

Nelasiorhynchites olivaceus (Gyllenhal, 1833) (col. pl. V: i, k)

Rhynchites olivaceus Gyllenhal, 1833: 228 Rhynchites comatus Gyllenhal, 1833: 229 Rhynchites pauciseta Wasmann, 1884: 252 Remarks. The lectotype is designated by the author -a female from the collection HNHM with labels “Im Wald bei Cleve auf Eichembusch”, “female, 23.5.1882”, Paratypus 1884 Rhynchites pauciseta Wasmann”, “Rhynchites pauciseta m., 23.5.82, Im Wald bei Cleve auf Eichembusch”, “Coll. Reitter”, “Lectotype Rhynchites

pauciseta Wasmann, 1884, A. Legalov design. 2009”. Distribution. Western Palaearctic.

Subtribe Rhynchitina Gistel, 1848 Genus Cartorhynchites Voss, 1958 Subgenus Cartorhynchoides Legalov, 2003

Cartorhynchites (Cartorhynchoides) breviusculus (Voss, 1939), placem.n.

Rhynchites breviusculus Voss, 1939b: 63 [RN]

Rhynchites brevirostris Voss, 1938b: 145 [non Roelofs,

1874]

Rhynchites breviusculus f. haematopus Voss, 1940: 85 Involvulus breviusculus ssp. bintamensis Voss, 1969: 145 Remarks. Previously, this species has been wrongly placed in Hyperinvolvulus Legalov, 2003.

Distribution. Indonesia (Java, Sumatra), Vietnam.

Subgenus Hyperinvolvulus Legalov, 2003

Cartorhynchites (Hyperinvolvulus) crassiusculus (Voss, 1938), placem.n.

Rhynchites crassiusculus Voss, 1938b: 145

Remarks. Previously, this species has been wrongly placed

in Cartorhynchoides Legalov, 2003.

Distribution. East India.

Genus Maculinvolvulus Legalov, 2003

Maculinvolvulus vestitoides (Legalov, 2002) (col. pl. V: l, n)

Cartorhynchites vestitoides Legalov, 2002: 91 [RN] Rhynchites vestitus Voss, 1938b: 144 [non Rey, 1893] Remarks. The lectotype is designated by the author

- a female from the collection ZMHB with labels “38996”, “Ceylon, Nietn.”, “Rhynchites vestitus n. sp.”, “SYNTYPUS Rhynchites vestitus Voss, 1938 labelled by MNHUB 2009”, “Lectotype Rhynchites vestitus Voss, 1938, A. Legalov design. 2009”.

Distribution. South India, Sri Lanka.

Genus Metarhynchites Voss, 1923 Subgenus Metarhynchites s. str.

Metarhynchites (Metarhynchites) longulus (Gyllenhal, 1833), comb.n., placem.n. (col. pl. VII: m)

Rhynchites longulus Gyllenhal, 1833: 234 Rhynchites pullus Voss, 1935b: 104, syn.n.

Rhynchites pullus Voss, 1938b: 140 [non Voss, 1935] Remarks. The lectotype is designated by the author - a female from the collection NHRS with labels “Typus”, “Java, Mellenb.”, “Lectotype Rhynchites longulus Gyllenhal, 1833, A. Legalov design. 2009”. Previously, this species has been placed in Rhynchitidae incertae sedis. Study of type specimens showed that Rhynchites pullus Voss, 1935, syn.n. is synonym to Metarhynchites longulus (Gyllenhal, 1833).

Distribution. South India, Indonesia (Java, Sumatra). Genus Pseudomechoris Legalov, 2003

Pseudomechoris aethiops (Bach, 1854) (col. pl. VI: d)

Rhynchites aethiops Bach, 1854: 172 Involvulus aethiops ssp. juraensis Voss, 1969: 250 Remarks. The lectotype is designated by the author - a female from the collection HNHM with labels “Dole, Jura”, “Paratypus 1954 Rhynchites aethiops ssp. juraensis

Voss”, “Paratypus Rh. aethiops ssp. juraensis m.”, “Rynchites aethiops Bach ssp. juraensis m.”, “Lectotype Involvulus aethiops ssp. juraensis Voss, 1969, A. Legalov design. 2009”.

Distribution. Western Palaearctic.

Genus Cyllorhynchites Voss, 1930

Cyllorhynchites (Cyllorhynchites) ursulus rostralis (Voss, 1930) (col. pl. VI: a-c)

Rhynchites rostralis Voss, 1930: 78 Rhynchites homalinus Voss, 1930: 76, syn.n.

Remarks. The lectotype of Rhynchites homalinus is designated by the author - a female from the collection ZMHB with labels “Yun-nan sen”, “Rhynchites homalinus Voss, Voss”, “Rhynchites homalinus Voss, China”, “SYNTYPUS Rhynchites homalinus Voss, 1930 labelled by MNHUB 2009”, “Lectotype Rhynchites homalinus Voss, 1930, A. Legalov design. 2009”.

Study of type specimens proved that Rhynchites homalinus Voss, 1930, syn.n. is synonym to Cyllorhynchites ursulus rostralis (Voss, 1930).

Distribution. China.

Genus Clinorhynchites Voss, 1969 Key to species of the genus Clinorhynchites

1. Clava of normal structure..............................2

- Clava strongly flattened and widened...................5

2. Femora black...........................................3

- Femora red-brown.......................................4

3. Eyes stronger convex. 2nd and 4th segments of the

funicle of equal length. Rostrum longer. Western and Central Africa.................C. nigripes (Faust, 1894)

- Eyes weaker convex. 2nd segment slightly longer than

4th segment. Rostrum shorter. Western Africa..............

...............................C. castaneus (Jekel, 1860)

4. Rostrum shorter. Antennae located before the middle of rostrum. Tibiae yellowish-brown. Rostrum black (except

basis). Pronotum narrow. Western and Central Africa.......

.........................C. distinguendulus (Voss, 1939)

- Rostrum longer. Antennae located in the first third of rostrum. Tibiae black or red-brown, dark. Rostrum black up to place of antennal attachment. Pronotum wide.

Western, Central, Eastern Africa..........................

.........................C. rufofemoratus (Voss, 1938)

5. Antennae in females located near the basis of rostrum.

Frons wider, flat. Central Africa.........................

...............................C. scheitzae (Voss, 1944)

- Antennae in females located further from the basis of

rostrum. Frons narrower, convex. Western and Central

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Africa....................C. despectus (Voss, 1938)

Clinorhynchites castaneus - group Clinorhynchites castaneus (Jekel, 1860) (col. pl. VI: p-q)

Rhynchites castaneus Jekel, 1860: 241 Remarks. The lectotype was studied: a female from the collection MCSN with labels “Rhynchitespicipes, Buques, Guinea”, “Castaneus Jekel”, “Syntypus Rhynchites castaneus Jekel, 1860”, “Museo Genova, coll. H. Jekel, via coll. A. Solari (acqusto 2000)”, “Lectotype Rhynchites castaneus Jek., A. Legalov design. 2008”.

This species is distributed in the Western Africa. All material from Zaire belongs to Clinorhynchites rufofemoratus. Distribution. Cameroon, Guinea.

Clinorhynchites distinguendulus (Voss, 1939) (col. pl. VI: g)

Rhynchites distinguendulus Voss, 1939b: 66 [RN] Rhynchites distans Voss, 1938b: 146 [non Sharp, 1889] Remarks. Specimen studied: a female from the collection MRAC with labels “Musee du Congo, Lulua: Kapanga, 09.1932, F.G. Overlaet”, “R. Det. Z 3415”, “Rhynchites distans m., Det. E. Voss”.

Distribution. Cameroon, Zaire.

Clinorhynchites nigripes (Faust, 1894) (col. pl. VI: e-f) Rhynchites nigripes Faust, 1894b: 528 Remarks. Specimens studied: the lectotype - a male from the collection SMTD with labels “gold small square”, “Gabun, Staudgr.”, “Coll. J. Faust, Ankauf 1900”, “Staatl. Museum fur Tierkunde, Dresden”, “Type”, “nigripes Faust”, “Rhynchites castaneus Jek. f nigripes Fst.”, “Lectotype Rhynchites nigripes Fst., A. Legalov design. 2005” - and a female from the collection MRAC with labels “Musee du Congo Belge, Mayumbe, Cabra”, “Rhynchites castaneus Jek., det. Gakm. 1909”, “R. Det. H 319”, “R. Det. Y 3415”, “Rhynchites castaneus Jek. v. nigripes Fst., Det. E. Voss”.This species is very close to Clinorhynchites castaneus but differs with minor characters in the armament of the endophallus, with convexity of eyes and shape of funicle segments. Probably, it is synonym of Clinorhynchites castaneus.

Distribution. Gabon, Zaire.

Clinorhynchites rufofemoratus (Voss, 1938), comb.n., stat.n. (col. pl. VI: j-k)

Rhynchites castaneus f. rufofemorata Voss, 1938b: 146 Material. 2 ex. (ISNB), 1 ex (SZMN), “Congo Belge, Ikela (Equateur), I. G. 20.536, Leg. R. Deguide”; 1 ex. (NMPC), “Kenya Colony, Africa orient, Shuling lgt.”; 1 ex. (MRAC), “Coll. Mus. Congo, Bambesa, 1.10.1938, J. Vrydag”; 1 ex. (MRAC), “Coll. Mus. Congo, Bambesa,

II-III.1938, J. Vrydagh”; 1 ex. (MRAC), “Coll. Mus. Congo, Bambesa, 4.10.1938, J. Vrydagh”; 1 ex. (MRAC), “Coll. Mus. Congo, Bambesa, 1.10.1938, J. Vrydagh”; 1 ex. (MRAC), “Parasite de fruit d’une Liane (Anonacee)”, “Musee du Congo, Sankuru: Komi, Jodja, 10.1929, J. Ghesquiere”, “Larvae in seeds of liana”, “Congo Bege, Lodia, 10.1929, J. Ghesquiere”, “Congo Belge, Lidja, 10.1925, Rec. J. Ghesquiere”; 1 ex. (MRAC), “Coll. Mus. Congo, Bambesa, 4.10.1938, J. Vrydagh”; 1 ex. (MRAC), “Coll. Mus. Congo, Bambesa, 11.V1938, J. Vrydagh”. Remarks. The lectotype is designated by the author - a male from the collection ZMHB with labels “461”, “Njam-Njam, Semnio, Bohndorff S.”, “Rhynchites castaneus f. rufofemorata m. Det. E. Voss”, “SYNTYPUS Rhynchites castaneusf. rufofemorata Voss, 1938 labelled by MNHUB 2009”, “Lectotype Rhynchites castaneus f. rufofemorata Voss, 1938, A. Legalov design. 2009”.

This species differs from C. castaneus with the colour of femora and minor differences in the armament of the endophallus.

Distribution. Western and Central Africa.

Clinorhynchites despectus - group Clinorhynchites despectus (Voss, 1938)

Rhynchites despectus Voss, 1938b: 146 Rhynchitesplatynotus Voss, 1938b: 147, syn.n.

