CC BY
14УДК 595.768.2
© Amurian zoological journal. I (3), 2009. 200-213.
ANNOTATED CHECKLIST OF FOSSIL AND RECENT SPECIES OF THE FAMILY NEMONYCHIDAE
(COLEOPTERA) FROM THE WORLD FAUNA A.A. Legalov
[Легалов А.А. Аннотированный список вымерших и рецентных видов семейства Nemonychidae (Coleoptera) мировой фауны] Institute of Animal Systematics and Ecology, Siberian Zoological Museum, Frunze street, 11, 630091, Novosibirsk, Russia Институт систематики и экологии животных СО РАН, Сибирский зоологический музей, Фрунзе, 11, 630091, Новосибирск, Россия. E-mail: [email protected]
Key words: Nemonychidae, Coleoptera, world fauna, annotated checklist.
Ключевые слова: Nemonychidae, Coleoptera, мировая фауна, аннотированный список.
Summary. An annotated checklist of the family Nemonychidae is presented, with a key to the supraspecific taxa provided. New subtribe Brarina Legalov, subtrib.n. (type genus: Brarus Kuschel, 1997) of the tribe Mecomacerini Kuschel, 1994, new genera Chinocimberis Legalov, gen.n. (type species: Brenthorrhinoides angustipecteris Liu, Ren, Tan, 2006) and Renicimberis Legalov, gen.n. (type species: Brenthorrhinoides latipeeteris Liu, Ren, Tan, 2006), new subgenus Amerocimberis Legalov, subgen.n. (type species: Rhinomacer pilosus LeConte, 1876) of the genus Cimberis des Gozis, 1881 are described. New synonym Paroxycorynoides L. Ar-noldi, 1977, syn.n. for Distenorrhinus L. Arnoldi, 1977 is presented. New systematic placements are proposed: Metrioxenoides Grat-shev, Zherikhin, Jarzembowski, 1997, placem.n. and Libanorhinus Kuschel & Poinar, 1993, placem.n. are transfered from Nemonychidae incertae sedis to the tribe Mecomacerini; Oxycorynoides lineatipunctatus (L. Arnoldi, 1977), placem.n. and O. retar-datus (Zherikhin, 1993), placem.n. are transfered from the genus Belonotaris to the genus Oxycorynoides. Change of status: Nano-phydini L. Arnoldi, 1977, stat.n. is downgraded from subfamily to tribe. Resurrected status: Procurculionini L. Arnoldi, 1977, stat.res. 6 new combinations are proposed: Distenorrhinus elegans (L. Arnoldi, 1977), comb.n., Oxycorynoides lineatipunctatus (L. Arnoldi, 1977), comb.n., O. retardatus (Zherikhin, 1993), comb.n., Chinocimberis angustipecteris (Liu, Ren, Tan, 2006), comb.n., Ch. magnoculi (Liu, Ren, Tan, 2006), comb.n. and Renicimberis latipeeteris (Liu, Ren, Tan, 2006), comb.n.
Резюме. Составлен систематический список семейства Nemonychidae. Представлен определитель надвидовых таксонов. Описаны новая подтриба Brarina Legalov, subtrib.n. (типовой род: Brarus Kuschel, 1997) трибы Mecomacerini Kuschel, 1994, новые рода Chinocimberis Legalov, gen.n. (типовой вид: Brenthorrhinoides angustipecteris Liu, Ren, Tan, 2006) и Renicimberis Legalov, gen.n. (типовой вид: Brenthorrhinoides latipeeteris Liu, Ren, Tan, 2006), а также новый подрод Amerocimberis Legalov, subgen.n. (типовой вид: Rhinomacer pilosus LeConte, 1876) рода Cimberis des Gozis, 1881. Paroxycorynoides L. Arnoldi, 1977, syn.n. сведен в синонимы к Distenorrhinus L. Arnoldi, 1977. Изменено систематическое положение родов Metrioxenoides Grat-shev, Zherikhin, Jarzembowski, 1997, placem.n. и Libanorhinus Kuschel & Poinar, 1993, placem.n. (из Nemonychidae incertae sedis в трибу Mecomacerini) и видов Oxycorynoides lineatipunctatus (L. Arnoldi, 1977), placem.n. и O. retardatus (Zherikhin, 1993), placem.n. (из рода Belonotaris в род Oxycorynoides). Изменен систематический статус Nanophydini L. Arnoldi, 1977, stat.n. (из подсемейства до трибы). Восстановлен систематический статус Procurculionini L. Arnoldi, 1977, stat.res. Установлены 6 новых комбинаций: Distenorrhinus elegans (L. Arnoldi, 1977), comb.n., Oxycorynoides lineatipunctatus (L. Arnoldi, 1977), comb.n., O. retardatus (Zherikhin, 1993), comb.n., Chinocimberis angustipecteris (Liu, Ren, Tan, 2006), comb.n., Ch. magnoculi (Liu, Ren, Tan, 2006), comb.n. и Renicimberis latipeeteris (Liu, Ren, Tan, 2006), comb.n.
INTRODUCTION
Family Nemonychidae, which had emerged in the Jurassic, is the most primitive family of the superfamily Curculionoidea [Kuschel, 1983; Legalov, 2006; Zherikhin,
1993]. The majority of recent Nemonychidae develop in the sporophylls inside dehiscing male conifer strobili of coniferous [Oberprieler, Marvaldi, Anderson, 2007].
Family Nemonychidae comprises 128 described species in 45 genera; the group is divisible into 6 subfamilies with 8 tribes. 72 recent species of the family Nemonychi-dae are distributed in North Africa, Europe, Caucasus, Asia minor, Middle, Central and North Asia, North, Central and South America, Australia, New Guinea, New Caledonia and New Zealand (fig. 1). The fossil forms (56 described and few undescribed) are known from Europe (Cretaceous: England, Spain), Central (Jurassic: Kazakhstan; Cretaceous: Mongolia) and North Asia (Cretaceous: Russia), North America (Eocene: USA: Green River) and South America (Cretaceous: Brazil) (fig. 2). 46 species
are described from the Jurassic, 7 species from the Cretaceous and 3 species from Jurassic or Cretaceous.
In this study the data from the following works were used: Alonso-Zarazaga & Lyal [1999]; Bedel [18821888]; Arnoldi [1977]; Biondi [1994]; Blatchley & Leng [1916]; Dalla Torre & Voss [1937]; Desbrochers des Loges [1869]; Dieckmann [1974]; Gratshev & Zherichin [1995, 1996, 2000, 2003]; Gratshev, Zherikhin, Jarzembowski [1997]; Hamilton [1983, 1994]; Kizub & Nazarenko [2005]; Kuschel [1954, 1959, 1983, 1989, 1993, 1994, 1995, 2000, 2003]; Kuschel & Poinar [1993]; Kuschel & May [1997]; Lea [1926]; LeConte [1876, 1880]; Legalov [1998, 2002, 2009]; Legalov & Opanas-senko [2000]; Legalov & Sitnikov [2000]; Legalov & Telnov [2004]; Liu, Ren, Shih [2006]; Liu, Ren, Tan [2006]; Morris [1990]; Morrone [1997]; O’Brien & Wibmer [1982]; Opanassenko [1973, 1976]; Ponomarenko, Zherikhin, Kirejtshuk [2004]; Reitter [1916]; Ren [1995]; Schilsky [1903]; Semenov [1900]; Solsky [1880]; Ter-Minassian [1984]; Thompson [1992]; Voss [1922, 1932, 1937, 1952, 1965, 1974]; Wibmer &
Fig. 1. Distribution of recent Nemonychidae.
