Catalogue of Afghanistan longhorn beetles (Coleoptera, Cerambycidae) with two descriptions of new Phytoecia (Parobereina Danilevsky, 2018) from central Asia Текст научной статьи по специальности «Биологические науки»

Humanity space International almanac VOL. 8, No 2, 2019: 104-140

http://zoobank.org/um:lsid:zoobank.org:pub:30F6FA0A-2D7A-4ED2-9EAE-AB7707FFBE61

Catalogue of Afghanistan Longhorn beetles (Coleoptera, Cerambycidae) with two descriptions of new Phytoecia (Parobereina Danilevsky, 2018) from Central Asia

M.A. Lazarev

State Budget Professional Educational Institution of the Moscow Region "Chekhov technical college"

Novaya str., 4, Novyi Byt village, Chekhov District, Moscow Region 142322 Russia e-mail: [email protected]; [email protected]

Key words: Coleoptera, Cerambycidae, taxonomy, distribution, new species, Afghanistan, Pakistan.

Abstract: The Catalogue includes all 78 Cerambycidae species of Afghanistan fauna known up to 2019 with the references to the original descriptions; 22 species were not mentioned for Afghanistan in Palaearctic Cerambycidae Catalogue by Löbl & Smetana (2010). Bibliography of each species usually includes the geographical information from corresponding publications. Many new taxonomy positions published after 2010 are used here without special remarks. Agapanthia (Epoptes) dahli ustinovi Danilevsky, 2013 stat. nov. is downgraded from the species level. Two species are described as new Phytoecia (Parobereina) pashtunica sp. n. from Afghanistan and Phytoecia (Parobereina) heinzi sp.n. from Pakistan.

The present work is an attempt to summarize all data published up to now on Cerambycidae of Afghanistan fauna.

Family CERAMBYCIDAE Latreille, 1802 subfamily Prioninae Latreille, 1802 tribe Macrotomini J. Thomson, 1861 genus Anomophysis Quentin & Villiers, 1981: 374 type species

Prionus spinosus Fabricius, 1787 inscripta C.O. Waterhouse, 1884: 380 (Macrotoma)

Heyrovsky, 1936: 211 - Wama; Tippmann, 1958: 41 - Kabul, OstAfghanistan, 1740; Sarobi, am Kabulflus, 900 m; Mangul, Bashgultal, Nuristan, Ost-Afghanistan, 1250 m; Fuchs, 1961: 259 -Sarobi 1100 m, O.-Afghanistan; Fuchs, 1967: 432 - Afghanistan, 25 km N von Barikot, 1800 m, Nuristan; Nimla, 40 km SW von Dschelalabad; Heyrovsky, 1967: 156 - Zentral-Afghanistan, Prov. Kabul: Kabul; Kabul-Zahir; Ost-Afghanistan, Prov. Nengrahar:

"Nuristan", ohne näheren Fundort; Kamu; Löbl & Smetana, 2010: 90 - Afghanistan.

plagiata C.O. Waterhouse, 1884: 381 (Macrotoma) vidua Lameere, 1903: 167 (Macrotoma)

Quentin & Villiers, 1981: 361, 376, 383 - Afghanistan: Kaboul; Sairobi; Darah-i-Nour, Nangahar; Löbl & Smetana, 2010: 90 -Afghanistan; Kariyanna et al., 2017: 267 - Afghanistan: Kaboul; Rapuzzi et al., 2019: 64 - Afghanistan.

tribe Prionini Latreille, 1802 genus Dorysthenes Vigors, 1826: 514 type species Prionus

rostratus Fabricius, 1793 subgenus Lophosternus Guérin-Méneville, 1844: 209 type species Lophosternus buquetii Guérin-Méneville, 1844 Cyrtosternus Guérin-Méneville, 1844: 210 type species Lophosternus hopei Guérin-Méneville, 1844 huegelii L. Redtenbacher, 1844: 550 (Cyrtognathus) falco J. Thomson, 1877: 262 (Cyrthognathus) palpalis Gahan, 1906: 12 (Lophosternus)

Note: D. huegelii was recorded for Afghanistan (Lobl & Smetana, 2010: 91; Rapuzzi et al., 2019: 65), and the occurance of the species in the country is rather probable, but no exact locality information was published up to now, and the corresponding specimens are not known.

genus Mesoprionus Jakovlev, 1887: 323 type species Prionus

asiaticus Faldermann, 1837 angustatus Jakovlev, 1887: 327 (Prionus) angheri Brancsik, 1899: 102 (Prionus) bucharicus Semenov, 1900: 328 (Prionus)

Löbl & Smetana, 2010: 92 - Afghanistan; Danilevsky, 2014b: 45 -North Afghanistan.

genus Miniprionus Danilevsky, 2000: 189 type species Prionus

pavlovskii Semenov, 1935 pavlovskii Semenov, 1935: 239 (Prionus)

Danilevsky, 2000: 190 - North Afghanistan.

genus Pogonarthron Semenov, 1900: 257 type species Polyarthron bedeli Semenov, 1900

subgenus Multicladum Danilevsky in Danilevsky & Komiya, 2014:

268 type species Prionus semenovianus Plavilstshikov, 1936 semenovianum Plavilstshikov, 1936: 92 (Prionus)

Löbl & Smetana, 2010: 93 - Afghanistan; Danilevsky, 2014b: 62 -North Afghanistan.

genus Prionus Geoffroy, 1762: 198 type species Cerambyx

coriarius Linnaeus, 1758 corpulentus Bates, 1878: 720

Fuchs, 1955: 272, 273 - Pashki, Nuristan; Löbl & Smetana, 2010: 93 - Afghanistan. elliotti Gahan, 1906: 16

Gahan, 1906: 16 - Baluchistan, near Quetta; Koh-i-Baba: Marrak; Fuchs, 1955: 271 - Marrak, Koh-i-Baba, Afghanistan; Shirparak, Koh-i-Baba, Afghanistan; Tippmann, 1958: 41 - Senna, Kokschatal, Badakschan, NO-Afghanistan; Löbl & Smetana, 2010: 94 -Afghanistan; Kariyanna et al., 2017: 273 - Afghanistan, Koh-i-Baba, Marrak.