Material. 1 ex. (SMTD), “Gabun”, “Samml. K. F. Hartmann, Ankauf, 1941”, “Staatl. Museum fur Tierkunde, Dresden”, “Rhynchites castaneus Jek.”, “Rhynchites despectus m., Det. E. Voss”, “Lectotype Rhynchites despectus Voss, A. Legalov design. 2005”; 1 ex. (SMTD), “Gabun, Richter”, “Coll. J. Faust, Ankauf 1900”, “Staatl. Museum fur Tierkunde, Dresden”, “Rhynchites despectus m., Det. E. Voss”, “Paralectotype Rhynchites despectus Voss, A. Legalov design. 2005”; 3 ex. (MMUE), 1 ex. (SZMN), “Old Calabar”; 2 ex. (MCSN), “Gabon, W. Africa (Mocquerys)”; 1 ex. (SMTD), “Gabon, Baden”; 1 ex. (ZIN), “Gabun”, “658”, “Staudinger 1927”; 1 ex. (ISNB), “R. Dem. Congo, Luki Biosphere reserve., Mayombe, Bas Congo, 05'37'16,7'S I 13'05'54,8 E, 266 m asl, Canopy Fogging, Fogging 1, 04-11-2006, Leg. D. De Bakkers & J. P. Michiels”; 1 ex. (ISNB), “Rep. Demo. Congo, Luki Biosphere reserve., Mayombe, 05'37'167'S I 13'05'548 E, 266 m ASL, Canopy Fogging, 05.XI.2006, Leg. D. De Bakkers & J. P. Michiels”; 1 ex. (MRAC), “Musee du Congo, Sankuru: faret de Lonkala, IV1925, Lt. J. Ghesquiere”, “R. Det. 1723 Q”, “R. Det. R 3415”; 1 ex. (MRAC), “Musee du Congo, Mayumbe: Zobe, 4 an 12 - 01. 1916, R. Mayne”, “R. Det. 1723 Q”, “R. Det. R 3415”; 1 ex. (MRAC), “Musee du Congo, Kiniati - Zobe, fin 12 - 1915, R. Mayne”, “R. Det. 1723 Q”, “R. Det. R 3415”; 1 ex. (MRAC), “Musee du Congo, Sankuru: Komi, 13.03.1930, J. Ghesquiere”, “R. Det. Q 3415”, “R. Det. R 3415”, “Rhynchites platynotus m., Det. E. Voss”; 1 ex. (MRAC), “Musee du Congo, Luluabourg, P. Callewaert”, “R. Det. 1723 Q”.

Remarks. A study of material from Africa has shown that Rhynchites platynotus Voss, 1938, syn.n. is synonym to Clinorhynchites despectus (Voss, 1938).

Distribution. Cameroon, Congo, Equatorial Guinea, Gabon, Togo, Zaire.

Clinorhynchites scheitzae (Voss, 1944) (col. pl. VI: h, l)

Rhynchites scheitzae Voss, 1944: 83 Remarks. The lectotype is designated by the author - a female from the collection MRAC with labels “Holotypus”, “Coll. Mus. Congo, Mongbwalu, 07 - 1938, Mme Scheitz”, “R. Det. Z 4970”, “Rhynchites scheitzae n. sp., det. E. Voss”, “Lectotype Rhynchites scheitzae Voss, 1944 A. Legalov design. 2009”.

Distribution. Zaire.

Genus Clinorhynchidius Legalov, 2003

Clinorhynchidius flexirostris (Voss, 1938)

Rhynchites flexirostris Voss, 1938b: 151 Rhynchites collarti Voss, 1938b: 152, syn.n.

Rhynchites benitoensis Voss, 1938b: 152, syn.n.

Remarks. The lectotype of Rhynchitesflexirostris - a male from the collection ZMHB with labels “Span. Guinea, Nkolentangan, 11,07-V,08., G. Tessmann S.G.” and paralectotypes (2 males and 3 females from the collection ZMHB) were studied as well as number of specimens from different collections: a male from the collection MCSN with labels “Gabon, W. Africa (Mocquerys)”, “Museo Genova, coll. Angelo Solari (acquisto 2000)”; a

female from the collection MRAC with labels “Musee du Congo, Equateur: Boende, 1928, R.P. Hulstaert”, “R. Det. 2320 g”, “R. collarti m.”, “R. Det. AA 3415”; a female from the collection MRAC with labels “Musee du Congo, Equateur: Flandria, 25.01.1933, R.P. Hulstaert”, “R. Det.

I 3251”, “Rynchites collarti m.”, “R. Det. AA 3415”; a female from the collection MRAC with labels “Musee du Congo, Basongo, 13-30.07.1921, Dr. H. Schouteden”, “R. Det. 1723 R”, “R. Det. 2320 g”, “R. collarti m., Hustache det.”, “R. Det. AA 3415”, “Rhynchites flexirostris m. v. collarti, Det. E. Voss”. Holotype of Rh. benitoensis was studied by the author - a female from the collection SMTD with labels “Benito”, “Samml. K. F. Hartmann, Ankauf, 1941”, “Staatl. Museum fur Tierkunde, Dresden”, “Typus”, “Rhynchites benitoensis n. sp., Det. E. Voss”, “Holotype Rhynchites benitoensis Voss, A. Legalov design. 2005” and one more specimen - a female from the collection MRAC with labels “Coll. Mus. Congo, Lokandu, 1937, Lt. Maree”, “R. Det. 22. 4066”, “Rhynchites benitoensis m., Det. E. Voss”. Study of type specimens and materials from Africa revealed that Rhynchites collarti Voss, 1938, syn.n. and Rh. benitoensis Voss, 1938, syn.n. are synonyms to Clinorhynchidius flexirostris (Voss, 1938). This species is reported for the first time for the fauna of Gabon. Distribution. Equatorial Guinea, Gabon, Guinea, Zaire. Genus Afrorhynchites Legalov, 2003 Subgenus Afrovolvulus Legalov, 2004

Afrorhynchites (Afrovolvulus) villosus (Boheman, 1845), comb.n., placem.n. (col. pl. VI: n-o)

Rhynchites villosus Boheman, 1845: 365 Rhynchites bipubescens Hustache, 1929a: 499, syn.n. Rhynchites rhodesianus Voss, 1938b: 156 Rhynchites methneri Voss, 1938b: 137 Rhynchites natalensis Voss, 1938b: 159 Afrovolvulus katonensis Legalov, 2004c: 64 Remarks. The lectotype is designated by the author - a female from the collection NHRS with labels “Caffraria”, “J. Wahlb.”, “Typus”, “Rhynchites villosus”, “villosus Boh.”, “7387 E91 +”, “Lectotype Rhynchites villosus Boheman, 1845, A. Legalov design. 2009”. Paralectotype - a female (NHRS) with labels “59”, “Paratypus”, “Mus. R. Holm., Capib. sp., Wahlberg”, “Paralectotype Rhynchites villosus Boheman, 1845, A. Legalov design. 2009”. Previously, this species has been wrongly placed in Pararhynchites Legalov, 2003. Study of type specimens of Rhynchites villosus Boheman, 1845 and material from Africa proved that Rhynchites bipubescens Hustache, 1929, syn.n. is synonym to Afrorhynchites villosus (Boheman, 1845). Distribution. Botswana, Kenya, Mozambique, Namibia, S-Africa, Tanzania, Zimbabwe.

Genus Proinvolvulus Legalov, 2003

Proinvolvulus rugosipennis (Voss, 1938) (col. pl. VII: a-b, d, f)

Rhynchites rugosipennis Voss, 1938b: 141 Rhynchites semiopacus Voss, 1939c: 53, syn.n.

Remarks. The holotype of Rhynchites rugosipennis was studied: a female from the collection ZMHB with labels “Span. Guinea, Nkolentangan, 11.07.-V08., G. Tessmann

S. G.”, “Rhynchites Metarhynchites rugosipennis n. sp.”, “HOLOTYPUS Rhynchites rugosipennis Voss,

1938 labelled by MNHUB 2009”, “Holotype Rhynchites rugosipennis Voss, 1938, A. Legalov det. 2009”. The lectotype of Rhynchites semiopacus is designated by the author - a male from the collection MRAC with labels “Paratypus”, “Musee du Congo, Haut - Uele: Watsa, 1922, L. Burgeon”, “R. Det. GG 3415”, “Lectotype Rhynchites semiopacus Voss, A. Legalov design. 2009”. Paralectotype - a female from the collection MRAC with labels “Holotypus”, “Musee du Congo, Mayumbe: Kiniati, 7-.06.1911, R. Mayne”, “R. Det. GG 3415”, “Rhynchites semiopacus n. sp., Det. E. Voss”, “Paralectotype Rhynchites semiopacus Voss, A. Legalov design. 2009”. All studied materials belong to one species. Voss [1938b, 1939c] put the described species in different subgenera.

Distribution. Guinea, Zaire.

Proinvolvulus flandriensis Legalov, sp.n. (col. pl. VII: c, e) Holotype. Female (MRAC), “Coll. Mus. Congo, Tshuapa: Flandria, 09.1946 - 08.1947, Rev. P. Hulstaert”,

“Rhynchites (Metarhynchites) rugosipennis Voss, Det. from deser. G.A.K. Marshall”.

Description. Female: Body black, with short dark semierect setae. Rostrum long, 5.67 times longer than wide, 1.21 times longer than pronotum, curved, thin, with carina from frons to place of antennal attachment, weakly widened to apex, lustrous, sparsely punctate. Antennae located before the rostrum middle. Eyes large, very weakly convex. Frons convex, almost matte, finely punctate. Vertex convex, finely rugosely punctate. Temples short. Antennae thin and long, reaching apical margin of pronotum. Scapus and 1st segment of funicle oval, almost equal in length. 2nd segment narrower and shorter than 1st segment. 3rd segment hardly shorter than 2nd segment. 4th segment almost equal to 3rd segment. 5th-7th segments short trapezoid. Clava long, shorter than funicle, almost compact. 1st segment longer than 2nd segment. 2nd segment wide trapezoid. 3rd segment pointed, longer than 1st segment. Pronotum almost trapezoid, of almost equal length and width. Sides almost straight. Disk convex, finely and densely rugosely punctate, with middle stria. Scutellum trapezoid. Elytra almost rectangular, 1.23 times longer than wide. Greatest width behind the middle. Humeri weakly smoothed. Intervals narrow, flat, densely punctate. Striae distinct. Points in them large and sparse. 9th stria merge with 10th at level of metacoxa. Thorax punctate. Metepisternum narrow. Abdomen convex, finely punctate. 1st and 2nd ventrites long. 3rd ventrite shorter. 4th ventrite shorter than 3rd ventrite. 5th ventrite very short. Pygidium convex, punctate. Legs long. Femora widened. Tibiae almost straight, long, slightly widened to apex. Tarsi long. 1st segment triangular. 2nd segment wide-triangular. 3rd segment bilobed. Clausal segment elongated. Claws with teeth. Length of body: 3.2 mm.

Diagnosis: The new species is similar to Proinvolvulus rugosipennis but differs with larger body, wider pronotum and longer rostrum.

Etymology: The name is derived from the location “Flandria” - “flandriensis”.

Distribution. Zaire.

Genus Teretriorhynchites Voss, 1938 Subgenus Aphlorhynchites Sawada, 1993

Teretriorhynchites (Aphlorhynchites) pubescens (Fabricius, 1775) (col. pl. VII: g)

Curculio pubescens Fabricius, 1775: 131 Rhynchites parellinus Gyllenhal, 1833: 224 Rhynchites multipunctatus Bach, 1858: 372 Rhynchitesparallinus var. fallax Starck, 1889: 55 Rhynchites parallelus var. flectirostris Pic, 1926: 9 Rhynchites pubescens ssp. byctiscoidiceps Voss, 1938b: 143 Remarks. The lectotype for Rhynchites parallinus var. fallax is designated by the author - a male from the collection HNHM with labels “Novorossijsk”, “Caucas occid., Novoros., 6.V.78, Starck”, “Holotypus 1889, Rhynchites parallinus Gyll. var. fallax Starck”, “Rhynchites parallinus Gyll. v. fallax Starck”, “Coll. Reitter”, “Lectotype Rhynchites parallinus var. fallax Starck, 1889, A. Legalov design. 2009”.

Distribution. Western and Central Palaearctic.