Рис. 1. Распространение современных видов Nemonychidae.
Fig. 2. Distribution of fossil Nemonychidae.
Circle - locations of the finds of Jurassic taxa, pentagon - locations of the finds of Cretaceous taxa, square - location of the find of Eocene taxa.
Рис. 2. Распространение ископаемых видов Nemonychidae.
Кружок - находки юрских таксонов, пятиугольник - меловых, квадрат - эоценовых таксонов.
O’Brien [1986]; Zherikhin [1986, 1993]; Zherichin &
Gratshev [2003, 2004] etc.
RESULTS Family Nemonychidae Bedel, 1820
Key to supraspecific taxa of the family Nemonychidae
1. Procoxae located in the middle or closer to apical
margin of prothorax. (Brenthorrhininae)............2
- procoxae located closer to basal margin of prothorax ...................................................9
2. Procoxae located closer to apical margin of prothorax. Mandibles narrow. (Procurculionini)...........3
- procoxae located in the middle of prothorax. Mandibles wide. (Brenthorrhinini).......................4
3. 1st-2nd segments of funicle widened. Femora narrow. Pseudobrenthorrhinus
- 1st-2nd segments of funicle not widened. Femora
widened..................................Procurculio
4. Antennae inserted subapically. Rostrum weakly elongated...................................Brenthorrhinus
- antennae inserted on the middle or beyond the middle of rostrum. Rostrum long..........................5
5. Body narrower. Pronotum without lateral carina or
with weak lateral carina. (Distenorrhinus).........6
- body wider. Pronotum with strong lateral carina .... 8
6. Protibiae widened, biconcave. Profemora strongly
widened........................Parabrenthorrhinus
- tibiae narrow, straight. Profemora weakly widened
or not widened.....................................7
7. Larger (greater than 5 mm in length). Rostrum shorter. Asternorrhinus
- smaller (less than 5 mm). Rostrum longer ..........
...............................Distenorrhinus s. str.
8. Anterior margin of prothorax strongly curved. La-
brum elongated.................Microbrenthorrhinus
- anterior margin of prothorax straight. Labrum not
elongated......................Megabrenthorrhinus
9. Antennae inserted in the base third of rostrum or
submedially........................................10
- antennae inserted subapically ..................... 22
10. Antennae inserted in the base third of rostrum. Elytra
without striae. Mandibles inserted dorsally. (Doy-dirhynchinae)................................. 11
- antennae inserted in the middle of rostrum. Elytra
with striae. (Eobelinae) ..........................12
11. Labrum large, subquadrate. Maxillary palps 4-
segmented. 4th ventrite in females with one or two sex patches..............................Lecontellus
- labrum very small, strongly transverse. Maxillary
palps 3-segmented. Ventrites in females without sex patches ............................. Doydirhynchus
12. Metathorax short. 5th ventrite elongated. (Nanophy-
dini) ................................... Nanophydes
- metathorax long. 5th ventrite not elongated........ 13
13. Rostrum thin and long. Pronotum with weak lateral
carinae. Larger (6.2-10.2 mm). (Eobelini)..........14
- rostrum thicker and short. Pronotum with sharper
lateral carinae. Smaller (2.1-5.5 mm). (Oxycorynoidi-ni)........................................... 18
14. Profemora not widened......................Eobelus
- profemora widened ................................ 15
15. 1st segment of protarsi widened and elongated. Archaeorrhynchus
- 1st segment of protarsi of usual shape................16
16. Metatibiae biconcave, with long mucro......Probelus
- metatibiae almost straight or weakly curved, without
long mucro.........................................17
17. Rostrum straight. Elytra flattened......Probelopsis
- rostrum curved. Elytra weakly convex ... Belonotaris
18. Body elongated........................Microprobelus
- body wider ....................................... 19
19. Tibiae curved.........................Scelocamptus
- tibiae straight .................................. 20
20. Profemora widened..........................Ampliceps
- profemora not widened.............................21
21. Eyes small. Tarsi not elongated.......Oxycorynoides
- eyes large. Tarsi elongated...........Cratomacer
22. Elytra without striae. Claws with or without teeth...
...................................................23
- elytra with distinct striae. Claws with teeth. (Rhino-
rhynchinae)........................................28
23. Rostrum short. Claws with teeth. Mesocoxal cavities open laterally to pleurites. (Nemonychinae) ...............................................Nemonyx
- rostrum long. Claws without teeth. Mesocoxal cavities
closed. (Cimberindinae)............................24
24. Pronotum wide.........................Chinocimberis
- pronotum narrower.................................25
25. Mandibles curved at the outer edge in dorsal and
lateral view, each armed with a strong tooth at the inner edge. (Cimberis) *........................26
- mandibles angulate at the outer edge in dorsal or lateral view, in both unarmed at the inner edge ...................................................27
26. Antennae inserted near apex of rostrum in males and females. Labrum almost rectangular ..........................................Cimberis s. str.
- antennae inserted closer to apex of rostrum in males than in females. Labrum almost trapezoidal ..........................................Amerocimberis