vartianorum Fuchs, 1967: 431

Fuchs,1967: 431 - Afghanistan: Paghman, 30 km NW von Kabul, 2500 m; Khurd-Kabul [34°23'N, 69°23'E], SO von Kabul, 1900 m; Löbl & Smetana, 2010: 94 - Afghanistan; Rapuzzi et al., 2019: 66 -Afghanistan.

genus Pseudoprionus Pic, 1898: 33 type species Polyarthron

bienerti Heyden, 1885 altimontanus Kabakow & Dolin, 1996: 45 (Prionus)

Kabakow & Dolin, 1996: 45 - Afghanistan: Prov. Uruzdan, N von Schachrestan; SW Panjav / Qonay Pass; Zabolj Prov., W von Argandab; Danilevsky, 2009: 293 - Central Afghanistan; Löbl & Smetana, 2010: 94 - Afghanistan.

subfamily Lepturinae Latreille, 1802 tribe Lepturini Latreille, 1802 genus Anastrangalia Casey, 1924: 280 type species Leptura

sanguinea LeConte, 1859 rubriola Bates, 1878: 720 (Leptura) kashmirica Plavilstshikov, 1927: 105

Heyrovsky, 1936: 211 - Parun-Tal; Schigi; Löbl & Smetana, 2010: 97 - Afghanistan; Rapuzzi et al., 2019: 66 - Afghanistan.

genus Pedostrangalia Sokolov, 1897: 461 type species Pedostrangalia kassjanowi Sokolov, 1897 (= Leptura imberbis Menetries, 1832)

subgenus Pedostrangalia Sokolov, 1897: 461 type species Pedostrangalia kassjanowi Sokolov, 1897 (= Leptura imberbis Menetries, 1832) afghanistana Sato & N. Ohbayashi, 1976: 485

Sato & N. Ohbayashi, 1976: 485 - Kabul, Aliabad; Lobl & Smetana, 2010: 110 - Afghanistan. imberbis Menetries, 1832: 231 (Leptura)

angulicollis Heyden, 1879: 323 [=1879: 67] (Strangalia) kassjanowi Sokolov, 1897: 461

Ozdikmen, 2004: 24, 28 - Afghanistan. subgenus Neosphenalia Lobl, 2010: 60 type species Leptura verticalis Germar, 1822

Sphenalia K. Daniel, 1904: 355 [HN] type species Leptura verticalis Germar, 1822

? kurda Sama, 1996: 104

Nota. The record of P.emmipoda (Mulsant, 1863) for Afghanistan by Ozdikmen (2004: 25, 28) had to be connected with P. kurda Sama, 1996, but later Ozdikmen (2014: 403) did not included Afghanistan in the general area of P .emmipoda.

genus Rutpela Nakane & K. Ohbayashi, 1957: 242 type species

Leptura maculata Poda von Neuhaus, 1761 inermis K. Daniel & J. Daniel, 1898: 74 (Strangalia) elboursensis Pic, 1905: 390 (Strangalia)

Heyrovsky, 1971: 82 - Herat, province d'Herat, Afghanistan septentrional.

tribe Rhagiini Kirby, 1837 genus Cortodera Mulsant, 1863: 572 type species Grammoptera spinosula Mulsant, 1839: 290 (= Leptura humeralis Schaller, 1783) bamiyana Danilevsky, 2014a: 256

Danilevsky, 2014a: 256 - Afghanistan, Bamiyan, Panjab Distr., Varas; Ozdikmen, 2016: 14 - Afghanistan.

Note. The records of Cortodera transcaspica persica Plavilstshikov, 1936 and C. pseudomophlus Reitter, 1889 for

Afghanistan by Özdikmen (2003) were most probably based on wrong determination of local species.

subfamily Apatophyseinae Lacordaire, 1869 tribe Apatophyseini Lacordaire, 1869 genus Apatophysis Chevrolat, 1860a: 95 type species Apatophysis toxotoides Chevrolat, 1860 (= Polyarthron barbarum P. H. Lucas, 1858)

subgenus Angustephysis Pic, 1956: 2 type species Apatophysis

richteri Pic, 1956 margiana Semenov & Shchegoleva-Barovskaya, 1936: 77 plavilstshikovi Miroshnikov, 1992: 392

Danilevsky, 2008: 37, 41 - North Afghanistan modica Gahan, 1906: 71

Danilevsky, 2006: 9 - south Afghanistan; Löbl & Smetana, 2010: 142 - Afghanistan.

subgenus Apatophysis Chevrolat, 1860a: 95 type species Apatophysis toxotoides Chevrolat, 1860 (= Polyarthron barbarum P. H.Lucas, 1858) afghanica Miroshnikov, 2014: 14

Miroshnikov, 2014: 14 - Afghanistan, Paktya, Spera. caspica Semenov, 1901: 31

Danilevsky, 2006: 2 - Afghanistan; 2008: 22, 41 - Afghanistan; Löbl & Smetana, 2010: 142 - Afghanistan.

genus Protapatophysis Semenov & Stschegoleva-Barovskaia,

1936: 26 type species Apatophysis kashmiriana Semenov, 1901 kabakovi Danilevsky, 2011: 102

Heyrovsky, 1936: 211 (as A .kashmirina Semenov, 1901) -Wama;Tippmann, 1958: 47 (as A . kashmirina Semenov, 1901) -Kamdesch, 2000 m, Bashgultal, Nuristan, O-Afghanistan; Fuchs, 1967: 433 - Afghanistan, Khurd-Kabul, SO von Kabul, 1900 m (as A. kashmirina Semenov, 1901) and Afghanistan, Paghman, 30 km NW von Kabul, 2500 m (as A. montana Gahan, 1906); Löbl & Smetana, 2010: 142 - Afghanistan (as A. vartianae Heyrovsky, 1971); Danilevsky, 2011: 101, 104 - Afghanistan, prov. Kunar, SW Pech-Dara; North-east Afghanistan, West of Konar province (Chapa Dara district).

subfamily Cerambycinae Latreille, 1802 tribe Achrysonini Lacordaire, 1868 genus Nortia J. Thomson, 1864: 252 type species Nortia cavicollis

J. Thomson, 1864 afghanica Heyrovsky, 1967: 158

Heyrovsky, 1967: 157, 158 - Ost-Afghanistan, Prov. Nengrahar: Kamu, 1450 m; Löbl & Smetana, 2010: 143 - Afghanistan.