Genus Parinvolvulus Legalov, 2003 Subgenus Nigroinvolvulus Legalov, 2003

Parinvolvulus (Nigroinvlovulus) apionoides (Sharp, 1889)

Rhynchites apionoides Sharp, 1889a: 67 Remarks. Specimens studied: 2 females from the collection BMNH with labels “Type, H.T.”, “Japan, G. Lewis”, “Sharp Coll., 1905-313”, “Rhynchites apionoides, Type D.S., Mijanoshita, 22.12.90, Lewis”.

Distribution. China, Japan, Russia.

Genus Heterorhynchites Voss, 1932 Subgenus Sawadaia Alonso-Zarazaga & Lyal, 1999

Heterorhynchites (Sawadaia) alcyoneus (Pascoe, 1875) (col. pl. VII: h)

Rhynchites alcyoneus Pascoe, 1875: 391 Rhynchites argutus Faust, 1882: 288, syn.n.

Material. 1 ex. (ISNB), “Ind. bor, Bacon”, “Coll. Castelnau, Coll. Roelofs”; 1 ex. (SZMN), “Himalaya, Sikkim”. Remarks. The lectotype is designated by the author - a female from the collection BMNH with labels “Type”, “India”, “Rhynchites alcyoneus Pasc., Type”, “Pascoe Coll., 93-60”, “Lectotype Rhynchites alcyoneus Pascoe, A. Legalov design. 2009”. The holotype of Rhynchites argutus was studied: a female from the collection SMTD with labels “brown small square”, “Darjeeling, Ribbe”, “Coll. J. Faust, Ankauf 1900”, “Staatl. Museum fur Tierkunde, Dresden”, “Type”, “argutus Faust”, “Holotype Rhynchites argutus Fst., A. Legalov design. 2005”.

Study of type specimens revealed that Rhynchites argutus Faust, 1882, syn.n. is synonym to Heterorhynchites (Sawadaia) alcyoneus (Pascoe, 1875).

Distribution. East India.

Heterorhynchites (Sawadaia) pruinosus (Voss, 1938), placem.n. (col. pl. VIII: k)

Rhynchites pruinosus Voss, 1938b: 167 Remarks. The lectotype is designated by the author - a female from the collection BMNH with labels “Holotype”, “Gng. Leo, Ned. Timor, 2 4000, Doherty, 11. 12.”, “G.A.K. Marshall Coll., B.M. 1950-255”, “Rhynchitespruinosus n. sp., Det. E. Voss”, “Lectotype Rhynchites pruinosus Voss, A. Legalov design. 2009”. Previously, this species has been placed wrongly in the subgenus Eosawadaia Legalov,

2004.

Distribution. Timor.

Heterorhynchites (Sawadaia) azureus (Olivier, 1807)

Rhynchites azureus Olivier, 1807: 23 Material. 1 ex. (ZMAN), “Nederlands Indie, W Java, J.M.A. v. Groenendael”; 1 ex. (ZMAN), “Nederlands Indie, W Java, Djampang Tengah, 10.12.1939, J.M.A. v. Groenendael”; 1 ex. (ZMAN), “Java or Sumatra”; 1 ex. (ZMAN), “Native Collectors.”, “Java, Buitenzong Gg. Pantjar, 800 m, 14.10.-5.11.1897”; 1 ex. (ZMAN), “Indonesia, Sumatra, Lauttador, 28.07.1949, C. v. Nidek”;

1 ex. (ZMAN), “Java, Babakan, 03.1911”, “coll. F.C. Drescher”; 1 ex. (MCSN), “Java orient., Montes Tengger,

2000, 1890, H. Fruhstorfer”; 1 ex. (MCSN), “Tji Salak, Wynkoopsbaai (Grelak)”; 2 ex. (MCSN), “Java occident, Sukabumi, 2000, 1893, H. Fruhstorfer”; 1 ex. (ZMUC), “J. Skovgaard, Java”, “Java 1905”; 1 ex. (ISNB), “Java”, “Javanus Buquet.”, “Rhynchites javanus Buquet, h. Java, D. Buquet”; 1 ex. (ISNB), “Java, Soekaboemi”; 1 ex. (ISNB), “Java, Soekaboemi, Coll. Le Mount”; 1 ex. (SZMN), “Java, Mt. Gedeh”.

Distribution. Indonesia (Java, Sumatra).

Heterorhynchites (Sawadaia) wahnesi (Hartmann, 1899)

Rhynchites wahnesi Hartmann, 1899: 24

Material. 1ex. (SZMN), Borneo, Sabah, env. Keningau,

05.1993.

Remarks. This species is reported for the first time for the fauna of Sabah.

Distribution. Indonesia (Kalimantan), Malaysia (Sabah).

Subgenus Eosawadaia Legalov, 2004

Heterorhynchites (Eosawadaia) philippensis (Chevrolat, 1841)

Rhynchites philippensis Chevrolat, 1841: 224 Material. 1 ex. (ZIN), “Rhynchites coelestinus, Philipp.”;

2 ex. (ZMAN), “Philippensis Chevr., Manilla”; 2 ex. (MCSN), “Mindanao, Surigao, G. Boettcher”; 1 ex. (MCSN), “Philippinen, Mindanao, 06.1915”.

Distribution. Philippines.

Heterorhynchites (Eosawadaia) subtectus (Voss, 1938) (col. pl. VIIj: l, n)

Rhynchites subtectus Voss, 1938b: 167 Material. 1 ex. (ZIN), Thailand, Surat Thani, env. Natn. Park Khao Sok, near Phanom City, 20-28.07.1996, Gorochov; male (SZMN), Thailand, 60 km of Nakhon Sawan, banana forest, N 15.42.27, E 100.06.81, 7-17.08.2009, A.VKorshunov.

Remarks. Lectotype is designated - a male from the collection BMNH with labels “Type”, “Assam, W.F. Basdgley, 1906-185”, “Rhynchites subtectus n. sp., Det. E. Voss”, “Lectotype Rhynchites subtectus Voss, A. Legalov design. 2009”.

Distribution. East India, Thailand.

Heterorhynchites (Eosawadaia) korshunovi Legalov, sp.n. (col. pl. VII: m, VIII: a)

Material. Holotype - male (ZIN), South Vietnam, 60 km N Ho Chi Minh, env. Phu Giao vill., 3-13.10.1994, A. Napolov. Paratype: 1 female (RDP), Vietnam N, 60 km SW from Hanoi, Chua Huong, 26-29.04.1991, Jendek;

1 female (ZIN), 1 female (SZMN), Vietnam, Cat Tien,

7-21.06.1995, A. Napolov; 1 female (ZIN), Vietnam, Ma-Da forest, 1.11.1990, S. Murzin; 1 male (ISNB), Cambodia, Siem Read Prov., Preah Khan Temple, Malaise Trap, 11-

18.12.2005, Oul Yothin; 1 male (ISNB), 3 females (ISNB),

2 females (SZMN), Cambodia, Angkor Thom, day catch, 1-3.11.2005, Oul Yothin; 1 male (SZMN), Cambodia, Siem Read Prov., Angkor, Preah Khan Temple, Malaise Trap, 18-25.12.2005, Oul Yothin; 2 females (ISNB), Cambodia, Siem Read Prov., Kbal Spean, Light Trap, 28.05.2005, Var & P. Grootaet; 1 female (ZMUC), Thailand, E coast Siam Gulf, Paklua, N of Pathaya, 11-13.11.1979, Zool. Mus. Copenhagen Exped.; 1 female (ISNB), Laos, 31.05.1915, R. Vitaljs de Salvaza; 1 female (ISNB), “Coll. Castelnau, Coll. Roelofs”; 1 female (ZMUM), S Vietnam, 120 rm NNE Ho Chi Minh, env. Cat Tien Nat. Park, 28.05.2005,

D. Fedorenko.

Description. Body dark with dark blue lustre, with short pale semierect setae. Male. Rostrum very long, strongly curved in topmost third, thin, with carina, slightly widened to apex, densely punctate. Antennae located before the middle of rostrum. Eyes large, weakly convex. Frons convex, punctate. Vertex convex, punctate. Temples short. Antennae thin and long, reaching the apical margin of pronotum. Scapus and 1st segment of funicle oval. Scapus longer than 1st segment. 2nd segment very long, longer than 1st segment. 3rd segment hardly shorter than 2nd segment. 4th segment shorter than 3rd segment. 5th segment shorter. 6th and 7th segments weakly trapezoid, shorter. 7th segment shorter than 6th segment. Clava short, almost compact. 1st and 2nd segments almost identical, wide. 3rd segment pointed, narrower and longer than 2nd segment. Pronotum campaniform. Sides weakly rounded. Pronotal groove wide. Greatest width near the basis. Disk convex, finely punctate. Scutellum trapezoid. Elytra almost rectangular. Greatest width in the middle. Humeri weakly smoothed. Intervals wide, weakly convex, punctate. Striae thin, with small points. 9th stria merge with 10th stria in the middle of elytra. Thorax punctate. Metepisternum almost wide. Abdomen convex, with small points. 1st and 2nd ventrites wide. 3rd ventrite narrower. 4th ventrite narrow.

5 th ventrite very narrow. Pygidium convex, punctate. Legs long. Femora widened. Tibiae almost straight, long, weakly widened to apex. Protibiae longer and narrower than meso-and metatibiae. Tarsi long. 1st segment long triangular. 2nd segment wide triangular. 3rd segment bilobed. Clausal segment elongated. Claws with long teeth. Length of body: 7.8-10.1 mm. Female. Rostrum longer. Antennae located near the middle of rostrum. Eyes weaker convex. Abdomen stronger convex. Length of body: 6.0-9.5 mm.

Diagnosis: The new species is similar to H. subtectus but differs with larger size, rostrum longer, pronotum wider, antennae located closer to the middle of rostrum and with the shape of basal sclerite of endophallus.

Etymology. The new species is named in honour of A.V Korshunov.

Distribution. Cambodia, Laos, Thailand, Vietnam.

Subgenus Heterorhynchites s. str.

Heterorhynchites (Heterorhynchites) pristis (Marshall, 1924)

Rhynchites pristis Marshall, 1924: 283

Material. 2 ex. (RDP), 1ex. (SZMN), NE India, Megalaya, 1 km E of Tura, 25.30 N, 90.14 E, 500-600 m, 2-5.V2002, M. Tryzna & P. Benda.

Remarks. Lectotype is designated - a male from the collection BMNH with labels “Type”, “India”, “Sylhet, Chandakhira, J.L. Shewill”, “Pres. by Imp. Bur. Ent., Brit. Mus. 1924-51”, “Rhynchites pristis Mshl., TYPE”, “Lectotype Rhynchites pristis Marshall, A. Legalov design. 2009”.

Distribution. East India.

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Heterorhynchites (Heterorhynchites) elysius (Pascoe, 1875)

Rhynchites elysius Pascoe, 1875: 391

Material. 1 ex. (RDP), Malaysia, Kampung uku Dong,

26-30.03.2001, O. Buzga; 1 ex. (MCSN), “Perak, Malacca

(Doherty)”; 1 ex. (MCSN), “Brunei, N. Borneo”; 1 ex.

(MCSN), “Java occident, Pengalengan, 4000, 1893, H.

Fruhstorfer”.

Remarks. Lectotype is designated - a female from the collection BMNH with labels “Type”, “Sumatra”, “Rhynchites elysius Pasc., Type”, “Pascoe Coll., 93-60”, “Lectotype Rhynchites elysius Pascoe, A. Legalov design. 2009”.

This species is reported for the first time for the faunae of Malaysia, Kalimantan and Java.

Distribution. Malaysia, Indonesia (Java, Sumatra, Kalimantan).