27. Labrum trapezoidal or broadly rounded at apex.
Setiferous sex patches usually present at one or two ventrites in females*....................Pityomacer
- labrum triangular, pointed at apex. Setiferous sex
patches absent in females...................Acromacer
28. Antennae inserted closer to apex of rostrum. Labrum
with 3 or more pairs of setae. (Mecomacerini)......29
- antennae inserted further from apex of rostrum. Labrum
with 2 pairs of setae. (Rhinorhynchini)............39
29. Rostrum reduced. Tibiae widened. (Brarina) .... Brarus
- rostrum long. Tibiae not widened. (Mecomacerina) .. ...................................................30
30. Pronotum wide. Striae of elytra weak .... Renicimberis
- pronotum narrower.................................31
31. Points in elytral striae large....................32
- points in elytral striae small....................33
32. Rostrum long. Eyes smaller. Head and pronotum
small punctuate ....................... Libanorhinus
- rostrum short. Eyes larger. Head and pronotum
largely punctuate......................Metrioxenoides
33. Rostrum much longer than pronotum
.....................................Rhynchitoplesius
- rostrum shorter than pronotum or of equal length...34
34. Rostrum narrowed before basis....Rhynchitomacerinus
- rostrum not narrowed before basis.................. 35
35. Rostrum not widened at apex, stronger flattened ... 36
- rostrum widened at apex, weaker flattened..........37
36. Frons considerably broader than rostrum behind an-
tennal insertions. Rostrum distinctly saddled against frons, cylindrical at middle of postrostrum, more than 1.7 times longer than apical width. Elytral colouring alike in both sexes**.....................Eutactobius
- frons as broad as width of rostrum behind antennal
insertions. Rostrum not saddled (not depressed in lateral view) against frons, tapering toward apex in dorsal and lateral views, less than 1.7 times longer than apical width. Elytra uniformly black in S and at least partly reddish brown in $ ............Bunyaeus
37. Pronotum transversal....................Aragomacer
- pronotum longer than wide..........................38
38. Rostrum narrower, stronger widened at apex ............................................Mecomacer
- rostrum wider, weaker widened at apex .... Notomacer
39. Rostrum strongly widened at apex. Antennae inserted
closer to rostrum apex...............Rhynchitomacer
- rostrum weakly widened at apex. Antennae inserted
further to rostrum apex.............................40
40. Mandibles bifurcated at apex ........... Nannomacer
- mandibles acuminate at apex........................41
41. Frons very wide....................................42
- frons less widened.................................43
42. Prosternal process in males directed forward. Larger
(3.0-4.2 mm). Body wider..................Atopomacer
- prosternal process in males directed to procoxae. Smaller (1.5-2.7 mm). Body narrower ... Rhinorhynchus
43. Mesonotum with 2 symmetrical stridulatory files** ............................................ Basiliorhinus
- mesonotum with 1 stridulatory file or 2 asymmetrical files 44
44. Rostrum deeply saddled at the base. Postrostrum with
forward-directed pubescence. Frons much wider between eyes than apex of rostrum. Internal sac of aedeagus with 1 pair of sclerites**.........Basiliogeus
- rostrum shallowly saddled at the base. Postrostrum
with pubescence directed backwards. Frons not wider between eyes than apex of rostrum. Internal sac of aedeagus with 2 pairs of sclerites..........Pagomacer
*by Kuschel [1989]
**by Kuschel [1994]
Subfamily Eobelinae L. Arnoldi, 1977
Eobelidae L. Arnoldi, 1977: 144
Tribe Eobelini L. Arnoldi, 1977
Eobelidae L. Arnoldi, 1977: 144
Genus Archaeorrhynchus Martynov, 1926
Archaeorrhynchus Martynov, 1926: 23
Type species: Archaeorrhynchus tenuicornis Martynov,
1926
Archaeorrhynchus acutirostris L. Arnoldi, 1977
Archaeorrhynchus acutirostris L. Arnoldi, 1977: 148
Distribution. Late Jurassic (Kazakhstan: Karatau). Archaeorrhynchus latitarsis L. Arnoldi, 1977
Archaeorrhynchus latitarsis L. Arnoldi, 1977: 150 Distribution. Late Jurassic (Kazakhstan: Karatau). Archaeorrhynchus paradoxopus L. Arnoldi, 1977 Archaeorrhynchus paradoxopus L. Arnoldi, 1977: 149 Distribution. Late Jurassic (Kazakhstan: Karatau). Archaeorrhynchus tenuicornis Martynov, 1926
Archaeorrhynchus tenuicornis Martynov, 1926: 25 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Belonotaris L. Arnoldi, 1977
Belonotaris L. Arnoldi, 1977: 154
Type species: Belonotaris punctatissimus L. Arnoldi,
1977
Belonotaris karatavicus L. Arnoldi, 1977
Belonotaris karatavicus L. Arnoldi, 1977: 156 Distribution. Late Jurassic (Kazakhstan: Karatau). Belonotaris punctatissimus L. Arnoldi, 1977 Belonotaris punctatissimus L. Arnoldi, 1977: 155 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Eobelus L. Arnoldi, 1977 (Col. pl. I, a) Eobelus L. Arnoldi, 1977: 147 Type species: Eobelus longipes L. Arnoldi, 1977
Eobelus longipes L. Arnoldi, 1977
Eobelus longipes L. Arnoldi, 1977: 147 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Probelopsis L. Arnoldi, 1977
Probelopsis L. Arnoldi, 1977: 153
Type species: Probelopsis acutiapex L. Arnoldi, 1977
Probelopsis acutiapex L. Arnoldi, 1977
Probelopsis acutiapex L. Arnoldi, 1977: 154 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Probelus L. Arnoldi, 1977
Probelus L. Arnoldi, 1977: 151
Type species: Probelus curvispinus L. Arnoldi, 1977
Probelus curvispinus L. Arnoldi, 1977
Probelus curvispinus L. Arnoldi, 1977: 151 Distribution. Late Jurassic (Kazakhstan: Karatau). Probelus longitarsus L. Arnoldi, 1977 Probelus longitarsus L. Arnoldi, 1977: 152 Distribution. Late Jurassic (Kazakhstan: Karatau). Probelus tibialis L. Arnoldi, 1977 Probelus tibialis L. Arnoldi, 1977: 152 Distribution. Late Jurassic (Kazakhstan: Karatau).
Tribe Oxycorynoidini L. Arnoldi, 1977
Oxycorynoidinae L. Arnoldi, 1977: 159
Genus Ampliceps L. Arnoldi, 1977
Ampliceps L. Arnoldi, 1977: 166
Type species: Ampliceps dentitibia L. Arnoldi, 1977
Ampliceps dentitibia L. Arnoldi, 1977
Ampliceps dentitibia L. Arnoldi, 1977: 166
Distribution. Late Jurassic (Kazakhstan: Karatau). Ampliceps furcitibia L. Arnoldi, 1977
Ampliceps furcitibia L. Arnoldi, 1977: 167 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Cratomacer Zherichin & Gratshev, 2004
Cratomacer Zherichin & Gratshev, 2004: 59 Type species: Cratomacer immersus Zherichin & Gratshev, 2004
Cratomacer ephippiger Zherichin & Gratshev, 2004
Cratomacer ephippiger Zherichin & Gratshev, 2004: 61 Distribution. Early Cretaceous (Brazil: Santana). Cratomacer immersus Zherichin & Gratshev, 2004 Cratomacer immersus Zherichin & Gratshev, 2004: 60 Distribution. Early Cretaceous (Brazil: Santana).
Genus Oxycorynoides L. Arnoldi, 1977 (Col. pl. I, d) Oxycorynoides L. Arnoldi, 1977: 159 Type species: Oxycorynoides similis L. Arnoldi, 1977
Oxycorynoides brevipes L. Arnoldi, 1977
Oxycorynoides brevipes L. Arnoldi, 1977: 160 Distribution. Late Jurassic (Kazakhstan: Karatau). Oxycorynoides lineatipunctatus (L. Arnoldi, 1977), comb.n., placem.n.
Belonotaris lineatipunctatus L. Arnoldi, 1977: 155 Distribution. Late Jurassic (Kazakhstan: Karatau). Remarks. This species and Belonotaris retardatus belong to the genus Oxycorynoides judging by shorter rostrum and by small size of body.
Oxycorynoides mongolicus Zherikhin, 1986 Oxycorynoides mongolicus Zherikhin, 1986: 107 Distribution. Early Cretaceous (Mongolia). Oxycorynoides ponomarenkoi L. Arnoldi, 1977 Oxycorynoides ponomarenkoi L. Arnoldi, 1977: 161 Distribution. Late Jurassic (Kazakhstan: Karatau). Oxycorynoides retardatus (Zherikhin, 1993), comb.n., placem.n.
Belonotaris retardatus Zherikhin, 1993: 21 Distribution. Early Cretaceous (Khabarovskii krai: Khetana).