tribe Callichromatini Swainson & Shuckard, 1840 genus Osphranteria L. Redtenbacher, 1850: 50 type species Osphranteria suaveolens L. Redtenbacher, 1850 Quettania Schwarzer, 1931: 62 type species Quettania coeruleipennis Schwarzer, 1931. coerulescens L. Redtenbacher, 1850: 50 inaurata Holzschuh, 1981: 98 coeruleipennis Schwarzer, 1931a: 63 (Quettania) mirabilis Podany, 1980: 231 (Polyzonus)

Note. The species must occur in Afghanistan as it is distributed in Pakistan and in Iran. lata Pic, 1956: 3

richteri Heyrovsky, 1959: 4

Löbl & Smetana, 2010: 149 - Afghanistan; Bentanachs, 2012a: 88,

89, 96 - Afghanistán; 2012b: 71 - Afghanistan; Rapuzzi et al., 2019: 72 - Afghanistan.

suaveolens L. Redtenbacher, 1850: 50

Heyrovsky, 1967: 157 - Ost-Afghanistan, Prov. Nengrahar: Kamu; Löbl & Smetana, 2010: 149 - Afghanistan; Bentanachs, 2012a: 88,

90, 96 - Afganistán; 2012b: 71 - Afghanistan.

tribe Callidiini Kirby, 1837 genus Turanium Baeckmann, 1922: 32 type species

Callidiumscabrum Kraatz, 1882 subgenus Turanium Baeckmann, 1922: 32 type species

Callidiumscabrum Kraatz, 1882 scabrum Kraatz, 1882a: 115 (Callidium) simplarium Heyden, 1885: 296 (Callidium)

Danilevsky, 2001: 580 - North Afghanistan (no specimens from the country known).

tribe Cerambycini Latreille, 1802 genus Trirachys Hope, 1843: 63 type species Trirachys orientalis Hope, 1843

Trirhachis Agassiz, 1846: 378 [unjustified emendation] Trirrhachys Gemminger, 1872: 2801 [unjustified emendation] indicola Bates, 1891: 21 (Neocerambyx)

Heyrovsky, 1936: 211 - Wama [35°11'N 70°48'E]; Löbl & Smetana, 2010: 158 - Afghanistan. sarta Solsky, 1871: 150 (Pachydissus)

Fuchs, 1955: 273 - Kabul; 1967: 433 - Afghanistan, Paghman, 30 km NW von Kabul, 2500 m; Tippmann, 1958: 45- Kabul, 1740 m, O-Afghanistan; Bashgultal, 1300 m, Nuristan, O-Afghanistan; Pagmangebirge, 2300 m, Ost-Afghanistan; Kandahar, 950 m, SüdAfghanistan; Darufulun, bei Kabul, O-Afghanistan, 1800 m; Fuchs, 1961: 259 - Kabul; Polichomri N.-Afghanistan, 700 m; Heyrovsky, 1967: 156 - Zentral-Afghanistan, Prov. Kabul: Kabul; Hua, 2002: 191 - Afghanistan; Löbl & Smetana, 2010: 158- Afghanistan; Kumawat et al., 2015: 7895 - Afghanistan;Vitali, Gouverneur & Chemin, 2017: 46, 48 - Afghanistan, Kaboul, Darulaman; Kariyanna et al., 2017: 36 - Afganistan: Kabul; Rapuzzi et al., 2019: 68 -Afghanistan.

genus Derolus Gahan, 1891: 26 type species Hammaticherus mauritanicus Buquet, 1840

Capnocerambyx Reitter, 1894: 356 type species Hammaticherus mauritanicus Buquet, 1840 iranensis Pic, 1956: 3 iranensis iranensis Pic, 1956: 3

iranensis Lepesme & Breuning, 1958: 179 [HN]

Löbl & Smetana, 2010: 159 - Afghanistan; Rapuzzi et al., 2019: 70 -Afghanistan.

genus Diorthus Gahan, 1891: 27 type species Hammaticherus

simplex A. White, 1853 (= Cerambyx cinereus Fabricius, 1793) subgenus Diorthus Gahan, 1891: 27 type species Hammaticherus simplex A. White, 1853 (= Cerambyx cinereus Fabricius, 1793) cinereus Fabricius, 1793: 265 (Cerambyx) [NP] holosericeus Olivier, 1795: 14 (Cerambyx) inclemensJ. Thomson, 1865: 576 (Pachydissus) simplexA. White, 1853: 130 (Hammaticherus) sordidusPascoe, 1888: 491 (Neocerambyx)

vernicosusPascoe, 1859: 19 (Cerambyx)

Fuchs, 1961: 259 - Kabul; Löbl & Smetana, 2010: 160 - Afghanistan; Rapuzzi et al., 2019: 69 - Afghanistan. genus Hoplocerambyx J. Thomson, 1864: 229 type species

Hammaticherus spinicornis Newman, 1842 spinicornis Newman, 1842a: 245 (Hammaticherus) minor Pic, 1923: 8 (Haplocerambyx) morosus Pascoe, 1857: 92 (Cerambyx) relictus Pascoe, 1866: 528

Gahan, 1906: 131 - South Afghanistan; Hayashi & Makihara, 1981: 188 - Afghanistan; Hüdepohl, 1990: 65 - Afghanistan; Niisato, 1990: 112- Afghanistan; Mukhopadhyay & Biswas, 2000: 51 - South Afghanistan; Hua, 2002: 211 - Afghanistan; Löbl & Smetana, 2010: 160 - Afghanistan;Nga, Long & Thinh, 2014: 434 - Afghanistan; Kumawat et al., 2015: 7895 - Afghanistan; Majumder et al., 2015: 8244 - Afghanistan; Kariyanna et al., 2017: 31 - South Afghanistan)

genus Neoplocaederus Sama, 1991: 123 [RN] type species

Plocaederus cyanipennis J. Thomson, 1861 scapularis Fischer von Waldheim, 1821: 15 (Cerambyx) tataricus Gebler, 1841: 375 (Hammaticherus)

Waterhouse, 1889: 130 - Afghanistan, Hari-rud valley; Fuchs, 1961: 259 - Polichomri N.-Afghanistan, 700 m; Fuchs, 1967: 433 -Afghanistan, Khurd-Kabul, SO von Kabul, 1900 m; Villiers, 1967: 354 - Nord de l'Afghanistan; Löbl & Smetana, 2010: 162 -Afghanistan; Kotan & Sama, 2011: 59 - Afghanistan.