Heterorhynchites (Heterorhynchites) subdentatus (Voss, 1938)

Rhynchites subdentatus Voss, 1938b: 168 Remarks. Specimen studied: a male from the collection BMNH with labels “Private Collected.”, “SARAWAK: Mt. Dulit, 4,000 ft., Moss forest, 16.10.1932”, “Oxford Univ. Exp. B.M. Hobby & A.W. Moore, B.M. 1933-254”, “Rhynchites subdentatus m., Det. E Voss”.

This species is reported for the first time for the fauna of Kalimantan.

Distribution. Indonesia (Sumatra, Kalimantan).

Tribe Byctiscini Voss, 1923 Subtribe Byctiscina Voss, 1923 Genus Byctiscus C.G. Thomson, 1859

Byctiscus minimus Legalov & Liu, 2005

Byctiscus minimus Legalov & Liu, 2005: 102 Material. 4 ex. (RDP), 1 ex. (SZMN), China, border Hebei

- Nei Mongolei, road Chengde - Chifeng, 41’6 N, 118’2 E, pass 1600 m, 1-2.06.2000, J. Turna.

Distribution. Central China.

Genus Aspidobyctiscus Schilsky, 1903 Subgenus Nepalobyctiscus Legalov, 2003

Aspidobyctiscus (Nepalobyctiscus) sculpturatus (Pascoe,

1875)

Rhynchites sculpturatus Pascoe, 1875: 394 Aspidobyctiscusgiganteus Legalov, 2003a: 332, syn.n. Material. Lectotype (BMNH), “Type”, “Rhynchites sculpturatus female Pasc.”, “India”, “Rhynchites sculpturatus Pasc.”, “Pascoe coll. 93-60”; holotype (ZIN), “Nepal, Expedition A. Konstantinov, S. Lingafelter, M. Volkovich 2000”, “Loc.no. 17, RASUWA,

Ka.ika.stha.n-Ra.mche, 27’59’60’N, 85’12’15’E, 15202100 m, 28.04.2000”, “Holotypus Aspidobyctiscus (Nepalobyctiscus) giganteus Legalov, sp.n.”; 2 ex. (ISNB), “Ind.Bor. Bacon”, “Coll. Castelnau, Coll. Roelofs”; 2 ex. (RDP), “NEPAL, Mustang distr., Gasa-Kalopani, 20002500 m, 20.IV.1986, CHs. leg.”; 1 ex. (SZMN), “NEPAL, Mustang distr., Kalopani, 2600 m, 21-25.06.1986, CHs. leg.”; 1 ex. (DEI),”Sikles Mts. upp. Garlang 2000 m, 29.7.”, “Nepal, Himalaya, Annapurna Mts., lg. Schmidt 1995”; 1 ex. (CKJU), “N India: Uttaranchal state, ca 30 km N of Bageshwar, Khati vill. env., Z. Kejval & M. Tryzna leg.”.

Remarks. Study of type specimens and materials from Nepal and India showed that Aspidobyctiscus giganteus Legalov, 2003, syn.n. is synonym to Aspidobyctiscus (Nepalobyctiscus) sculpturatus (Pascoe, 1875). Distribution. North India, Nepal.

Family Attelabidae Billberg, 1820 Subfamily Attelabinae Billberg, 1820 Supertribe Attelabitae Billberg, 1820 Tribe Euopsini Voss, 1925 Subtribe Suniopsina Legalov, 2003 Genus Suniops Voss, 1928 Subgenus Vietsuniops Legalov, 2007

Suniops (Vietsuniops) gorochovi Legalov, 2003

Suniops gorochovi Legalov, 2003a: 367

Material. 1 ex. (APB), Thailand, Do Inthanon N.P., Pha

sum Waterfall, 30.X.2004, A. Ozoroz.

Remarks. This species is reported for the first time for the fauna of Thailand.

Distribution. Vietnam, Thailand.

Subtribe Synaptopsina Legalov, 2003 Genus Pseudoeuops Legalov, 2003

Pseudoeuops clarus (Sawada & Morimoto, 1985)

Euops clarus Sawada & Morimoto, 1985: 180 Material. 1 ex. (HNHM), Taiwan, Nantou coutv., Kao-Leng Dyi, 18 km W of Wushe, 24.4.561 N, 121.8.046 E, 1945 m, swept from vegetation, 18-19.04.2002, D. Anstine, Gy. Faban, O. Merkl.

Distribution. China (Taiwan).

Subtribe Parasynaptopsisina Legalov, 2007 Genus Kobusynaptops Kano, 1927

Kobusynaptops verrucosus Legalov & X. Zhang, 2007

Kobusynaptops verrucosus Legalov & Zhang, 2007: 231 Material. 1 ex. (ZMUM), China, W Sichuan, Li Xian, 2500 m, 20.06.2004, V. Patrikeev.

Remarks. This species is reported for the first time for the fauna of Sichuan.

Distribution. China (Sichuan, Yunnan).

Genus Pseudosynaptops Legalov, 2003

Pseudosynaptops barbieri (Marshall, 1953)

Euops barbieri Marshall, 1953: 91 Material. 1 ex. (APB), Thailand, Pachin prov., Bun, Sakaerat Environmental Research Station, 1-16.06.2001, E. Horvath, Gy. Sziraki.

Remarks. This species is reported for the first time for the fauna of Thailand.

Distribution. Vietnam, Thailand.

Genus Riedeliops Alonso-Zarazaga & Lyal, 2002 Subgenus Riedeliopsis Legalov, 2007 Riedeliops (Riedeliopsis) zolotarenkoi Legalov, 2003

Riedeliops zolotarenkoi Legalov, 2003a: 386 Material. 1 ex. (TAUI), Thailand, 105 0m, Chiang Mai Prov., Doi Inthanon N. P., Mae Pan waterfall, 27.05.2004, N 18’31, E 98’37, Ilan Yarom.

Remarks. This species is reported for the first time for the fauna of Thailand.

Distribution. Vietnam, Thailand.

Subgenus Vieteuops Legalov, 2007

Riedeliops (Vieteuops) darevskyi Legalov, 2003

Riedeliops darevskyi Legalov, 2003a: 385 Material. 1 ex. (ZMUM), S Vietnam, 120 rm NNE Ho Chi Minh, env. Cat Tien Nat. Park, 11.2004, D. Fedorenko. Distribution. Vietnam.

Subgenus Levoeuops Legalov, 2007

Riedeliops (Levoeuops) nepalensis Legalov, 2003

Riedeliops nepalensis Legalov, 2003a: 385 Material. 4 ex. (ZIN), 9 ex. (USNM), Nepal, Lantang Nat. Park, env. of Dhunche, 28’07’00, 85’17’00, 30.04.2000, 19000-2100m, Konstantinov, Lingafelter, Volkovitsh. Distribution. Nepal.

Riedeliops (Levoeuops) vietnamensis Legalov, 2003

Riedeliops vietnamensis Legalov, 2003a: 388 Material. 1 ex. (SZMN), Vietnam, Tam Dao, 900 m,

06.1981, L. Medvedev.

Distribution. Vietnam.

Genus Parasynaptopsis Legalov, 2003

Parasynaptopsis chinensis (Voss, 1922)

Euops chinensis Voss, 1922b: 166 Euops chinensis f. purpurea Kano, 1927: 39 Material. 1 ex. (VRP), China, SW Anhui, Tianzhushan env., 30.75 N, 116.45 E, 11-14.05.2004, V. Ryjacek; 1 ex. (VRP), China, W Anhui, Tianzhushan, 05.2004, V. Ryjacek; 1 ex. (VRP), China, Hubei, Lucongpo, 05.06.2004, V Ryjacek.

Remarks. This species is reported for the first time for the fauna of Anhui.

Distribution. East Asia.

Parasynaptopsis cuprifulgens (Voss, 1942)

Euops cuprifulgens Voss, 1942b: 102

Material. 1 ex. (DEI), China, Guangdong prov., Tsuholin

Nat. Park, 21.06.1990, Kushetzov.

Distribution. China (Fujian, Guangdong). Parasynaptopsis lespedezae koreanus (Voss, 1924)

Euops splendens f. koreana Voss, 1924: 43 Euops lespedezae f cuprinipennis Voss, 1941b: 118 Material. 1 ex. (VRP), China, E. Hubei, Dabie Shan, Wujashan forest park, 31.1 N, 115.8 E, 7-10.05.2004, V. Ryjacek; 1 ex. (ZMUM), “China, Nankin, 07.1934, N.N. Filippov”; 5 ex. (ZMHB), “China, 1956, 98, Honan-Shantung, Prof. Zimmermann”; 2 ex. (ZMHB), “China, 1956, 109, Honan-Shantung, Prof. Zimmermann”; 1 ex. (ZMHB), “China, 1956, 110, Honan-Shantung, Prof. Zimmermann”.

Remarks. This species is reported for the first time for the fauna of Hubei.

Distribution. Eastern Asia.

Parasynaptopsis lespedezae lespedezae (Sharp, 1889)

Euops lespedezae Sharp, 1889a: 55 Euops splendens f. unicolorata Voss, 1924: 43 Material. 1ex. (HNHM), Ichinotani, Suma-Kobe, Japan, 19.06.32, J. Fodor; 1 ex. (MZLU), Japan, Osaka, Mt. Iwowaki, 15.09.1946, S. Ueono.

Distribution. Japan.

Parasynaptopsis nigrum (Kano, 1927)

Euops splendens f. nigra Kano, 1927: 39

Attelabus splendens Roelofs, 1874: 139 [non Gyllenhal,

1839]

Euops splendidus Dalla Torre & Voss, 1930: 56 [RN] Material. 1 ex. (NMPC), Japan, Kyoto pref., Kyoto city, 23.V1978, W. Sunzuki; 3 ex. (ISNB), Gifu pref., Suhara Hondo, 16.05.-20.07.54; 9 ex. (ISNB), Gifu pref., Suhara Hondo, 05.-07.56; 1 ex. (ZIN), Central Japan, Hiroyu, 21.07.1964, H. Ohira; 1 ex. (ISNB), “Mt. Izugatake, Saitama-ken, May-5, 1952, Coll. Masaru Ohtake”; 1 ex. (DEI), Japan, Yunohana spa., Fukushima Pref., 14.06.1990; 1 ex. (HNHM), Ichinotani, Suma-Kobe, Japan, 19.06.32, J. Fodor; 1 ex. (MZLU), Japan, Osaka, Mt. Iwowaki,

15.09.1946, S. Ueono; 1 ex. (MZLU), Japan, Kohara, Shiga Pref., 24.05.1952, S. Ueono.

Distribution. Japan.

Genus Parasynatops Legalov, 2003 Subgenus Parasynatops s. str.

Parasynatops (Parasynatops) beijingensis Legalov, 2003

Parasynatops beijingensis Legalov, 2003a: 379 Material. 1 male (APB), China, Beijing, Mentougou Distr., Beijing, 130 km NW Xiaolongmen station, 1905 m, 39.59’, 115.31’, 28.07.2002, G. Melika, Gebusch, Trockenrasen. Remarks. This species has been previously wrongly placed in the genus Sawadaeuops Legalov, 2003 [Legalov, 2007]. Distribution. China (Beijing, Hebei).

Parasynatops (Parasynatops) konoi (Sawada & Morimoto, 1985)

Euops konoi Sawada & Morimoto, 1985: 181 Material. 1 ex. (MMUE), “Mt. Takao, Musashi, Japan, 13-

8-1926”.

Distribution. Japan, Russia (Kuril Isl.).

Subgenus Neparasynatops Legalov, 2007

Parasynatops (Neparasynatops) championi (Voss, 1929)

Euops championi Voss, 1929c: 214 Material. 3 ex. (HNHM), 1 ex. (SZMN), Taiwan, Nantou coutv., Kao-Leng Dyi, 18 km W of Wushe, 24.4.561 N, 121.8.046 E, 1945 m, swept from vegetation, 18-

19.04.2002, D. Anstine, Gy. Faban, O. Merkl. Distribution. South-eastern Asia.