Oxycorynoides rohdendorfi L. Arnoldi, 1977
Oxycorynoides rohdendorfi L. Arnoldi, 1977: 161 Distribution. Late Jurassic (Kazakhstan: Karatau). Oxycorynoides similis L. Arnoldi, 1977 Oxycorynoides similis L. Arnoldi, 1977: 159 Distribution. Late Jurassic (Kazakhstan: Karatau). Oxycorynoides zherichini L. Arnoldi, 1977 Oxycorynoides zherichini L. Arnoldi, 1977: 162 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Microprobelus Liu, Ren, Shih, 2006
Microprobelus Liu, Ren, Shih, 2006: 885
Type species: Microprobelus liuae Liu, Ren, Shih, 2006
Microprobelus liuae Liu, Ren, Shih, 2006
Microprobelus liuae Liu, Ren, Shih, 2006: 886 Distribution. Late Jurassic (China: Liaoning Prov.).
Genus Scelocamptus L. Arnoldi, 1977
Scelocamptus L. Arnoldi, 1977: 164
Type species: Scelocamptus tenuirostris L. Arnoldi, 1977
Scelocamptus curvipes L. Arnoldi, 1977
Scelocamptus curvipes L. Arnoldi, 1977: 165 Distribution. Late Jurassic (Kazakhstan: Karatau). Scelocamptus tenuirostris L. Arnoldi, 1977 Scelocamptus tenuirostris L. Arnoldi, 1977: 164 Distribution. Late Jurassic (Kazakhstan: Karatau).
Tribe Nanophydini L. Arnoldi, 1977, stat.n.
Nanophydinae L. Arnoldi, 1977: 173
Genus Nanophydes L. Arnoldi, 1977 (Col. pl. I, c) Nanophydes L. Arnoldi, 1977: 173 Type species: Nanophydes ovatus L. Arnoldi, 1977
Nanophydes ovatus L. Arnoldi, 1977
Nanophydes ovatus L. Arnoldi, 1977: 173 Distribution. Late Jurassic (Kazakhstan: Karatau).
Subfamily Doydirhynchinae Pierce, 1916 Doydirhynchoidea Pierce, 1916: 463 Type genus: Doydirhynchus Dejean, 1821
Genus Doydirhynchus Dejean, 1821
Doydirhynchus Dejean, 1821: 79
Type species: Rhynchites austriacus Olivier, 1807
Diodyrhynchus Schoenherr, 1833: 240
Type species: Rhynchites austriacus Olivier, 1807
Doedycorhynchus Labram & Imhoff, 1843: 3
Type species: Rhynchites austriacus Olivier, 1807
Doedycorrhynchus Bedel, 1883: 18
Type species: Rhynchites austriacus Olivier, 1807
Doydirhynchus austriacus (Olivier, 1807)
Rhynchites austriacus Olivier, 1807: 27 Doydirhynchus austriacus v. castaneus Germar, 1833:
242
Diodyrhynchus karamani Stierlin, 1886: 229; Diodyrrhynchus austriacus cilicicus Daniel & Daniel, 1903: 327;
Diodyrhynchus austriacus v. lutescens Schilsky, 1903: 92 Diodyrrhynchus austriacus v. pallidicolor Pic, 1905: 190 Diodyrrhynchus austriacus v. fulvipennis Reitter, 1916: 260
Diodyrrhynchus austriacus f. testacea Voss, 1932: 72 Distribution. Western Palaearctic.
Host plants. Pinus halepensis, P. pinaster, P. sylvestris, P. uncinata [Dieckmann, 1974; Kuschel, 1993]. Doydirhynchus bicolor Pic, 1905 Diodyrrhynchus austriacus v. bicolor Pic, 1905: 190 Diodyrrhynchus austriacus v. testaceus Pic, 1905: 190 Distribution. Northern Africa.
Host plants. Unknown.
Genus Lecontellus Kuschel, 1989
Lecontellus Kuschel, 1989: 125
Type species: Doydirhynchus byturoides LeConte, 1880
Lecontellus byturoides (LeConte, 1880)
Dioyrhynchus byturoides LeConte, 1880: 215 Distribution. USA.
Host plants. Pinus ponderosa, P. radiata [Kuschel, 1989].
Lecontellus pinicola Kuschel, 1989
Lecontellus pinicola Kuschel, 1989: 156 Distribution. USA (California).
Host plants. Pinussabiniana [Kuschel, 1989].
Lecontellus sleveni (Martin, 1930)
Diodyrhynchus sleveni Martin, 1930: 130 Distribution. USA (California).
Host plants. Unknown.
Subfamily Nemonychinae Bedel, 1820
Nemonychidae Bedel, 1820: 3
Type genus: Nemonyx Redtenbacher, 1845
Genus Nemonyx Redtenbacher, 1845
Nemonyx Redtenbacher, 1845: 96
Type species: Rhinomacer lepturoides Fabricius, 1801
Nematonyx Agassiz, 1846: 247
Type species: Rhinomacer lepturoides Fabricius, 1801
Nemonyx canescens Solsky, 1881
Nemonyx canescens Solsky, 1881: 263 Distribution. Kazakhstan, Uzbekistan, Turkmenistan. Host plants. Consolida camptocarpa [Ter-Minassian, 1984; Kuschel, 1993].
Nemonyx lepturoides (Fabricius, 1801)
Rhinomacer lepturoides Fabricius, 1801: 429 Nemonyx grisescens Reitter, 1899: 209 Distribution. Western Palaearctic.
Host plants. Consolida regalis, C. paniculata, C. divariacata [Dieckmann, 1974; Ter-Minassian, 1984; Biondi, 1994; Kuschel, 1993].
Nemonyx scutellatus Abeille, 1901 Nemonyx scutellatus Abeille, 1901: 234 Distribution. Tunisia.
Host plants. Unknown.
Nemonyx semirufus Pic, 1898 Nemonyx semirufus Pic, 1898: 74 Nemonyx variicolor Abeille, 1898: 258 Distribution. Algeria.
Host plants. Delphiniumperegrinum [Kuschel, 1993].
Subfamily Cimberindinae des Gozis, 1882
Cimberidae des Gozis, 1882: 58 Type genus: Cimberis des Gozis, 1881
Tribe Cimberindini des Gozis, 1882
Cimberidae des Gozis, 1882: 58 Type genus: Cimberis des Gozis, 1881
Genus Chinocimberis Legalov, gen.n.
Type species: Brenthorrhinoides angustipecteris Liu,
Ren, Tan, 2006
Remarks. For the description, see that of Brenthorrhin-oides angustipecteris and B. magnoculi [Liu, Ren, Tan, 2006: 608-609.
Diagnosis. The new genus is close to genus Cimberis but differs by the wide pronotum.
Etymology. The name is formed from the words “Chinese” and “cimberis”.
Chinocimberis angustipecteris (Liu, Ren, Tan, 2006), comb.n.
Brenthorrhinoides angustipecteris Liu, Ren, Tan, 2006: 607
Distribution. Late Jurassic or Early Cretaceous (China: Liaoning Prov.).
Chinocimberis magnoculi (Liu, Ren, Tan, 2006), comb.n.
Brenthorrhinoides magnoculi Liu, Ren, Tan, 2006: 607 Distribution. Late Jurassic or Early Cretaceous (China: Liaoning Prov.).