genus Xenopachys Sama, 1999: 46 type species Dissopachys

matthiesseni Reitter, 1907 matthiesseni Reitter, 1907: 218 (Dissopachys)

Fuchs, 1967: 433 - Afghanistan, 25 km N von Barikot, 1300 m, Nuristan; Heyrovsky, 1967: 157 - Ost-Afghanistan, Prov. Nengrahar: Kamu.

schurmanni Sama, 1999: 46

Sama, 1999: 46 - Afghanistan: Ghov., Jam; Löbl & Smetana, 2010: 162- Afghanistan.

tribe Clytini Mulsant, 1839 genus Chlorophorus Chevrolat, 1863: 290 type species Callidium

annulare Fabricius, 1787 subgenus Humeromaculatus Özdikmen, 2011: 536 type species

Cerambyx figuratus Scopoli, 1763 quatuordecimmaculatus Chevrolat, 1863: 295 (Anthoboscus) guerryi Pic, 1902: 30 (Clytanthus)

Tippmann, 1958: 54 - Firgamu, Kokschatal, Badakschan, 2300 m, NO-Afghanistan; Löbl & Smetana, 2010: 168 -Afghanistan;Kariyanna et al., 2017: 46 - Afganistan. subgenus Immaculatus Özdikmen, 2011: 536 type species

Chlorophorus kanoi Hayashi, 1963 faldermanni Faldermann, 1837: 269 (Clytus) caucasicus Pic, 1897: 262 (Clytanthus) mesminiPic, 1908: 3 (Clytanthus) punctomaculatusPic, 1893: 26 (Clytus) quinquemaculatusGebler, 1845: 104 (Clytus)

Tippmann, 1958: 51 - Kamdesch, Bashgultal, 2000 m, Nuristan, O-Afghanistan; Kutiau, 1500 m, Ost-Nuristan, Afghanistan; Villiers, 1967: 361 - Nord Afghanistan; Löbl & Smetana, 2010: 166-Afghanistan) varius O. F. Müller, 1766: 188 varius varius O. F. Müller, 1766: 188

Tippmann, 1958: 51 - Schiva, Hochsteppe 2800 m, Badakschan, NO-Afghanistan.

genus Perissus Chevrolat, 1863: 262 type species Perissus x-littera

Chevrolat, 1863 delerei Tippmann, 1958: 49

Tippmann, 1958: 49 - Bashgultal, 1100 m, Nuristan, O. Afghanistan; Löbl & Smetana, 2010: 175 - Afghanistan.

genus Xylotrechus Chevrolat, 1860b: 456 type species Clytus

sartorii Chevrolat, 1860 subgenus Turanoclytus Sama, 1994: 325 type species Clytus

namanganensis Heyden, 1885 namanganensis Heyden, 1885: 297 (Clytus) bucharensis Semenov, 1893: 501 (Clytus) subcrucifer Pic, 1903: 4

Gressitt, 1951: 249 - Afghanistan; Hua, 2002: 236 - Afghanistan.

subgenus Xylotrechus Chevrolat, 1860b: 456 type species Clytus

sartorii Chevrolat, 1860 badachschanicus Tippmann, 1958: 48

Tippmann, 1958: 48 - Firgamu, Kokschatal, 2300 m, Badakschan, NO-Afghanistan; Löbl & Smetana, 2010: 181- Afghanistan. smei Laporte & Gory, 1841: 37 (Clytus)

Ahmad, 1946: 35, 44 - Kabul; Heyrovsky, 1971: 82 - Afghanistan oriental, 30 km au Nord de Kaboul; Majumder et al., 2015: 8245 -Afghanistan; Kariyanna et al., 2017: 79 - Afghanistan (Kabul); Rapuzzi et al., 2019: 72 - Afghanistan. stebbingi Gahan, 1906: 244

Tippmann, 1958: 48 - Kabul, 1740 m, Ost-Afghanistan; Fuchs, 1961: 259 - Kabul; Löbl & Smetana, 2010: 183 - Afghanistan; Kariyanna et al., 2017: 63 - Afganistan.

tribe Hesperophanini Mulsant, 1839 subtribe Daramina Sama, 2008 genus Zoodes Pascoe, 1867: 319 type species Stromatium maculatum A. White, 1855

Cornutaemospila Pic, 1927: 30 type species Cornutaemospila piceosignata Pic, 1927

compressus Fabricius, 1787: 153 (Callidium)

Fuchs, 1967: 433 - Afghanistan, 25 km N von Barikot, 1300 m, Nuristan; Löbl & Smetana, 2010: 187 - Afghanistan.

tribe Oemini Lacordaire, 1868 genus Noserius Pascoe, 1857: 95 type species Noserius tibialis

Pascoe, 1857 indicus Gahan, 1906: 104 (Hypoeschrus)

Löbl & Smetana, 2010: 194- Afghanistan; Kariyanna et al., 2017: 79 - Afghanistan; Rapuzzi et al., 2019: 67 - Afghanistan.

tribe Purpuricenini J. Thomson, 1861 genus Afghanicenus Heyrovsky, 1941: 36 type species

Purpuricenus nuristanicus Heyrovsky, 1936 dreesi Tippmann, 1958: 57

Tippmann, 1958: 57- Kamu, Bashgultal, 1500 m, Nuristan, O-Afghanistan;Bashgultal, 1200 m, Nuristan, O-Afghanistan; Kutiau, 1450 m, Nuristan, O-Afghanistan; Löbl & Smetana, 2010: 196 -Afghanistan.

nuristanicus Heyrovsky, 1936: 212 (Purpuricenus)

Heyrovsky, 1936: 211, 212 - Gulan-Tal; Brubrutz im Kti-Tal; Kti-Tal; Heyrovsky, 1941: 37, 39 - Afghanistan or., Nuristan; Tippmann, 1958: 56 - Pagman-Gebirge, 2300 m, O-Afghanistan; Bashgultal, 1200 m, Nuristan, O-Afghanistan; Löbl & Smetana, 2010: 196 -Afghanistan.

genus Purpuricenus Dejean, 1821: 105 type species Cerambyx kaehleri Linnaeus, 1758