Parasynatops (Neparasynatops) moanus Legalov, 2003 Parasynatops moanus Legalov, 2003a: 382 Material. 1 ex. (CJPM), China, Yunnan, Yulongsue Shan, 3300-3900 m, 14-19.06.1996, S. Murzin.

Remarks. This species is reported for the first time for the fauna of Yunnan.

Distribution. China (Sichuan, Yunnan).

Subtribe Sawadaeuopsina Legalov, 2007 Genus Sawadaeuops Legalov, 2003 Subgenus Sawadaeuops s. str.

Sawadaeuops (Sawadaeuops) centralchinensis Legalov

& Liu, 2005

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Sawadaeuops centralchinensis Legalov & Liu, 2005: 122 Material. 1 ex. (RDP), China, Shaanxi, Qing Ling Shan mts., road Baoji - Taibai vill., pass 35 km S of Baoji, 21-

23.06.1998, O. Safranek, M. Tryzna.

Distribution. China (Hubei, Shaanxi).

Sawadaeuops (Sawadaeuops) punctatostriatus (Motschulsky, 1860)

Atellabus punctatostriatus Motschulsky, 1860: 22 Euops phaedonius Sharp, 1889a: 56 Euopspuncticollis Schilsky, 1906: 92 [non Boheman, 1858] Euops schilskyi Voss, 1922c: 174 [RN]

Euops aceri Kano, 1926: 223 Euops punctatostriata f awana Kano, 1927: 39 Material. 20 ex. (ZMUC), “Kamikochi, 500 ft, Northern Alps, Japan, 14.6.1939, E. Suenson”; 1 ex. (DEI), Japan, Yamanashi Pref., Fujiyama N., Subaru-line, 2000 m,

1.08.1999, V Puthz; 2 ex. (DEI), Japan, Maruseppu, Hokkaido, 8.07.1987, S. Ohmomo; 2 ex. (DEI), Japan, Mts. Daibosatsu, Yamanashi Pref., 29.07.1987, H. Akiyama; 1 ex. (HNHM), Ichinotani, Suma-Kobe, Japan, 19.06.32, J. Fodor; 2 ex. (MZLU), Japan, Osaka, Mt. Iwowaki,

15.09.1946, S. Ueono.

Distribution. Japan, Korea, Russia (Sakhalin, Kuril Isl.).

Subtribe Euopsina Voss, 1925 Genus Euops Schoenherr, 1839

Euops falcatus (Guerin-Meneville, 1833)

Attelabus falcatus Guerin-Meneville, 1833: 137 Euops australasie Fahraeus, 1839: 319 Material. 1 ex. (ACD), Australia, N.S.W., 17 km SE Bombla, Wog Wog, 02.1994, Margueles.

Distribution. Australia.

Tribe Euscelini Voss, 1925 Subtribe Euscelina Voss, 1925 Genus Alleuscelus Voss, 1937 Subgenus Alleuscelus s. str.

Alleuscelus (Alleuscelus) violaceipennis Voss, 1937

Alleuscelus violaceipennis Voss, 1937: 159 Remarks. Lectotype was studied - a male from the collection BMNH with labels “Type”, “Peru”, “56018”, “Fry coll. 1905.100”, “Euscelus violaceipennis sp.n., Det.

E. Voss”.

Distribution. Peru.

Subgenus Paralleuscelus Legalov, 2004, stat.n.

Alleuscelus (Paralleuscelus) deletangi (Hustache, 1924), comb.n. (col. pl. VIII: b, e, i)

Euscelus deletangi Hustache, 1924: 170 Euscelus insignis Voss, 1925: 39

Remarks. Lectotype was designated - a male from the collection DEI with labels “Bolivien, Germain”, “Coll. Kraatz”, “Syntypus”, “Euscelus insignis”, “Euscelus insignis n. sp., Det. E. Voss”, “Dtsch. Entomol. Institut

Berlin”, “coll. DEI Muncheberg”, “Euscelus insignis Voss”, “Lectotype Euscelus insignis Voss, 1925, A. Legalov design. 2009”.

Distribution. Bolivia.

Tribe Hybolabini Voss, 1925 Genus Omolabus Jekel, 1860 Subgenus Asternolabus Legalov, 2007

Omolabus (Asternolabus) callosus (Sharp, 1889)

Attelabus callosus Sharp, 1889b: 7

Material. 1 ex. (ACD), CA, Guat. Baja Verap., 3 m E

Purulha, 5000', 16-17.10.2006, W.H. Tyson.

Distribution. Costa Rica, Mexico, Guatemala, Honduras, Nicaragua, Panama.

Subgenus Sternolaboides Legalov, 2007 Omolabus (Sternolaboides) bowringi Voss, 1938 (col. pl. VIII: d, l-n)

Omolabus bowringi Voss, 1938c: 157 Remarks. The lectotype was designated by the author -a male from the collection HNHM with labels “Brasil, Sao Paulo”, “Bowring 63 47”, “Bates Biras”, “Paratypus Omolabus bowrimgi Voss”, “Paratypus Omolabus bowringi m.”, “Omolabus bowringi m.”, “Lectotype Omolabus bowringi Voss, 1938, A. Legalov design. 2009”. Distribution. Brazil.

Subgenus Pseudomolabus Legalov, 2004

Omolabus (Pseudomolabus) centomyrciae (Voss, 1925), placem.n. (col. pl. VIII: c, j-k)

Xestolabus centomyrciae Voss, 1925: 268 Remarks. The lectotype was designated - a male from the collection HNHM with labels “Paraguay, San Berhardino, Fiebrig”, “Paratypus Xestolabus centomyrciae Voss”, “Paratypus Xestolabus centomyrciae m.”, “Lectotype Xestolabus centomyrciae Voss, 1925, A. Legalov design. 2009”. This species has been wrongly placed previously in the subgenus Paralabus Legalov, 2004.

Distribution. Paraguay.

Subtribe Hybolabina Voss, 1925

Genus Hybolabus Jekel, 1860

Hybolabus amazonicus Voss, 1925 (col. pl. VIII: f-h) Hybolabus amazonicus Voss, 1925: 193 Remarks. The lectotype was designated by the author - a female from the collection DEI with labels “Amasonas”, “Coll. Kraatz”, “Syntypus”, “Hybolabus amazonicus n. sp., Det. E. Voss”, “coll. DEI Muncheberg”, “Lectotype Hybolabus amazonicus Voss, 1925, A. Legalov design. 2009”. Paralectotypes: female (DEI) with labels

“Amasonas”, “Coll. Kraatz”, “Syntypus”, “Voss det.”, “coll. DEI Muncheberg”, “Paralectotype Hybolabus amazonicus Voss, 1925, A. Legalov design. 2009” and female (DEI) with labels “Amasonas”, “Coll. Kraatz”, “Syntypus”, “Voss det.”, “coll. DEI Muncheberg”, “Hybolabus amazonicus Voss”, “Paralectotype Hybolabus amazonicus Voss, 1925, A. Legalov design. 2009”. Distribution. Brazil.

Tribe Attelabini Billberg, 1820 Subtribe Lamprolabina Voss, 1925 Genus Lamprolabus Jekel, 1860

Lamprolabus bispinosus (Gyllenhal, 1833)

Attelabus bispinosus Gyllenhal, 1833: 204 Lamprolabus bispinosus ssp. tabangensis Voss, 1961: 244 Material. 1 ex. (ACD), Malaysia, Taiping, 09.1982. Distribution. South-eastern Asia.

Lamprolabus spiculatus (Boheman, 1845)

Attelabus spiculatus Boheman, 1845: 359

Attelabus corallipes Pascoe, 1883: 90

Lamprolabus latispinosus Voss, 1929c: 209

Material. 1 ex. (ACD), Laos, Oudomxai prov., Namo,

1-6.07.2004, Li Jingki.

Distribution. South-eastern Asia.

Lamprolabus trapezicollis (Heller, 1922)

Attelabus trapezicollis Heller, 1922b: 16

Material. 1 ex. (CKJU), N Laos, Louang Namtha distr.,

15km NW of Louang Namtha, 5.-11.05.1997, Strba &

Hergovitsch.

Distribution. Laos, Myanmar, Thailand.

Subtribe Paramecolabina Legalov, 2003 Genus Catalabus Voss, 1925 Subgenus Catalabus s. str.

Catalabus (Catalabus) quadriplagiatus (Voss, 1953)

Paramecolabus quadriplagiatus Voss, 1953: 49 Remarks. The lectotype was designated by the author -a female from the collection ZFMK with labels “Kuatun (2300 m), 27.40n. Br., 117.408 L., J. Klapperich, 6.08.1938”, “Type”, “Paramecolabus quadriplagiatus n. sp.”, “Lectotype Paramecolabus quadriplagiatus Voss, 1953, A. Legalov design. 2009”.

Distribution. China (Fujian, Hunan).

Subtribe Isolabina Legalov, 2007 Genus Isolabus Voss, 1925

Isolabus indigaceus (Pascoe, 1883)

Attelabus indigaceus Pascoe, 1883: 90 Material. 4 ex. (MCSN), Laos, Muong Qu, Vitalis. Distribution. Laos, Vietnam.

Isolabus jekeli Legalov, 2002

Isolabus jekeli Legalov, 2002: 92 [RN]

Attelabus caeruleus Jekel, 1860: 202 [non Fabricius, 1798] Material. 1 ex. (CBN), China, Fujian rp., Shaowu, 27.06.1991.

Distribution. Eastern Asia.

Isolabus magnus Voss, 1925

Isolabus magnus Voss, 1925: 215

Attelabus longicollis Fairmaire, 1894: 222 [non Fabricius, 1801]

Material. 1 ex. (ACD), China, Sichuan, 50 km E of Chengkoi, 1900 m, 5.07.1995.

Distribution. Eastern and South-eastern Asia.

Subtribe Henicolabina Legalov, 2007 Genus Henicolaboides Legalov, 2007

Henicolaboides ruficeps (Voss, 1948)

Henicolabus haematideus ssp. ruficeps Voss, 1948: 159 Remarks. The lectotype was designated by the author - a male from the collection ZFMK with labels “Kuatun (2300 m), 27.40n. Br., 117.408 L., J. Klapperich, 27.04.1938”,

“Type”, “Henicolabus haematideus m. f n. ruficeps”,

“Lectotype Henicolabus haematideus ssp. ruficeps Voss, 1948, A. Legalov design. 2009”.

Distribution. China (Fujian, Hunan).

Henicolaboides sapansis Legalov, 2007 Henicolaboides sapansis Legalov, 2007: 284 Material. 1 ex. (ACD), N Vietnam, Sa pa, 06.2001. Distribution. Vietnam.

Genus Allolabus Voss, 1925 Subgenus Jekelilabus Legalov, 2003

Allolabus (Jekelilabus) octomaulatus (Jekel, 1860)

Attelabus octomaulatus Jekel, 1860: 190

Attelabus octospilotus Jekel, 1860: 201

Material. 2 ex. (CKJU), S India, Kerala state, Kallar env.,

30 km NE of Trivandrum valley of river Kallar, 300-500

m, 7-13.05.1999, Kejval, Tryzna.

Distribution. South Asia.

Subgenus Allolabus s. str.

Allolabus (Allolabus) lewisi (Sharp, 1889)

Attelabus lewisi Sharp, 1889a: 53 Henicolabus lewisi var. maculatus Kano, 1927: 36 Material. 1 ex. (MZLU), Japan, Osaka, Mt. Minoo, 30.03.1949, S. Ueno; 1 ex. (ISNB), Japan, Tochigi Pref., Kaiko-bashi, Shiobara-machi, 14.05.1988, S. Ohmomo. Distribution. Eastern Asia.