Genus Cimberis des Gozis, 1881
Cimberis des Gozis, 1881: 112
Type species: Rhinomacer attelaboides Fabricius, 1787
Rhinomacer Olivier, 1807: 457 nec Geoffroy, 1762
Type species: Rhinomacer attelaboides Fabricius, 1787
Neocimberis O'Brien & Wibmer, 1982: 3, 18
Type species: Rhinomacer attelaboides Fabricius, 1787
Subgenus Cimberis s. str.
Cimberis (Cimberis) attelaboides (Fabricius, 1787)
Rhinomacer attelaboides Fabricius, 1787: 123 Curculio rhinomacer Paykull, 1792: 126 Cimberis attelaboides v. canescens Semenov, 1900: 129 Remarks. Male from the collection of Zoological Institute, Russian Academy of Sciences, St. Petersburg, with label “N. Korea, prov., South Hampyong, Toxong-Omdonzi, 1800 m, 27.V.1990, S.V. Murzin” is the easternmost record of Nemonychidae in Eurasia. Distribution. Eurasia.
Host plants. Pinus nigra, P. pinaster, P. sylvestris, P. uncinata [Dieckmann, 1974; Opanassenko, 1976; Perris, 1856; Kuschel, 1993].
Cimberis (Cimberis) elongata (LeConte, 1876)
Rhinomacer elongatus LeConte, 1876: 2 Distribution. Canada, USA.
Host plants. Pinus banksiana, P. contorta, P. strobes, P. taeda, P. virginiana [Kuschel, 1989].
Cimberis (Cimberis) pallidipennis (Blatchley & Leng, 1916)
Rhinomacer pallidipennis Blatchley & Leng, 1916: 50 Distribution. Canada, USA.
Host plants. Unknown.
Cimberis (Cimberis) decipiens Kuschel, 1989
Cimberis decipiens Kuschel, 1989: 133 Distribution. Canada, USA.
Host plants. Pinus monticola [Kuschel, 1989].
Subgenus Amerocimberis Legalov, subgen.n.
Type species: Rhinomacer pilosus LeConte, 1876 Description. Body brown or black, with semierect light setae. Labrum almost trapezoidal. Rostrum long, thin, weakly curved, small punctate, without carina. Apex widened, sparsely punctate, flattened. Antennae inserted closer to apex of rostrum in males than in females. Frons wide, convex, small and densely punctate. Eyes strongly convex. Vertex densely and small punctate. Temples short. Antennae long, reaching humeri. Scapus and funicle segments trapezoidal, more or less elongated. Clava not compact, elongated. 1st and 2nd segments al-
most trapezoidal. 3rd segment tear-shaped, longer than 2nd segment, weakly pointed. Pronotum almost rectangular, of equal length and width or weakly transversal, weakly narrowed to basis and apex, lustrous, small and densely punctate. Greatest width in middle. Sides almost direct. Scutellum almost rectangular, small. Elytra elongated. Humeri weakly convex. Greatest width in middle or behind middle. Intervals wide, flat, lustrous, punctate. Scutellar striole and striae absent. Thorax small and densely punctate. Prothorax in males with weak prosternal process before procoxae. Precoxal part longer than post-coxal part. Metepisternum narrow. Abdomen convex, small punctate. 1st and 2nd ventrite longer. 3rd-5th ven-trite shorter. 3rd and 4th ventrites in females with sex patches. 5th ventrite wide, with weak impression in males. Legs long. Femora weakly widened. Protibiae almost straight, weakly widened to apex. Meso- and metatibiae shorter, weakly curved. Tarsi long. 1st segment elongated. 2nd segment triangular. 3rd segment bilobed. Clausal segment elongated. Claws with teeth. Length of body: 2.0-4.2 mm.
Diagnosis. The new subgenus differs from subgenus Cimberis s. str. by the antennae inserted closer to apex of rostrum in males than in females and almost trapezoidal labrum.
Etymology. The name is derived from the words “American” and “cimberis”.
Cimberis (Amerocimberis) pilosa (LeConte, 1876) Rhinomacer pilosus LeConte, 1876: 2 Distribution. Canada, USA.
Host plants. Pinus banksiana, P. palustris, P. taeda, P. virginiana [Kuschel, 1989].
Cimberis (Amerocimberis) compta (LeConte, 1876)
Rhinomacer comptus LeConte, 1876: 2 Cimberis parvulus Hatch, 1971: 336 Distribution. Canada, USA.
Host plants. Pinus ponderosa, P. sibiniana, P. virginiana [Kuschel, 1989].
Cimberis (Amerocimberis) bihirsuta (Hatch, 1971)
Rhinomacer bihirsuta Hatch, 1971: 336 Distribution. Canada, USA.
Host plants. Pinus contorta, P. ponderosa [Kuschel, 1989].
Cimberis (Amerocimberis) turbans Kuschel, 1989
Cimberis turbans Kuschel, 1989: 133 Distribution. USA.
Host plants. Pinus contorta [Kuschel, 1989].
Genus Pityomacer Kuschel, 1989
Pityomacer Kuschel, 1989: 125
Type species: Pityomacer carmelites Kuschel, 1989
Pityomacer carmelites Kuschel, 1989
Pityomacer carmelites Kuschel, 1989: 147 Distribution. USA (California).
Host plants. Unknown.
Pityomacer pix Kuschel, 1989 Pityomacer pix Kuschel, 1989: 147 Distribution. Canada, USA.
Host plants. Unknown.
Pityomacer nugax Kuschel, 1989 Pityomacer nugax Kuschel, 1989: 147
Distribution. USA (California).
Host plants. Pinus sp. [Kuschel, 1989].
Genus Acromacer Kuschel, 1989
Acromacer Kuschel, 1989: 125
Type species: Rhinomacer bombifrons (LeConte, 1876)
Acromacer bombifrons (LeConte, 1876)
Rhinomacer bombifrons LeConte, 1876: 412 Distribution. Canada, USA.
Host plants. Pinus contorta, P. jeffreyi [Kuschel, 1989].
Subfamily Rhinorhynchinae Voss, 1922
Rhinorhynchini Voss, 1922: 2
Type genus: Rhinorhynchus Sharp, 1882
Tribe Mecomacerini Kuschel, 1994
Mecomacerini Kuschel, 1994: 576 Type genus: Mecomacer Kuschel, 1954 Remarks. Brenthorrhinoides latipeeteris Liu, Ren, Tan, 2006, Metrioxenoides pusillus Gratshev, Zherikhin, Jarzembowski, 1997 and Libanorhinus succinus Kuschel & Poinar, 1993 belong to the tribe Mecomacerini because of morphological characters, the antennal attachment in the first place.
Genus Renicimberis Legalov, gen.n.
Type species: Brenthorrhinoides latipeeteris Liu, Ren, Tan, 2006: 607
Remarks. For the description, see that of Brenthorrhin-oides latipeeteris [Liu, Ren, Tan, 2006: 608.
Diagnosis. The new genus differs from Libanorhinus and Metrioxenoides by the wide pronotum and weak striae of elytra.
Etymology. The new genus is named in honour of D. Ren.
Renicimberis latipeeteris (Liu, Ren, Tan, 2006), comb.n.