Acanthoptera Latreille, 1829: 114 type species Cerambyx budensis Götz, 1783 Cyclodera A. White, 1846: 510 type species Cyclodera quadrinotata A. White, 1846

Hamadrias Gistel, 1848: 130 [unnecessary substitute name]

Philagathes J. Thomson, 1864: 196 type species Philagathes laetus J. Thomson,

1864

Porphyrocenus Reitter, 1913: 34 type species Purpuricenus spectabilis Motschulsky, 1858

Sternoplistes Guerin-Meneville, 1844: 224 type species Purpuricenus temminckii Guerin-Meneville, 1844 indus Semenov, 1908: 261 [RN] haussknechti Gahan, 1906: 186

Ahmad, 1946: 35, 44 - Afghanistan, Darra-i-Khail; Tippmann, 1958: 55 - Bashgultal, Nuristan, 1200 m, O-Afghanistan; Heyrovsky, 1967: 157-Ost-Afghanistan, Prov. Nengrahar: Kamu, 1450 m; Miroshnikov & Lobanov, 1990: 17 - Afghanistan; Löbl & Smetana, 2010: 198 -Afghanistan; Ambrus & Tichy, 2017: 95 - Afghanistan; Kariyanna et al., 2017: 78 - Afganistan; Rapuzzi et al., 2019: 72 - Afghanistan. kabakovi Miroshnikov & Lobanov, 1990: 15

Miroshnikov & Lobanov, 1990: 15 - Afghanistan: Nuristan, Pech River, SW Chapadar, 2000-2400 m; Löbl & Smetana, 2010: 198 -Afghanistan; Danilevsky, 2012b: 149 - Afghanistan; Kariyanna et al., 2017: 78 - Afghanistan: Nuristan, SW Chanadar; Rapuzzi et al., 2019: 71 - Afghanistan. montanus A. White, 1853: 138

Ambrus & Tichy, 2017: 96 - Afghanistan; Löbl & Smetana, 2010: 199 - Afghanistan; Rapuzzi et al., 2019: 71 - Afghanistan.

tribe Tillomorphini Lacordaire, 1868 genus Cleroclytus Kraatz, 1884: 226 type species Cleroclytus semirufus Kraatz, 1884

subgenus Cleroclytus Kraatz, 1884: 226 type species Cleroclytus

semirufus Kraatz, 1884 semirufus Kraatz, 1884: 225 semirufus collaris Jakovlev, 1885: 290 manifestus Jakovlev, 1900: 664 strigicollis Jakovlev, 1900: 663

Löbl & Smetana, 2010: 206 - Afghanistan. semirufus semirufus Kraatz, 1884: 225

Heyden, 1894: 83 - Afghanistan, Kischlak-Tusch.

subfamily Lamiinae Latreille, 1825

tribe Agapanthiini Mulsant, 1839 genus Agapanthia Audinet-Serville, 1835: 35 type species

Cerambyx cardui Linnaeus, 1767 subgenus Epoptes Gistel, 1857a:93 [1857b: 605] type species Lamia asphodeli Latreille, 1804

Agapanthiella Pesarini & Sabbadini, 2004: 126 type species Cerambyx villoso viridescens DeGeer, 1775 dahli C. F. W. Richter, 1821: pl. 12 dahli ustinovi Danilevsky, 2013: 35, stat. nov.

Note. Afghanistan, Badakhshän province. The taxon was described from Tadzhikistan, but just from the borderline with Afghanistan. detrita Kraatz, 1882b: 336

Heyden, 1894: 83 - Afghanistan, Kischlak-Tusch; Tippmann, 1958: 60.

nigriventris C. O. Waterhouse, 1889: 130

Waterhouse, 1889: 130 -Afghanistan, Hari-rud valley; Winkler, 1929: 1212 - Afghanistan; Löbl & Smetana, 2010: 216 - Afghanistan. paki Rapuzzi, 2012: 958

Rapuzzi, 2012: 958 - Afghanistan: Ghor prov., 16 km E Chagcharan, Bandi-Ali vill. env. subgenus Stichodera Pesarini & Sabbadini, 2004: 126 type species

Saperda irrorata Fabricius, 1787 soror Kraatz, 1882b: 336

Tippmann, 1958: 61 - Schiva, Hochsteppe, 2800 m, Badakschan, NO-Afghanistan.

tribe Apodasyini Lacordaire, 1872 genus Cylindilla Bates, 1884: 250 type species Cylindilla grisescens Bates, 1884

Anaesthetomorphus Pic, 1929a: 6 [RN] type species Pseudanesthetis apicalis Pic, 1929 (= Cylindilla grisescens Bates, 1884)

Ascoldatimura Breuning, 1960: 27 [RN] type species Atimura ascoldensis Heyden, 1884 (= Cylindilla grisescens Bates, 1884)

Microestola Gressitt, 1940: 180 type species Microestola bidentata Gressitt, 1940

Mimatimura Breuning, 1958b: 492 [HN] type species Atimura ascoldensis Heyden, 1884 (= Cylindilla grisescens Bates, 1884)

Pseudanesthetis Pic, 1929b: 119 [HN] type species Pseudanesthetis apicalis Pic,

1929 (= Cylindilla grisescens Bates, 1884) parallela Gressitt, 1951: 515 (Microestola)

Tippmann, 1958: 59 - Asmar, 900 m, Kunartal, Nuristan, O.Afghanistan; Bashgultal, 1100 m, Nuristan, O-Afghanistan; Lobl & Smetana, 2010: 222 - Afghanistan.