Subgenus Eoallolabus Legalov, 2003

Allolabus (Eoallolabus) geniculatus (Heller, 1908)

Attelabus unoformis var. geniculatus Heller, 1908: 155 Material. 1 ex. (MCSN), “Borneo, Drunoc, Pall”. Distribution. Indonesia (Kalimantan).

Allolabus (Eoallolabus) javensis (Voss, 1961) Henicolabus uniformis ssp. javensis Voss, 1961: 243 Material. 2 ex. (MCSN), Java occident, Sukabumi, 2000, 1893, H. Fruhstorfer; 1 ex. (MCSN), Java orient, Montes Teneger, 4000, 1890, H. Fruhstorfer.

Distribution. Indonesia (Java).

Tribe Lagenoderini Voss, 1925 Subtribe Phymatopsinina Legalov, 2003 Genus Phymatopsinus Voss, 1925

Phymatopsinus pustula (Ancey, 1881)

Attelabus pustula Ancey, 1881: 469 Apoderus dromedarius Faust, 1883: 472 Phymatopsinus pustula ssp. affinis Voss, 1939c: 56 Material. 2 ex. (ZMHB), “Uamgebiat, Bosum, 1-10.04.14, Tessmann S.”; 1 ex. (MCSN), “Zanguebar, V. de Poll”; 1 ex. (MCSN), “Zanzibar”; 1 ex. (ISNB), “Congo Francais, Fort Crampel”, “Coll. on Le Moult Naturaliste, Paris”; 1 ex. (ISNB), “Fort Crampel, Congo Francais”, “Coll. K.”; 1 ex. (ISNB), 1ex. (SZMN), “Rhodesie du Nord, Mweru -Wantipa, 22.01.1944, H.J. Bredo”; 1 ex. (ISNB), “Rhodesie du Nord, Abercorn, 4.06.1944, H.J. Bredo”; 1 ex. (TAUI), Malawi, South Zomba Plateau, Changwa Dam., 3.10.1998,

F. Kaplan et A. Freidberg; 1 ex. (ACD), Tanzania, Wamuia area, Mivomelo distr., Morogoro, 3.01.2007, C. Jeromo. Distribution. Congo, Guinea, Malawi, Mozambique, Senegal, Tanzania, Zaire, Zambia, Zimbabwe.

Subtribe Pleurolabina Legalov, 2003

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Genus Apleurolabus Legalov, 2007

Apleurolabus evanescens (Voss, 1928), comb.n., placem.n.

Anisolabus evanescens Voss, 1928: 113

Remarks. This species has been wrongly placed previously

in the genus Pleurolabus Jekel, 1860.

Distribution. S-Africa.

Apleurolabus spectator (Marshall, 1932)

Attelabus spectator Marshall, 1932: 2

Material. 1 ex. (SMWN), “Nyika Nat. Park, Malawi, SE

1033 pd, 6.12.1986, E. Holm, E. Marais”.

Distribution. Malawi, Zimbabwe.

Subtribe Lagenoderina Voss, 1925 Genus Lagenoderus White, 1841 Subgenus Lagenoderus s. str.

Lagenoderus (Lagenoderus) fairmairei Hustache, 1922 (col. pl. VIII: p)

Lagenoderus fairmairei Hustache, 1922: 418 Material. 2 males (MRAC), “Madagascar: Fempanambo,

03.1961, J. Vadon”, “Lagenoderes fairmairei Hust., E. Voss det. 1966”; male (ISNB), “Madagascar, Antsihanaka, Leg Perrot I. G. 18.293, ex. Coll. Oberthur”; male (HNHM), “Madagascar, Ambaton dralaka”.

Distribution. Madagascar.

Lagenoderus (Lagenoderus) dentipennis (Gyllenhal, 1839) (col. pl. VIII: o, q-r)

Attelabus dentipennis Gyllenhal, 1839: 315 Lagenoderus gnomoides White, 1841: 183 Lagenoderus brevicollis Fairmaire, 1897: 186, syn.n. Lagenoderus coniferus Fairmaire, 1902: 382 Lagenoderus vadoni Voss, 1966: 378, syn.n.

Material. 1ex. (HNHM), “Madagascar, Ambaton dralaka”;

3 ex. (ISNB), 2 ex. (SZMN), “Madagascar, Region de Mananjary, Leg A. Mathiaux Ex. Coll. Oberthur”; 1ex. (ISMB), 1 ex. (SZMN), “Ex Museo R. Oberthur, Madagascar, Mandritsara”; 2 ex. (ISNB), 2 ex. (SZMN), “Madagascar, Antsihanaka, Leg Perrot I. G. 18.293, ex. Coll. Oberthur”; 3 ex. (ISNB), “Madagascar, 1898, Ex Oberthur”; 1 ex. (ISNB), “Madagascar, Coll. Castelnau, Coll. Roelofs”; 1 ex. (MCSN), “Fenerive, Mad., Coll. v. de Poll”.

Remarks. Type specimens of L. gnomoides were studied: a male from the collection SMTD with labels “Madagascar”, “cotype”, “Paratype”, “Samml. K. F. Hartmann, Ankauf, 1941”, “Staatl. Museum fur Tierkunde, Dresden”, “Lagenoderus gnomoides Frm.”, “Lectotype Lagenoderus gnomoides White, A. Legalov design. 2005” and a male with labels “Madagascar”, “Paratype”, “Samml. K. F. Hartmann, Ankauf, 1941”, “Staatl. Museum fur Tierkunde, Dresden”, “Paralectotype Lagenoderus gnomoides White, A. Legalov design. 2005”. The paratypes of L. vadoni were studied: a male from the collection MRAC with labels “Allotypus male”, “Coll. Mus. Congo, Madagascar: Andranofotsy, 07-1937, J. Vadon”, “male”, “Lagenoderes vadoni n. sp., E. Voss det. 1966”; a male (MRAC) with labels “Paratypus female”, “Coll. Mus. Congo, Madagascar: Atakotako, 15.01.1939, J. Vadon”, “Lagenoderes vadoni n. sp., E. Voss det. 1966”; a male (MRAC) with labels “Paratypus female”, “Coll. Mus. Congo, Madagascar:

Andranofotsy, 07.1937, J. Vadon”, “Lagenoderes vadoni n. sp., E. Voss det. 1961”. More specimens studied: a male from the collection MRAC with labels “Coll. Mus. Congo, Madagascar: Mt Sandrasoa, S. W. Maroantsetra, J. Vadon, 23.02.1939”, “Lagenoderes brevicollis Fm., E. Voss det. 1966”; a male (MRAC) with labels “Coll. Mus. Congo, Madagascar: Andranofotsy, 07.1937, J. Vadon”, “male”, “Lagenoderes brevicollis Fm., E. Voss det. 1966”; a male (MRAC) with labels “Coll. Mus. Tervuren, N.E. Madagascar: Ambodivoangy, 10.1959, J. Vadon”, “Lagenoderes brevicollis Fairm., Ferragu det.”.

The study of type specimens and large materials from Madagascar revealed that L. brevicollis Fairmaire, 1897, syn.n. and L. vadoni Voss, 1966, syn.n. were synonyms to L. (L.) dentipennis (Gyllenhal, 1839).

Distribution. Madagascar.

Subgenus Lagenoderoides Legalov, 2007

Lagenoderus (Lagenoderoides) ferrumequinum (Fairmaire, 1897)

Apoderus ferrumequinum Fairmaire, 1897: 186 Material. 4 ex. (ISNB), 1 ex. (SZMN), “Madagascar, 1898, Ex Oberthur”.

Distribution. Madagascar.

Tribe Euscelophilini Voss, 1925 Subtribe Metocalolabina Legalov, 2003 Genus Trachelolabus Jekel, 1860

Trachelolabus floridus (Zhang, 1993)

Himatolabusfloridus Zhang, 1993: 198

Euscelophilus qinni Liang, 1994: 488

Material. 1 ex. (CKJU), China, Yunnan, Canghan Mts.,

Near Dali, 2500 m, 11.06.1998, S. Murzin.

Distribution. China (Xingjian, Yunnan).

Subtribe Euscelophilina Voss, 1925 Genus Euscelophilus Voss, 1925

Euscelophilus vitalisi (Heller, 1922)

Trachelolabus vitalisi Heller, 1922a: 13 Material. 1 ex. (ACD), N Vietnam, Sa pa, 06.2001. Distribution. China (Yunnan), Vietnam, Cambodia.

Genus Euscelophilidius Legalov, 2003 Subgenus Euscelophilidius s. str.

Euscelophilidius (Euscelophilidius) gibbicollis (Schilsky, 1906)

Euscelus gibbicollis Schilsky, 1906: 90 Euscelophilus hidakai Liang, 1994: 493 Material. 1 ex. (ACD), “China, #21, from Moretto”. Distribution. Eastern Asia.

Subfamily Apoderinae Jekel, 1860 Tribe Clitostylini Voss, 1926 Subtribe Allapoderina Legalov, 2003 Genus Allapoderus Voss, 1927 Subgenus Biallapoderus Legalov, 2003

Allapoderus (Biallapoderus) rubriventris (Hustache, 1923)

Apoderus rubriventris Hustache, 1923: 151

Material. 1 ex. (ACD), Tanzania, Moro Goro PR, Wami

Vil Sekoine area, Mvomoro Distr., 19-31.03.2008, G.

Mpoyda.

Distribution. Congo, Guinea, Malawi, Tanzania, Uganda, Zaire, Zimbabwe.

Subgenus Allapoderus s. str.

Allapoderus (Allapoderus) giganteus Legalov, 2007

Allapoderus giganteus Legalov, 2007: 294

Material. 2 ex. (ACD), N Vietnam, Koang Lien,

18.05.2002.

Distribution. Vietnam.

Allapoderus (Allapoderus) bhutanensis Legalov, sp.n.

Material. Holotype - female (ACB), Bhutan, Wangdue Phodrang, 07.2005, Li Jingki.

Diagnosis. The new species is very close to Allapoderus manaliensis (Voss, 1920) but differs with wider body without metallic sheen, densely shagreen pronotum, densely punctate frons, distinct transversal-wrinkled vertex and larger teeth on femora. Body black. Length of body: 4.7 mm.

Etymology. The name is derived from the location “Bhutan” - “bhutanensis”.

Distribution. Bhutan.

Subtribe Clitostylina Voss, 1926 Genus Trachelismus Motschulsky, 1870 Subgenus Eoclitostyloides Legalov, 2007

Trachelismus (Eoclitostyloides) prolixus (Voss, 1929), stat.n.

Clitostylus tenuissimus f. prolixa Voss, 1929c: 199 Distribution. Philippines.

Tribe Hoplapoderini Voss, 1926 Subtribe Hoplapoderina Voss, 1926 Genus Agomadaranus Voss, 1958 Subgenus Agomadaranus s. str.

Agomadaranus (Agomadaranus) bihumeratus (Jekel, 1860)

Apoderus bihumeratus Jekel, 1860: 180 Paroplapoderus breviceps Voss, 1926: 43 Material. 1 ex. (ACD), Myanmar, Monghkok, Shan Hinghland, 18-23.07.2005, Li Jingki.

Distribution. Myanmar, Eastern India, Nepal.

Agomadaranus (Agomadaranus) perakensis (Voss, 1935)

Paroplapoderus perakensis Voss, 1935d: 514 Material. 1 ex. (ACD), W Malaysia, Tapah, Cameroon Highland, 02.2005.

Distribution. Malaysia.