Brenthorrhinoides latipeeteris Liu, Ren, Tan, 2006: 607 Distribution. Late Jurassic or Early Cretaceous (China: Liaoning Prov.).
Genus Metrioxenoides Gratshev, Zherikhin, Jarzembowski, 1997, placem.n.
Metrioxenoides Gratshev, Zherikhin, Jarzembowski,
1997: 323
Type species: Metrioxenoides pusillus Gratshev, Zherikhin, Jarzembowski, 1997
Metrioxenoides pusillus Gratshev, Zherikhin, Jarzembowski, 1997
Metrioxenoides pusillus Gratshev, Zherikhin,
Jarzembowski, 1997: 324
Distribution. Early Cretaceous (England).
Genus Libanorhinus Kuschel & Poinar, 1993, placem.n.
Libanorhinus Kuschel & Poinar, 1993: 144
Type species: Libanorhinus succinus Kuschel & Poinar,
1993
Libanorhinus succinus Kuschel & Poinar, 1993
Libanorhinus succinus Kuschel & Poinar, 1993: 144 Distribution. Early Cretaceous (Lebanon, Amber).
Genus Mecomacer Kuschel, 1954
Mecomacer Kuschel, 1954: 104
Type species: Mecomacer scambus Kuschel, 1954
Mecomacer collaris (Voss, 1952)
Rhynchitomacer collaris Voss, 1952: 177 Distribution. Argentina, Chile.
Host plants. Araucaria araucana [Kuschel, 1959]. Mecomacer hirticeps Kuschel, 1954 Mecomacer hirticeps Kuschel, 1954: 109 Distribution. Chile.
Host plants. Araucaria araucana [Kuschel, 2000]. Mecomacer ruficornis Kuschel, 1954 Mecomacer ruficornis Kuschel, 1954: 109 Distribution. Chile.
Host plants. Araucaria araucana [Kuschel, 2000]. Mecomacer scambus Kuschel, 1954 Mecomacer scambus Kuschel, 1954: 108 Distribution. Argentina, Chile.
Host plants. Araucaria araucana [Kuschel, 2000].
Genus Notomacer Kuschel, 1994
Notomacer Kuschel, 1994: 577
Type species: Notomacer araucariae Kuschel, 1994
Notomacer araucariae Kuschel, 1994
Notomacer araucariae Kuschel, 1994: 581 Distribution. New Caledonia.
Host plants. Araucaria birmulata [Kuschel, 1994]. Notomacer australiae (Lea, 1926)
Rhinomacer australiae Lea, 1926: 362 Distribution. Australia.
Host plants. Agathis robustus, A. atropurpurea [Kuschel,
1994].
Notomacer brittoni Kuschel, 1994
Notomacer brittoni Kuschel, 1994: 582 Distribution. Australia.
Host plants. Unknown.
Notomacer caledonicus Kuschel, 1994 Notomacer caledonicus Kuschel, 1994: 581 Distribution. New Caledonia.
Host plants. Araucaria culumnaris (Kuschel, 1994). Notomacer eximius Kuschel, 1994 Notomacer eximius Kuschel, 1994: 580 Distribution. Australia.
Host plants. Araucaria cunninghamii [Kuschel, 1994]. Notomacer hirtulus Kuschel, 1994 Notomacer hirtulus Kuschel, 1994: 580 Distribution. Australia.
Host plants. Araucaria biramulata [Kuschel, 1994]. Notomacer reginae Kuschel, 1994 Notomacer reginae Kuschel, 1994: 592 Distribution. Australia.
Host plants. Araucaria cunninghamii [Kuschel, 1994]. Notomacer zimmermani Kuschel, 1994 Notomacer zimmermani Kuschel, 1994: 581 Distribution. Australia.
Host plants. Araucaria cunninghamii [Kuschel, 1994].
Genus Aragomacer Kuschel, 1994
Aragomacer Kuschel, 1994: 577
Type species: Aragomacer leai Kuschel, 1994
Aragomacer grayi Kuschel, 1994
Aragomacer grayi Kuschel, 1994: 601 Distribution. Papua New Guinea.
Host plants. Araucaria cunninghamii [Kuschel, 1994]. Aragomacer leai Kuschel, 1994 Aragomacer leai Kuschel, 1994: 601 Distribution. Australia.
Host plants. Agathis robusta, A. atropurpurea, Araucaria cunninghamii [Kuschel, 1994].
Aragomacer munus Kuschel, 1994 Aragomacer munus Kuschel, 1994: 601 Distribution. Papua New Guinea.
Host plants. Araucaria hunsteinii [Kuschel, 1994]. Aragomacer papuae Kuschel, 1994 Aragomacer papuae Kuschel, 1994: 601 Distribution. Papua New Guinea.
Host plants. Araucaria cunninghamii [Kuschel, 1994]. Aragomacer uniformis Kuschel, 1994 Aragomacer uniformis Kuschel, 1994: 601 Distribution. Australia.
Host plants. Araucaria cunninghamii [Kuschel, 1994].
Genus Eutactobius Kuschel, 1994
Eutactobius Kuschel, 1994: 577
Type species: Eutactobius puellus Kuschel, 1994
Eutactobius puellus Kuschel, 1994
Eutactobius puellus Kuschel, 1994: 610 Distribution. Asutralia.
Host plants. Araucaria cunninghamii [Kuschel, 1994].
Genus Bunyaeus Kuschel, 1994
Bunyaeus Kuschel, 1994: 578
Type species: Bunyaeus monteithi Kuschel, 1994
Bunyaeus eutactae Kuschel, 1994
Bunyaeus eutactae Kuschel, 1994: 612 Distribution. Asutralia.
Host plants. Araucaria cunninghamii [Kuschel, 1994]. Bunyaeus monteithi Kuschel, 1994 Bunyaeus monteithi Kuschel, 1994: 612 Distribution. Asutralia.
Host plants. Araucaria bidwilii [Kuschel, 1994].
Genus Rhynchitomacerinus Kuschel, 1954
Rhynchitomacerinus Kuschel, 1954: 104
Type species: Rhynchitomacer kuscheli Voss, 1952
Rhynchitomacerinus kuscheli (Voss, 1952)
Rhynchitomacer kuscheli Voss, 1952: 177 Distribution. Argentina, Chile.
Host plants. Araucaria araucana [Kuschel, 1954].
Genus Rhynchitoplesius Voss, 1952
Rhynchitoplesius Voss, 1952: 177
Type species: Rhynchitomacer eximius Voss, 1937
Rhynchitoplesius eximius (Voss, 1937)
Rhynchitomacer eximius Voss, 1937: 202 Distribution. Brazil.
Host plants. Unknown.
Subtribe Brarina Legalov, subtrib.n.
Type genus: Brarus Kuschel, 1997
Remarks. For the description, see that of Brarus
[Kuschel & May, 1997: 16-18].
Diagnosis. The new subtribe Brarina differs from nominative subtribe by the reduced rostrum and widened tibiae.
Genus Brarus Kuschel, 1997
Brarus Kuschel, 1997: 16
Type species: Brarus mystes Kuschel, 1997
Brarus mystes Kuschel, 1997
Brarus mystes Kuschel, 1997: 19 Distribution. Brazil.