Note. The occurance of the species in Afghanistan is very doubtful as it was decsribed from China (Hubei).

genus Sophronica Blanchard, 1845: 160 type species Sophronica calceata Chevrolat, 1855

Dasyo Pascoe, 1858: 253 type species Dasyo lineata Pascoe, 1858 Dasystola Kolbe, 1894: 63 type species Dasystola hirta Kolbe, 1894 (= Sophronica carbonaria Pascoe, 1864)

Dimbrokoa Pic, 1944: 16 type species Dimbrokoa apicalis Pic, 1944 Elithiotes Pascoe, 1864: 279 type species Elithiotes hirsuta Pascoe, 1864 Eupogonioides Fisher, 1930: 275 type species Eupogonioides gardneri Fisher,

1930 (= Phunginus apicalis Pic, 1922)

Lasiapheles Bates, 1873: 382 type species Lasiapheles obrioides Bates, 1873 Mimanaesthetis Pic, 1926: 16 type speciesMimanaesthetis atripennis Pic, 1926 Phunginus Pic, 1922: 15 type species Phunginus apicalis Pic, 1922 egena Holzschuh, 2006: 488

Holzschuh, 2006: 488 - Est-Afghanistan, Nuristan, Baschgultal; Lobl & Smetana, 2010: 227 - Afghanistan.

tribe Apomecynini J. Thomson, 1860 genus Apomecyna Dejean, 1821: 108 type species type species Saperda

alboguttata Megerle, 1802 (= Lamia histrio Fabricius, 1793) subgenus Apomecyna Dejean, 1821: 108 type species Saperda alboguttata Megerle, 1802 (= Lamia histrio Fabricius, 1793) Anapomecyna Pic, 1925: 29 type species Anapomecyna luteomaculata Pic, 1925

Crassapomecyna Breuning, 1958b: 492 type species Apomecyna crassiuscula Fairmaire, 1896

Mecynapus J. Thomson, 1858: 187 type species Apomecyna parumpunctata Chevrolat, 1856

Pseudoalbana Pic, 1895: 77 type species Pseudoalbana lameerei Pic, 1895 Vocula Lacordaire, 1872: 587 type species Vocula irrorata Lacordaire, 1872 (= Apomecyna parumpunctata Chevrolat, 1856) leucosticte Hope, 1831: 28 (Callidium) lineate Pic, 1937: 14

Löbl & Smetana, 2010: 228 - Afghanistan.

tribe Batocerini J. Thomson, 1864 genus Apriona Chevrolat, 1852: 414 type species Lamia germari Hope, 1831

subgenus Apriona Chevrolat, 1852: 414 type species Lamia germari Hope, 1831

Anapriona Breuning, 1949: 8 type species: Apriona submaculosa Pic, 1917 Cylindrapriona Breuning, 1949: 8 type species Monochamus cylindricus J. Thomson, 1857

Humeroapriona Breuning, 1949: 8 type species Lamia swainsoni Hope, 1840 Mesapriona Breuning, 1949: 8 type species Apriona punctatissima Kaup, 1866 Parapriona Breuning, 1948: 17 type species: Parapriona brunneomarginata Breuning, 1948 cinerea Chevrolat, 1852: 416 newcombei Gilmour, 1958: 112

Breuning, 1967a: 435 -Afghanistan, 25 km N von Barikot, 1300 m, Nuristan; Heyrovsky, 1967: 157-Ost-Afghanistan, Prov. Nengrahar: Kamu, 1450 m; Löbl & Smetana, 2010: 237 - Afghanistan; Rapuzzi et al., 2019: 76 - Afghanistan.

tribe Exocentrini Pascoe, 1864 genus Exocentrus Dejean, 1835: 339 type species Cerambyx balteatus Fabricius sensu Dejean, 1835 (= Cerambyx lusitanus Linnaeus, 1767)

Bicolorihirtus Kusama & Tahira, 1978: 9 type species Exocentrus venatoides Kusama & Tahira, 1978

Formosexocentrus Breuning, 1958a: 322 type species Exocentrus variepennis Schwarzer, 1925 parrotiae Fisher, 1932: 304-306 klapperichi Breuning, 1957: 12

Breuning, 1957: 12 - Afghanistan: Nuristan, Kutiau, 1500 m; Tippmann, 1958: 59 - Kutiau, 1500 m, Nuristan, O-Afghanistan;

Löbl & Smetana, 2010: 310 - Afghanistan; Lingafelter et al., 2014: 84 - Afghanistan: Nuristan, Kutiau; Holzschuh, 2015: 48 -Afghanistan.

tribe Phytoeciini Mulsant, 1839 genus Paramallosia Fuchs, 1955: 273 type species Paramallosia

afghanica Fuchs, 1955 afghanica Fuchs, 1955: 274

Fuchs, 1955: 274 - Tarapas, Koh-i-Baba, Afghanistan; Panjao, Kohi-Baba, Afghanistan; Ghilzai, Koh-i-Baba, Afghanistan; Schirparak, Koh-i-Baba, Afghanistan; Löbl & Smetana, 2010: 302 - Afghanistan. genus Phytoecia Dejean, 1835: 351 type species Cerambyx

cylindricus Linnaeus, 1758 subgenus Parobereina Danilevsky, 2018: 131 type species

Phytoecia vittipennis Reiche, 1877 pashtunica Lazarev, sp.n.

Afganistan, Bamian province, Waras district, 5 km SW Waras near Dahane Denawak village, 34°15'42"N, 66°52'43"E, 2680 m.

povolnyi Heyrovsky, 1971: 82

Heyrovsky, 1971: 82 - Darunta, province de Nengrahar, Afghanistan oriental; Skrylnik, 2010: 260, 261 - Afghanistan: Kaboul prov., Nangarhar prov.; Löbl & Smetana, 2010: 302 - Afghanistan. ochraceipennis Kraatz, 1882b: 337

Tippmann, 1958: 61 - Kutiau, Nuristan; 1500 m, O-Afghanistan; Kabul, 1740 m, O-Afghanistan; Bashgultal, 1200-1300 m. Nuristan, O-Afghanistan; Löbl & Smetana, 2010: 302 - Afghanistan. tatyanae Skrylnik, 2010: 260

Skrylnik, 2010: 260 - Afghanistan: Bamian prov., Band-e Amir env. [34°50'59.64"N, 67°12'8.10"E]; Kabul prov., Paghman. tekensis Semenov, 1896: 258

Fuchs, 1961: 259 - Gulbahar 1700 m, O.-Afghanistan; Löbl & Smetana, 2010: 302 - Afghanistan. subgenus Fulgophytoecia Pic, 1900: 16 type species Phytoecia circumdata Kraatz, 1882

Pseudomallosia Breuning, 1967b: 1 type species Pseudomallosia parterufipennis Breuning, 1967 circumdata Kraatz, 1882b: 337

parterufipenis Breuning, 1967b: 2 (Pseudomallosia) sellata Ganglbauer, 1887: 296

Heyrovsky, 1936: 212 - Gulan-Tal; Tippmann, 1958: 61 - Bazarak, Panchirtal, 2200 m, O-Afghanistan; Breuning, 1967b: 2 -Afghanistan: Hazaradjat, Koh-i-Baba, Pandjao Umgebung; Löbl & Smetana, 2010: 303 - Afghanistan. valentinae Skrylnik, 2010: 258