Genus Echinapoderus Voss, 1926

Echinapoderus enoplus (Brancsik, 1893)

Apoderus enoplus Brancsik, 1893: 239 Apoderus aculeatus Faust, 1899b: 14 Apoderus aculeatus var. decolor Faust, 1899b: 14 Apoderus aculeatus ssp. ebeninus Kuntzen, 1915: 138 Echinapoderus madegassus Janczyk, 1960: 41 Material. 1 ex. (ACD), Madagascar, Antsiranana prov., Ambodidimaka env., 15-16.12.2002, E. Ambanja, I. Jenis. Distribution. Madagascar.

Genus Hoplapoderus Jekel, 1860

Hoplapoderus echinatus (Gyllenhal, 1833)

Apoderus echinatus Gyllenhal, 1833: 195

Material. 1 ex. (ACD), S India, Kumili, Tamil nadu st.,

06.1986, Trs. Nathan.

Distribution. India, Laos, Malaysia, Myanmar, Nepal, Sri Lanka, Vietnam.

Hoplapoderus hystrix (Fabricius, 1801)

Attelabus hystrix Fabricius, 1801: 419 Hoplapoderus hystrix f. penangincola Voss, 1929a: 363 Material. 1 ex. (ACD), Malaysia, Geopeng, 10.1982. Distribution. Indonesia (Java, Sumatra), Malaysia (Perak).

Tribe Trachelophorini Voss, 1926 Genus Metriotracheloides Legalov, 2008

Metriotracheloides regularis (Ter-Minassian, 1986), comb.n., placem.n.

Trachelophoridius regularis Ter-Minassian, 1986: 725 Remarks. The holotype was studied: a male from the collection ZFMK with labels “Madagascar, Antsianaka, 7.92, Sig. R. Oberthur Eing. Nr. 4, 1956”, “Museum Koenig Bonn”, “Holotypus Trachel. regularis T.-M.”, “Holotype Trachelophoridius regularis Ter-Minassian, 1986, A. Legalov det. 2009”. This species has been wrongly placed previously in genus Trachelophoridius Voss, 1929. Distribution. Madagascar.

Genus Madagasocycnelus Legalov, 2003

Madagasocycnelus humeralis (Olivier, 1807)

Apoderus humeralis Olivier, 1807: 17 Trachelophorus humeralis ssp. pygmaeus Voss, 1929b: 159 Material. 1 ex. (ACD), Madagascar, Antsiranana prov., Ambodidimaka env., 15-16.12.2002, E. Ambanja, I. Jenis. Distribution. Madagascar.

Tribe Apoderini Jekel, 1860 Subtribe Leptapoderina Legalov, 2003 Genus Heterapoderus Voss, 1927 Subgenus Pseudoheterapoderus Legalov, 2003

Heterapoderus (Pseudoheterapoderus) crenatus (Jekel, 1860)

Apoderus crenatus Jekel, 1860: 173

Material. 1 ex. (ACD), N Laos, Vientiane, Vang Vieng

prov., 19.02.2005, Li Jingki.

Distribution. South-Eastern Asia.

Subtribe Anisonychina Legalov, 2003 Genus Anisonychus Voss, 1927 Subgenus Anisonychus s. str.

Anisonychus (Anisonychus) atropterus atropterus (Voss, 1927)

Apoderus atropterus Voss, 1927: 7 Apoderus atropterus f. varipes Voss, 1927: 45 Tomapoderus nigrosculpturatus Janczyk, 1960: 52 Material. 1 ex. (ACD), Malaysia, Taiping, X.1983. Distribution. Indonesia (Java, Sumatra, Kalimantan), Malaysia (Penang, Perak, Sabah, Sarawak).

Subtribe Centrocorynina Legalov, 2003 Genus Eocentrocorynus Legalov, 2003 Subgenus Eocentrocorynus s. str.

Eocentrocorynus (Eocentrocorynus) aemulus (Faust,

1894)

Apoderus aemulus Faust, 1894a: 158 Material. 2 ex. (ACD), Laos, Sing, Louang Nantha, 11-

27.05.2006.

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Distribution. Laos, Myanmar, Thailand, Vietnam.

Subtribe Cycnotrachelina Legalov, 2003 Genus Paratrachelophorus Voss, 1924 Subgenus Paratrachelophorus s. str.

Paratrachelophorus (Paratrachelophorus) gigas Legalov, 2003

Paratrachelophorus gigas Legalov, 2003a: 581 Material. 1 ex. (ACD), N Vietnam, Sa pa, 06.2001. Distribution. China (Yunnan), Vietnam.

ACKNOWLEDGEMENTS

I wish to thank A. Allen (Boise), M.VL. Barclay (London), L. Behne (Muncheberg), R. Borovec (Nechanice), B. Brugge (Amsterdam), R. Danielsson (Lund), M. De Meyer (Tervuren), R. Dunda (Prague),

D. Efimov (Kemerovo), A.-L.-L. Friedman (Tel Aviv), J. Frisch (Berlin), A.A. Gusakov (Moscow), M. Hartmann (Erfurt), J. Hajek (Prague), O. Jaeger (Dresden), A.G. Kirejchuk (St.-Petersburg), K.-D. Klass (Dresden), A. Korshunov (Kemerovo), P. Kresl (Janovice nad Uhlavou), P. Limbourg (Bruxelles), S. Lingafelter (Washington), D. Logunov (Manchester), B.A. Korotyaev (Saint Petersburg),

E. Marais (Windhoek), O. Martin (Copenhagen), O. Merkl (Budapest), N.B. Nikitsky (Moscow), J. Pelletier (Monnaie), H. Perrin (Paris), A. Podlussany (Budapest), R. Poggi (Genova), V.Yu. Savitsky (Moscow), A. Solodovnicov (Copenhagen), M. Schmitt (Bonn), K. Ulmen (Bonn), B. Viklund (Stockholm) and J. Willers (Berlin), for their help with the work.

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Rhynchitidae gen. spp.: a - Rhinocartus tessmanni, male (dorsal view), b-Australetobius incostans, male, lectotype (dorsal view), с — A. incostans, male, lectotype, head, rostrum, antenna (lateral view), d - A. incostans, male, lectotype, mandible and apex of rostrum (dorsal vie-w),e-Auletobius aeneus, female, holotype (dorsal view), f -A. aeneus, female, holotype (frontal view), f-A. aeneus, female, holotype (lateral view), h —A. albipilosus, female, holotype (dorsal view), i—A. albipilosus, female, holotype (lateral view), j —A. ebenus, male, holotype (dorsal view).

Представители Rhynchitidae gen. spp.: a - Rhinocartus tessmanni, самец (вид сверху), b -Australetobius incostans, самец, лектотип (вид сверху), с A. incostans, самец, лектотип, голова, головотрубка, усик (вид сбоку), d - A. incostans, самец, лектотип, мандибулы и вершина головотрубки (вид сверху), е -Auletobius aeneus, самка, голотип (вид сверху), f-A. aeneus, самка, голотип (вид спереди), f-A. aeneus, самка, голотип (вид сбоку), h — A. albipilosus, самка, голотип (вид сверху), i — A. albipilosus, самка, голотип (вид сбоку), j -A. ebenus, самец, голотип (вид сверху).

Australetobius anci Auletobius spp.: a-Australetobius rubricollis, female, holotype (lateral view), b — A. rubricollis, female, holotype (dorsal view), с — A. rubricollis, female, holotype (frontal view), d - Auletobius imitator, female, holotype (lateral view), с-A. imitator, female, holotype (dorsal view), f-A. imitator, male (dorsal view), g -A. laterirostris, male, lectotype (lateral view), h —A. laterirostris, male, lectotype (dorsal view), i — A. montrouzieri, male, lectotype (dorsal view), j-A. montrouzieri, male, lectotype (frontal view), к —A. montrouzieri, female (dorsal view), I-A. melanocephalus, female, lectotype (dorsal view), m —A. montrouzieri, female (lateral view), n-A. pygmaeus, male, holotype (lateral view), о -A. melanocephalus, female, lectotype (frontal view), p -A. pygmaeus, male, holotype (dorsal view).

Представители Australetobius and Auletobius spp.: a Australetobius rubricollis, самка, голотип (вид сбоку), b - A. rubricollis, самка, голотип (вид сверху), с - A. rubricollis, самка, голотип (вид спереди), d -Auletobius imitator, самка, голотип (вид сбоку), е — A. imitator, самка, голотип (вид сверху), f-A. imitator, самец (вид сверху), g - A. laterirostris, самец, лектотип (вид сбоку), h -A. laterirostris, самец, лектотип (вид сверху), i -A. montrouzieri, самец, лектотип (вид сверху), j —A. montrouzieri, самец, лектотип (вид спереди), к -A. montrouzieri, самка (вид сверху), 1 -A. melanocephalus, самка, лектотип (вид сверху), m —A. montrouzieri, самка (вид сбоку), n -A. pygmaeus, самец, голотип (вид сбоку), о A. melanocephalus, самка, лектотип (вид спереди), р -A. pygmaeus, самец, голотип (вид сверху).

Auletini gen. spp.: a - Auletobius iviei, male, holotype (dorsal view), b - A. iviei, male, holotype (lateral view), с — A. montanus, male, holotype (lateral view), d - A. montanus, male, holotype (dorsal view), e -Pseudominurus tanganyikus, female (dorsal view), f-P. tanganyikus, female (lateral view), g-Pseudauletesluceus, female, lectotype (dorsal view), h - P. luceus, female, lectotype (lateral view), i - Auletobius maculipennis var. concolor, female, lectotype (dorsal view), j —A. cubanus, female, lectotype (dorsal view), к -A. cubanus, female, paralectotype (dorsal view), 1 — Pseudomesauletespodocarpi, male, lectotype (dorsal view).

Представители Auletini gen. spp.: a-Auletobius iviei, самец, голотип (вид сверху), b - A. iviei, самец, голотип (вид сбоку), с -A. montanus, самец, голотип (вид сбоку), d-A. montanus, самец, голотип (вид сверху), е - Pseudominurus tanganyikus, самка (вид сверху), f P. tanganyikus, самка (вид сбоку), g - Pseudauletes luceus, самка, лектотип (вид сверху), h - P. luceus, самка, лектотип (вид сбоку), i - Auletobius maculipennis var. concolor, самка, лектотип (вид сверху), j - A. cubanus, самка, лектотип (вид сверху), к-A. cubanus, самка, паралектотип (вид сверху), 1 -Pseudomesauletespodocarpi, самец, лектотип (вид сверху).

Pseudomesauletina gen. spp.: a - Tanzanauletes hustachei, male, lectotype (dorsal view), b - T. hustachei, male, lectotype (lateral view), с - Pseudomesauletes subsignatus, male, lectotype (dorsal view), d -P. subsignatus, female, paralectotype (dorsal view), e - P. collarti, male, holotype (dorsal view), f -P. subsignatus, male, lectotype (lateral view), g - Auletobius hirtellus, male, lectotype (dorsal view), h — Pseudomesauletes punctipennis, male, lectotype (lateral view), i - P. punctipennis, male, lectotype (dorsal view), j - P. punctipennis, male, lectotype, head and rostrum (dorsal view), к - P.friedmani, male, holotype (dorsal view), 1- P.friedmani, female, paratype (dorsal view), m P. ankaratraensis, female, holotype (dorsal view), n — P. ankaratraensis, female, holotype (dorsal view).

Представители Pseudomesauletina gen. spp.: a - Tanzanauletes hustachei, самец, лектотип (вид сверху), b - Т. hustachei, самец, лектотип (вид сбоку), с - Pseudomesauletes subsignatus, самец, лектотип (вид сверху), d — P. subsignatus, самка, паралектотип (вид сверху), е — P. collarti, самец, голотип (вид сверху), f-Р. subsignatus, самец, лектотип (вид сбоку), g - Auletobius hirtellus, самец, лектотип (вид сверху), h -Pseudomesauletes punctipennis, самец, лектотип (вид сбоку), i-P. punctipennis, самец, лектотип (вид сверху), j

— P. punctipennis, самец, лектотип, голова и головотрубка (вид сверху), k-P. friedmani, самец, голотип (вид сверху), 1 - P. friedmani, самка, паратип (вид сверху), m - P. ankaratraensis, самка, голотип (вид сверху), п - Р. ankaratraensis, самка, голотип (вид сверху).