Host plants. Araucaria angustifolia [Kuschel, May,
1997].
Tribe Rhinorhynchini Voss, 1922
Rhinorhynchini Voss, 1922: 2 Type genus: Rhinorhynchus Sharp, 1882 Rhynchitomacerini May, 1993: 15 Type genus: Rhynchitomacer Voss, 1937
Genus Atopomacer Kuschel, 1989
Atopomacer Kuschel, 1989: 125
Type species: Atopomacer ites Kuschel, 1989
Atopomacer ites Kuschel, 1989 Atopomacer ites Kuschel, 1989: 127 Distribution. USA.
Host plants. Unknown.
Atopomacer hondurasensis Legalov, 2009 Atopomacer hondurasensis Legalov, 2009: 51 Distribution. Honduras.
Host plants. Unknown.
Atopomacer hoplites Kuschel, 1989 Atopomacer hoplites Kuschel, 1989: 127 Distribution. Mexico.
Host plants. Pinus culminicola [Kuschel, 1989]. Atopomacer orites Kuschel, 1989 Atopomacer orites Kuschel, 1989: 127 Distribution. Mexico.
Host plants. Pinus culminicola [Kuschel, 1989].
Genus Rhinorhynchus Sharp, 1882
Rhinorhynchus Sharp, 1882: 88
Type species: Rhinorhynchus zealandicus Sharp, 1882
Rhinorrhynchus Kirby, 1884: 102
Type species: Rhinorhynchus zealandicus Sharp, 1882
Rhinomacer subgen. Listrorhinus Voss, 1922: 9
Type species: Rhinomacer rufulus Broun, 1880
Rhinorhynchus halli Kuschel, 2003
Rhinorhynchus halli Kuschel, 2003: 21 Distribution. New Zealand.
Host plants. Phyllocladus alpinus [Kuschel, 2003].
Rhinorhynchus halocarpi Kuschel, 2003
Rhinorhynchus halocarpi Kuschel, 2003: 20 Distribution. New Zealand.
Host plants. Halocarpus bidwillii, Phyllocladus alpinus [Kuschel, 2003].
Rhinorhynchus phyllocladi Kuschel, 2003
Rhinorhynchus phyllocladi Kuschel, 2003: 20 Distribution. New Zealand.
Host plants. Phyllocladus alpinus, Ph. trichomanoides, Manoao colensoi, Halocarous bidwillii, Podocarpus totara [Kuschel, 2003].
Rhinorhynchus rufulus (Broun, 1880)
Rhinomacer rufulus Broun, 1880: 467 Rhinorhynchus zealandicus Sharp, 1882: 88 Distribution. New Zealand.
Host plants. Phyllocladus sp., Dacrycarpus sp., Dacrydium sp., Halocarpus sp., Lepidothamnus sp., Mamoao sp., Podocarpus sp., Prumnopitys sp. [Kuschel, 2003].
Genus Nannomacer Kuschel, 1954
Nannomacer Kuschel, 1954: 105
Type species: Rhynchitomacer germaini Voss, 1952
Nannomacer germaini (Voss, 1952)
Rhynchitomacer germaini Voss, 1952: 177 Distribution. Chile.
Host plants. Unknown.
Nannomacer wittmeri Kuschel, 1954
Nannomacer wittmeri Kuschel, 1954: 115 Distribution. Argentina, Chile.
Host plants. Saxegothaea conspicua [Kuschel, 1959].
Genus Basiliorhinus Kuschel, 1994
Basiliorhinus Kuschel, 1994: 588
Type species: Basiliorhinus araucariae Kuschel, 1994
Basiliorhinus araucariae Kuschel, 1994
Basiliorhinus araucariae Kuschel, 1994: 618 Distribution. Australia.
Host plants. Araucaria bidwillii [Kuschel, 1994].
Genus Basilogeus Kuschel, 1994
Basilogeus Kuschel, 1994: 578
Type species: Basilogeus prasinus Kuschel, 1994
Basilogeus prasinus Kuschel, 1994
Basilogeus prasinus Kuschel, 1994: 621 Distribution. Australia.
Host plants. Araucaria bidwillii [Kuschel, 1994]. Basilogeus striatopunctatus (Lea, 1926)
Auletes striatopunctatus Lea, 1926: 352 Distribution. Australia.
Host plants. Araucaria cunninghamii [Kuschel, 1994].
Genus Pagomacer Kuschel, 1994
Pagomacer Kuschel, 1994: 578
Type species: Basilogeus deceptus Kuschel, 1994
Basilogeus deceptus Kuschel, 1994
Basilogeus deceptus Kuschel, 1994: 625 Distribution. Australia.
Host plants. Unknown.
Genus Rhynchitomacer Voss, 1937
Rhynchitomacer Voss, 1937: 201
Type species: Rhynchitomacer flavus Voss, 1937
Stenomacer Kuschel, 1954: 105
Type species: Stenomacer vernus Kuschel, 1954
Rhynchitomacer apionoides Kuschel, 1959
Rhynchitomacer apionoides Kuschel, 1959: 241 Distribution. Argentina, Chile.
Host plants. Unknown.
Rhynchitomacer brevicollis Voss, 1965 Rhynchitomacer brevicollis Voss, 1965: 329 Distribution. Argentina.
Host plants. Unknown.
Rhynchitomacer cortesi Kuschel, 1959 Rhynchitomacer cortesi Kuschel, 1959: 241 Distribution. Chile.
Host plants. Nothofagus oblicua [Kuschel, 1959]. Rhynchitomacer errans Kuschel, 1959
Rhynchitomacer errans Kuschel, 1959: 240 Distribution. Chile.
Host plants. Unknown.
Rhynchitomacer flavus Voss, 1937 Rhynchitomacer flavus Voss, 1937: 201 Distribution. Chile.
Host plants. Unknown.
Rhynchitomacer fuscus (Kuschel, 1954) Stenomacer fuscus Kuschel, 1954: 124 Distribution. Argentina.
Host plants. Unknown.
Rhynchitomacer luridus Kuschel, 1954 Rhynchitomacer luridus Kuschel, 1954: 118 Distribution. Argentina.
Host plants. Unknown.
Rhynchitomacer nigrinus Kuschel, 1954 Rhynchitomacer nigrinus Kuschel, 1954: 118 Distribution. Argentina.
Host plants. Unknown.
Rhynchitomacer nitidus Kuschel, 1959
Rhynchitomacer nitidus Kuschel, 1959: 240 Distribution. Chile.
Host plants. Nothofagus nitida [Kuschel, 1959]. Rhynchitomacer puberulus Kuschel, 1959 Rhynchitomacer puberulus Kuschel, 1959: 241 Distribution. Argentina, Chile.
Host plants. Nothofagus dombeyi [Kuschel, 1959]. Rhynchitomacer rostralis Kuschel, 1959 Rhynchitomacer rostralis Kuschel, 1959: 240 Distribution. Argentina.
Host plants. Unknown.
Rhynchitomacer rufus Kuschel, 1954 Rhynchitomacer rufus Kuschel, 1954: 118 Distribution. Argentina.
Host plants. Unknown.