Skrylnik, 2010: 258 - C. Afghanistan, Bamian prov., Band-e Amir env.

subgenus Opsilia Mulsant, 1863: 387 type species Phytoecia

flavicans Mulsant, 1851 (= Leptura coerulescens Scopoli, 1763) brevicornis Kasatkin, 2013: 265

Kasatkin, 2013: 265 - Afghanistan, Bamiyan Prov., 8 km South of Bamiyan, Kohi-Baba Mts., Khushkak vill. env. bucharica Breuning, 1943: 101 breuningi Dahlgren, 1988: 114

Löbl & Smetana, 2010: 301 - Afhganistan.

Note. A series of specimens with the label: "Nurestan, N Waigal riv., 2000-3000m, IV-VII, 1971-73, O.Kabakov leg." is preserved in the collection of M.L. Danilevsky (Moscow). subgenus Pseudocoptosia Pic, 1900: 16 type species Phytoecia

eylandti Semenov, 1891 cinerascens Kraatz, 1882b: 337 (Phytoecia) sokolowi Semenov, 1895: 206, 209 (Phytoecia)

Heyden, 1894: 83 - Afghanistan; Heyrovsky, 1967:158 - NordAfghanistan, Prov. Herat: Bala Murghab, 470 m; Löbl & Smetana, 2010: 292 - Afghanistan. eylandti Semenov, 1891: 380 (Phytoecia) glasunowi Semenov, 1895b: 207, 210 (Phytoecia)

Note. The species is undoubtedly distributed in Afghanistan, as it is very numerous in Turkmenia just on Afghanistan boder (Kushka environs).

tribe Pteropliini J. Thomson, 1860 genus Niphona Mulsant, 1839: 169 type species Niphona

picticornis Mulsant, 1839 subgenus Niphona Mulsant, 1839: 169 type species Niphona

picticornia Mulsant, 1839 indica Breuning, 1938: 239 grisea Breuning, 1938: 241

Heyrovsky, 1967: 157 - Ost-Afghanistan, Prov. Nengrahar: Samarchel, 900 m; Löbl & Smetana, 2010: 316 - Afghanistan; Kariyanna et al., 2017: 195 - Afghanistan; Rapuzzi et al., 2019: 73 -Afghanistan.

genus Pterolophia Newman, 1842c: 370 [NP] type species Mesosa

bigibbera Newman, 1842 subgenus Pterolophia Newman, 1842c: 370 [NP] type species Mesosa bigibbera Newman, 1842

Acroptycha Quedenfeldt, 1888: 209 type species Acroptycha spinifera Quedenfeldt, 1888

Alyattes J. Thomson, 1864: 48 type species Alyattes guineensis J. Thomson, 1864

Cormia Pascoe, 1864: 281 type species Cormia ingrata Pascoe, 1864 Eurycotyle Blessig, 1873: 210 type species Eurycotyle maacki Blessig, 1873 Incamelomorpha Pic, 1926: 4 type species Pterolophia depensis Pic, 1926 Paralophia Aurivillius, 1925: 30 [HN] type species Paralophia quadrinodosa Aurivillius, 1925

Praonetha Dejean, 1835: 344 [NO] type species Lamia crassipes Wiedemann, 1823

Praonetha Pascoe, 1862a: 348 [HN, RN] type species Prioneta albosignata Blanchard, 1853

Prioneta Blanchard, 1853: 292 [RN] type species Prioneta albosignata Blanchard, 1853

Prionetopsis J. Thomson, 1864: 49 type species Prionetopsis balteata J. Thomson, 1864 (= Lamia inaequalis Fabricius, 1801)

Theticus J. Thomson, 1858: 190 type species Theticus biarcuatus J. Thomson, 1858

apicefusca Breuning, 1938: 319

Tippmann, 1958: 58 - Bashgultal, 1200 m, Nuristan, O-Afghanistan; Löbl & Smetana, 2010: 319 - Afghanistan; Kariyanna et al., 2017: 199 - Afghanistan.

tribe Saperdini Mulsant, 1839 genus Glenea Newman, 1842b: 301 type species Saperda

novemguttata Guérin-Méneville, 1831 subgenus Glenea Newman, 1842b: 301 type species Saperda novemguttata Guérin-Méneville, 1831

Accola Jordan, 1894: 503 [HN] type species Accola citrina Jordan, 1894 (= Glenea astathiformis Breuning, 1958)

Accolona E. Strand, 1942: 392 [RN] type species Accola citrina Jordan, 1894 (= Glenea astathiformis Breuning, 1958)

Cryllis Pascoe, 1867: 363 type species Cryllis clytoides Pascoe, 1867 Hapochoron Gistel, 1848: x [RN]

Sphenura Dejean, 1835: 350 [HN] type species Saperda novemguttata Guérin-Méneville, 1831 afghana Breuning, 1971: 44

Breuning, 1971: 44 - Afghanistan: Shogran; Löbl & Smetana, 2010: 324 - Afghanistan; Rapuzzi et al., 2019: 78 - Afghanistan.

genus Saperda Fabricius, 1775: 184 type species Cerambyx

carcharias Linnaeus, 1758 subgenus Compsidia Mulsant, 1839: 182 type species Cerambyx

populneus Linnaeus, 1758 populnea Linnaeus, 1758: 394 populnea populnea Linnaeus, 1758: 394 (Cerambyx) betulina Geoffroy, 1785: 78 (Leptura) bilineata Scopoli, 1772: 102 (Leptura) decempunctata DeGeer, 1775: 78 (Cerambyx) populi Duméril, 1860: 607 salicis Zetterstedt, 1818: 258

Krivolutskaya & Lobanov, 1996: 132 - Afghanistan.

tribe Tetropini Portevin, 1927 genus Lenotetrops Danilevsky, 2012a: 5 type species Lenotetrops

ivanovae Danilevsky, 2012 ivanovae Danilevsky, 2012a: 6

Danilevsky, 2012a: 6 - Kabul prov., 13 km NW Kabul, Qargha environs, Band-e-Qargha; Wardak prov., W Maydan.