Rhynchitidae gen. spp.: a - Pseudomesauletes gibbipennis, female, holotype (dorsal view), b - P. gibbipennis, female, holotype (lateral view), с - P. jizushanensis, male, holotype (dorsal view), d - Biblarodepus solitarius, male, holotype (lateral view), e - Caenorhinus rufiventris, male, lectotype (dorsal view), f -Capylavodepopsis conjinis, male, lectotype (dorsal view), g - Biblarodepus solitarius, male, holotype (dorsal view), h - B. solutus, female, holotype (lateral view), i - Rhynchites pauciseta, female, lectotype (dorsal view), j -Biblarodepus solutus, female, holotype (lateral view), к - Rhynchites pauciseta, female, lectotype (lateral view), 1 -Maculinvolvulus vestitoides, female, lectotype (dorsal view), m - Caenorhinus rufiventris, male, lectotype (lateral view), n— Maculinvolvulus vestitoides, female, lectotype (lateral view).

Представители Rhynchitidae gen. spp.: a-Pseudomesauletes gibbipennis, самка, голотип (вид сверху), b-P. gibbipennis, самка, голотип (вид сбоку), с-P. jizushanensis, самец, голотип (вид сверху), d - Biblarodepus solitarius, самец, голотип (вид сбоку), е - Caenorhinus rufiventris, самец, лектотип (вид сверху), f -Capylarodepopsis conjinis, самец, лектотип (вид сверху), g - Biblarodepus solitarius, самец, голотип (вид сверху), h - В. solutus, самка, голотип (вид сбоку), i - Rhynchites pauciseta, самка, лектотип (вид сверху), j -Biblarodepus solutus, самка, голотип (вид сбоку), k - Rhynchites pauciseta, самка, лектотип (вид сбоку), 1 -Maculinvolvulus vestitoides, самка, лектотип (вид сверху), m - Caenorhinus rufiventris, самец, лектотип (вид сбоку), n - Maculinvolvulus vestitoides, самка, лектотип (вид сбоку).

Rhynchitina gen. spp.: a - Rhynchites homalinus, female, lectotype (dorsal view), b - Rh. homalinus, female, lectotype (frontal view), с - Rh. homalinus, female, lectotype (lateral view), d - lnvolvulus aethiops ssp. juraensis, female, lectotype (dorsal view), e - Clinorhyncliites nigripes, female (lateral view), f- C. nigripes, female (dorsal view), g - C. distinguendulus, female (dorsal view), h C. scheitzae, female, lectotype (lateral view), і -Metarhynchites longulus, female, lectotype (dorsal view), j - Clinorhynchites rufofemoratus, male, lectotype (dorsal view), к - С. rufofemoratus, male, lectotype (lateral view), 1 - C. scheitzae, female, lectotype (dorsal view), m -Metarhynchites longulus, female, lectotype (lateral view), n - Afrorhynchites villosus, female, lectotype (lateral view), о - A. villosus, female, lectotype (dorsal view), p - Clinorhynchites castaneus, female, lectotype (lateral view), q-C. castaneus, female, lectotype (dorsal view).

Представители Rhynchitina gen. spp.: a - Rhynchites homalinus, самка, лектотип (вид сверху), b - Rh. homalinus, самка, лектотип (вид спереди), с - Rh. homalinus, самка, лектотип (вид сбоку), d - Involvulus aethiops ssp .juraensis, самка, лектотип (вид сверху), е - Clinorhynchites nigripes, самка (вид сбоку), f - С. nigripes, самка (вид сверху), g - С. distinguendulus, самка (вид сверху), h — С. scheitzae, самка, лектотип (вид сбоку), і - Metarhynchites longulus, самка, лектотип (вид сверху), j - Clinorhynchites rufofemoratus, самец, лектотип (вид сверху), к- С. rufofemoratus, самец, лектотип (вид сбоку), 1 - С. scheitzae, самка, лектотип (вид сверху), m - Metarhynchites longulus, самка, лектотип (вид сбоку), n - Afrorhynchites villosus, самка, лектотип (вид сбоку), о -A. villosus, самка, лектотип (вид сверху), р - Clinorhynchites castaneus, самка, лектотип (вид сбоку),q-C. castaneus,самка, лектотип (видсверху).

Rhynchitina gen. spp.: a - Proinvolvulus rugosipennis, female, lectotype (dorsal view), b - Rhynchites semiopacus, female, paralectotype (dorsal view), с - ProinvolvulusJlandriensis, female, holotype (dorsal view), d -P. rugosipennis, female, lectotype (lateral view), e - P. Jlandriensis, female, holotype (lateral view), f- Rhynchites semiopacus, male, lectotype, aedeagus (dorsal view), g - Rhynchitesparallinus var. J'allax, male, lectotype (dorsal view), h - Heterorhynchites alcyoneus female, lectotype (dorsal view), i - H. pruinosus, female, lectotype (dorsal view),j —H. subtectus, male, lectotype, aedeagus (lateral view), k-H.pruinosus, female, lectotype (lateral view), 1 -

H. subtectus, male, lectotype (dorsal view), m - H. korshunovi, male, holotype, basal sclerite of endophallus (dorsal view), n—H. subtectus, male, lectotype (lateral view).

Представители Rhynchitina gen. spp.: a-Proinvolvulus rugosipennis, самка, лектотип (вид сверху), b -Rhynchites semiopacus, самка, паралектотип (вид сверху), с - Proinvolvulus Jlandriensis, самка, голотип (вид сверху), d - P. rugosipennis, самка, лектотип (вид сбоку), е — P. Jlandriensis, самка, голотип (вид сбоку), f-Rhynchites semiopacus, самец, лектотип, эдеагус (вид сверху), g - Rhynchites parallinus var. fallax, самец, лектотип (вид сверху), h - Heterorhynchites alcyoneus самка, лектотип (вид сверху), i - Н. pruinosus, самка, лектотип (вид сверху), j - Н. subtectus, самец, лектотип, эдеагус (вид сбоку), к - Я. pruinosus, самка, лектотип (вид сбоку), I — Н. subtectus, самец, лектотип (вид сверху), m - II. korshunovi, самец, голотип, базальный склерит эндофаллуса (вид сверху), n-II. subtectus, самец, лектотип (видсбоку).

Rhynchitidae and Attelabidae gen. spp.: a - Heterorhynchites korshunovi, male, holotype (dorsal view), b -Euscelus insignis, male, lectotype (dorsal view), с - Omolabus centomyrciae, male, lectotype (dorsal view), d - O. bowringi, male, lectotype (dorsal view), e — Euscelus insignis, male, lectotype, aedeagus (dorsal view), f -Hybolabus amazonicus, female, lectotype (dorsal view), g - //. amazonicus, female, lectotype (frontal view), h - If. amazonicus, female, lectotype (lateral view), i - Euscelus insignis, male, lectotype, aedeagus (lateral view), j -Omolabus centomyrciae, male, lectotype, aedeagus (dorsal view), к - Omolabus centomyrciae, male, lectotype, aedeagus (lateral view), 1 - O. bowringi, male, lectotype, aedeagus (dorsal view), m - 0. bowringi, male, lectotype, armament of endophallus (lateral view), n - 0. bowringi, male, lectotype, armament of endophallus (dorsal view), о

- Lagenoderus brevicollis, male (dorsal view), p - L. fairmairei, female (lateral view), q - L. vadoni, male, paratype (dorsal view), r- L. vadoni, male, paratype, aedeagus (dorsal view).

Представители Rhynchitidae and Attelabidae gen. spp.: a - Heterorhynchites korshunovi, самец, голотип (вид сверху), b - Euscelus insignis, самец, лектотип (вид сверху), с - Omolabus centomyrciae, самец, лектотип (вид сверху), d -О. bowringi, самец, лектотип (вид сверху), е - Euscelus insignis, самец, лектотип, эдеагус (вид сверху), f - Hybolabus amazonicus, самка, лектотип (вид сверху), g - Н. amazonicus, самка, лектотип (вид спереди), h - Н. amazonicus, самка, лектотип (вид сбоку), i - Euscelus insignis, самец, лектотип, эдеагус (вид сбоку), j - Omolabus centomyrciae, самец, лектотип, эдеагус (вид сверху), k - Omolabus centomyrciae, самец, лектотип, эдеагус (вид сбоку), 1 - О. bowringi, самец, лектотип, эдеагус (вид сверху), m - О. bowringi, самец, лектотип, вооружение эндофаллуса (вид сбоку), п О. bowringi, самец, лектотип, вооружение эндофаллуса (вид сверху), о - Lagenoderus brevicollis, самец (вид сверху), р - L. fairmairei, самка (вид сбоку), q - L. vadoni, самец, паратип (вид сверху), т-L. vadoni, самец, паратип, эдеагус (вид сверху).

Aedeagus of Rhynchitidae gen. spp.: 1 - Rhinocartus tessmanni, 2-3 -Australetobius incostans, lectotype, 4—5 - Auletobius imitator, 6-7 -A. laterirostris, lectotype, 8-9 —A. montrouzieri, lectotype, 10-11 — A. pygmaeus, holotype, 12-A.ebenus, holotype, 13-/1. iviei, holotype, \4-A. montanus, holotype, 15-Pseudauletes luceus, 16-Tanzanauletes hustachei, lectotype, 17-18 - Pseudomesauletes subsignatus, lectotype, 19-20 — P. podocarpi, lectotype, 21-22 - Auletobius hirtellus, lectotype, 23 - Pseudomesauletes punctipennis, lectotype, 24 — P. jizushanensis, holotype, 25-P. punctipennis, lectotype,26-P. jizushanensis, holotype, 21-P. collarti, holotype, 28

— P. friedmani, holotype, 29 - Capylarodepopsis conjinis, lectotype, 30 - Biblarodepus solitarius, lectotype, 31 -Caenorhinus rufiventris, lectotype.

Эдеагусы Rhynchitidae gen. spp.: 1 Rhinocartus tessmanni, 2-3 - Australetobius incostans, лектотип, 4-5 - Auletobius imitator, 6-7-A. laterirostris, лектотип, 8-9 - A. montrouzieri, лектотип, 10-11 - A. pygmaeus, голотип, 12-/4. ebenus, голотип, 13-A.iviei, голотип, \4-A. montanus, голотип, 15-Pseudauletes luceus, 16-Tanzanauletes hustachei, лектотип, 17-18 - Pseudomesauletes subsignatus, лектотип, 19-20 - P. podocarpi, лектотип, 21-22 - Auletobius hirtellus, лектотип, 23 - Pseudomesauletes punctipennis, лектотип, 24 — P. jizushanensis, голотип, 25 -P.punctipennis, лектотип, 26-P. jizushanensis, голотип, 21 -P. collarti, голотип,28 -P.friedmani, голотип, 29 - Capylarodepopsis conjinis, лектотип, 30 - Biblarodepus solitarius, лектотип, 31 -Caenorhinus rufiventris, лектотип.

Contribution to the knowledge of the leaf-rolling weevils (Coleoptera, Rhynchitidae, Attelabidae) Текст научной статьи по специальности «Биологические науки»

Аннотация научной статьи по биологическим наукам, автор научной работы — Легалов А. А.

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Текст научной работы на тему «Contribution to the knowledge of the leaf-rolling weevils (Coleoptera, Rhynchitidae, Attelabidae)»