Phytoecia (Parobereina) pashtunica sp.n Figs 1-2

Type locality. Afganistan, Bamian province, Waras district, 5 km SW Waras near Dahane Denawak village, 34°15'42"N, 66°52'43"E, 2680 m.

Description. A single female available; head, antennae and legs totally black; lower eye lobes about two times wider than genae; antennae a little shorter than body, not reaching elytral apices; 1st antennal joint short, about as long as 5 th, 4th joint much longer, but shorter than 3rd; prothorax elongated, about 1.1 times longer than basal width; pronotum shining, glabrous, without glabrous callosities, without pale recumbent pubescence, without longitudinal setae stripe, with small sparse punctation; solitary white setae are distributed along sides of prothorax and under humeral elytral angles; scutellum triangula, without pale recumbent pubescence, with sparse small erect setae; elytra yellow with wide black area along middle; curved elytral margin with narrow black stripe, as well as anterior elytral margin; elytral punctation relatively big arranged in distinct longitudinal rows; elytral pubescence hardly visible, consisting of very short erect and recumbent setae; metepisternae with dense semierect short pale pubescence; claws brown, with very narrow sharp long denticles; ventral body side covered by shorte moderately dense pale pubescence, and looks black; last abdominal tergite rounded, sternite truncated; body length: 6.2 mm, width: 1.3 mm.

Differencial diagnosis. The species is similar to 3 other Ph. (Parobereina) of the region: Ph. (P.) ochraceipennis Kraatz, 1882, Ph. (P.)povolnyi Heyrovsky, 1971, Ph. (P.) tatyanae Skrylnik, 2010, but each of them has partly yellow tibiae or (in Ph. ochraceipennis) partly yellow femora; pronotum in all 3 species usually more or less covered by recumbent pale pubescence, with more or less distinct glabrous shining callosities; anterior elytral margin partly yellow (black tranverse stripe interrupted in the middle; black stripe of curved elytral margin never reaching humeri; pale elyral pubescence rather distinct; ventral body margin with very dense recumbent pubescence, looks white.

Material studied. 1 female, each with 2 labels: 1) "AFGANISTAN 5 km WS Waras / Bamian prov., Waras distr. / near Dahane Denawak vill. 2680 m / 34°15'42.46"N, 66°52'43.20"E / Yu.Ye. Skrylnik 25.VI.2016."; 2) "HOLOTYPUS / Phytoecia (Parobereina) / PASHTUNICA sp.n. / M.Lazarev det. 2019" - collection of M.L. Danilevsky.

Distribution. Central Afganistan, Bamian province.

Etymology. The Pashtuns are the largest ethnic group in

Afghanistan.

Phytoecia (Parobereina) heinzi sp.n

Figs 3-4

Type locality. Pakistan, Khyber Pakhtunkhwa province, Swat district, Utrot, about 35°29'N, 72°28'E, 2300/2600 m. Description. A single male available; head and antennae totally black; lower eye lobes more than two times wider than genae; antennae longer than body, surpassing elytral apices by 2 apical joints; 1st antennal joint short, shorter than 5th, 4th joint much longer than 5th and about equal to 3rd; prothorax about 1.1 times shorter than basal width; pronotum with hardly distinct microsculpture and rather small sparse punctation, with very fine recumbent pale pubescence concentrated along middle forming pale central line poorly pronounced; without glabrous callosities; scutellum semicircula, with dense white recumbent pubescence; elytra black with elongated humeral yellow spots protruding backwards along anterior elytral third; curved elytral margin black; elytral punctation rather small, longitudinal rows are not regular, partly mixed; elytral pubescence hardly visible, consisting of very short erect and recumbent setae; metepistrnae with dense recumbent white pubescence; legs yellow with partly darckened tarsi; claws with very narrow long sharp denticles; ventral body side covered by dense white pubescence, and looks white; pygidium shallowly emarginated, postpygidium rounded; last abdominal sternite slightly exposed apically, concave along middle; body length: 8.2 mm, width: 2.0 mm. Differencial diagnosis. The species is very close to rather variable Ph. (Parobereina) vittipennis Reiche, 1877, because of similarly colored antennae and pronotum, as well as similar pronotal sculpture,

but in Ph. (P.) vittipennis prothorax more elongated, elytral punctation considerably larger strongly longitudinally arranged; many populations of Ph. (P.) vittipennis are characterized by long and dense pronotal and elytral pale erect pubescens; middle and hind femora in all populations of Ph. (P.) vittipennis with black bases; pygidium in the nominative subspecies and in Ph. (P.) v. pravei Plavilstshikov, 1926 deeply emarginated.

Material studied. 1 male, each with 2 labels: 1) "Pakistan (Swat): Utrot 2300/2600 m / Heinz leg. / 15/19.VII.1997"; 2) "HOLOTYPUS / Phytoecia (Parobereina) / HEINZI sp.n. / M.Lazarev det. 2019" - collection of M.L. Danilevsky. Distribution. North Pakistan.

Etimology. The new species is dedicated to Walter Heinz, who collected the holotype.

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AFGHANISTAN 5 km NW Waras Bamian prov., Waras disir. near Daliane Denawak vill. 2680m 34°15'42.46"N, 66°52'43.20"E Yu. Ye. Skrvlnik25.VI.2016

'Pakistan^ (Swat):

Uti -of Z3co/-26oom

//ejhz /eq. I'S.//9M J J9??

HOLOTYPUS Phytoecia (Parobereuta) PASHTUNICA sp. it. M.Lazarev det. 2018

HOLOTYPUS Phytoecia (Parobereuta) HEINZ I s p. n. M.Lazarev det. 2018

2 4

Figs 1-2. Phytoecia (Parobereina) pashtunica sp.n.: 1 - Afganistan, Bamian province, Waras district, 5 km SW Waras near Dahane Denawak village, 34°15'42"N, 66°52'43"E, 2680 m; 2 - Labels of the holotype.

Figs 3-4. Phytoecia (Parobereina) heinzi sp.n.: 3 - Pakistan, Khyber Pakhtunkhwa province, Swat district, Utrot, about 35°29'N, 72°28'E, 2300/2600 m; 4 - Labels of the holotype. Received: 28.12.2018

Accepted: 14.01.2019