Rhynchitomacer vernus (Kuschel, 1954) Stenomacer vernus Kuschel, 1954: 124 Distribution. Chile.
Host plants. Araucaria araucana [Kuschel, 1954]. Rhynchitomacer viridulus Kuschel, 1954
Rhynchitomacer viridulus Kuschel, 1954: 118
Rhinomacer frustatus Voss, 1974: 43 Distribution. Argentina, Chile.
Host plants. Unknown.
Subfamily Brenthorrhininae L. Arnoldi, 1977
Brenthorrhininae L. Arnoldi, 1977: 171
Tribe Procurculionini L. Arnoldi, 1977, stat.res.
Procurculionini L. Arnoldi, 1977: 157 Eccoptothoracini L. Arnoldi, 1977: 158 Remarks. This tribe is a monophyletic group. It differs from the tribe Brenthorrhinini by the subapical located procoxae and narrow mandibles. Therefore I restore it from the synonym of the tribe Brenthorrhinini.
Genus Procurculio L. Arnoldi, 1977 (Col. pl. 1, e) Procurculio L. Arnoldi, 1977: 157 Type species: Procurculio fortipes L. Arnoldi, 1977 Eccoptothorax L. Arnoldi, 1977: 158 Type species: Eccoptothorax latipennis L. Arnoldi, 1977
Procurculio fortipes L. Arnoldi, 1977
Procurculio fortipes L. Arnoldi, 1977: 157 Distribution. Late Jurassic (Kazakhstan: Karatau). Procurculio latipennis (L. Arnoldi, 1977)
Eccoptothorax latipennis L. Arnoldi, 1977: 158 Distribution. Late Jurassic (Kazakhstan: Karatau). Procurculio pallens Gratshev & Zherikhin, 1995 Procurculio pallens Gratshev & Zherikhin, 1995: 93 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Pseudobrenthorrhinus Gratshev & Zherikhin, 1996
Pseudobrenthorrhinus Gratshev & Zherikhin, 1996: 114 Type species: Pseudobrenthorrhinus crassicornis Gratshev & Zherikhin, 1996
Pseudobrenthorrhinus crassicornis Gratshev &
Zherikhin, 1996
Pseudobrenthorrhinus crassicornis Gratshev &
Zherikhin, 1996: 114
Distribution. Late Jurassic (Kazakhstan: Karatau). Pseudobrenthorrhinus magnus Gratshev & Zherikhin, 1996
Pseudobrenthorrhinus magnus Gratshev & Zherikhin, 1996: 114
Distribution. Late Jurassic (Kazakhstan: Karatau). Pseudobrenthorrhinus tenuicornis Gratshev &
Zherikhin, 1996
Pseudobrenthorrhinus tenuicornis Gratshev & Zherikhin, 1996: 114
Distribution. Late Jurassic (Kazakhstan: Karatau).
Tribe Brenthorrhinini L. Arnoldi, 1977
Brenthorrhininae L. Arnoldi, 1977: 171 Distenorrhinini L. Arnoldi, 1977: 170
Genus Brenthorrhinus L. Arnoldi, 1977 (Col. pl. I, b) Brenthorrhinus L. Arnoldi, 1977: 172 Type species: Brenthorrhinus mirabilis L. Arnoldi, 1977
Brenthorrhinus brevirostris Gratshev & Zherikhin, 1996
Brenthorrhinus brevirostris Gratshev & Zherikhin, 1996: 115
Distribution. Late Jurassic (Kazakhstan: Karatau). Brenthorrhinus longidigitatus Ren, 1995
Brenthorrhinus longidigitatus Ren, 1995: 90. Distribution. Late Jurassic (Kazakhstan: Karatau). Brenthorrhinus mirabilis L. Arnoldi, 1977 Brenthorrhinus mirabilis L. Arnoldi, 1977: 172 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Distenorrhinus L. Arnoldi, 1977
Distenorrhinus L. Arnoldi, 1977: 170
Type species: Distenorrhinus antennatus L. Arnoldi,
1977
Paroxycorynoides L. Arnoldi, 1977: 168, syn.n.; type species: Paroxycorynoides elegans L. Arnoldi, 1977
Subgenus Distenorrhinus s. str.
Distenorrhinus (Distenorrhinus) angulatus L. Arnoldi, 1977
Distenorrhinus angulatus L. Arnoldi, 1977: 170 Distribution. Late Jurassic (Kazakhstan: Karatau). Distenorrhinus (Distenorrhinus) antennatus L. Arnoldi, 1977
Distenorrhinus antennatus L. Arnoldi, 1977: 171 Distribution. Late Jurassic (Kazakhstan: Karatau). Distenorrhinus (Distenorrhinus) arnoldii Gratshev & Zherikhin, 1995
Distenorrhinus arnoldii Gratshev & Zherikhin, 1995: 85 Distribution. Late Jurassic (Kazakhstan: Karatau). Distenorrhinus (Distenorrhinus) pallidirostris Gratshev & Zherikhin, 1995
Distenorrhinus pallidirostris Gratshev & Zherikhin, 1995: 85
Distribution. Late Jurassic (Kazakhstan: Karatau). Distenorrhinus (Distenorrhinus) rotundicollis Gratshev & Zherikhin, 1995
Distenorrhinus rotundicollis Gratshev & Zherikhin, 1995: 85
Distribution. Late Jurassic (Kazakhstan: Karatau).
Subgenus Parabrenthorrhinus Gratshev & Zherikhin, 1996
Parabrenthorrhinus Gratshev & Zherikhin, 1996: 85 Type species: Parabrenthorrhinus sinuatipes Gratshev & Zherikhin, 1995
Distenorrhinus (Parabrenthorrhinus) sinuatipes
Gratshev & Zherikhin, 1995
Distenorrhinus sinuatipes Gratshev & Zherikhin, 1995: 85
Distribution. Late Jurassic (Kazakhstan: Karatau). Distenorrhinus (Parabrenthorrhinus) xavieri
(Zherichin & Gratshev, 2003)
Distenorrhinus xavieri Zherichin & Gratshev, 2003: 70 Distribution. Late Jurassic (Kazakhstan: Karatau).
Subgenus Astenorrhinus Gratshev & Zherikhin, 1995
Astenorrhinus Gratshev & Zherikhin, 1995: 85
Type species: Distenorrhinus elongatus Gratshev & Zherikhin, 1995
Distenorrhinus (Astenorrhinus) elongatus Gratshev & Zherikhin, 1995
Distenorrhinus elongatus Gratshev & Zherikhin, 1995: 85 Distribution. Late Jurassic (Kazakhstan: Karatau). Distenorrhinus (Astenorrhinus) major Gratshev & Zherikhin, 1995
Distenorrhinus major Gratshev & Zherikhin, 1995: 85 Distribution. Late Jurassic (Kazakhstan: Karatau).
Subgenus incertae sedis Distenorrhinus elegans (L. Arnoldi, 1977), comb.n.
Paroxycorynoides elegans L. Arnoldi, 1977: 168 Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Megabrenthorrhinus Gratshev & Zherikhin, 1996
Megabrenthorrhinus Gratshev & Zherikhin, 1996:
Type species: Megabrenthorrhinus grandis Gratshev & Zherikhin, 1996
Megabrenthorrhinus grandis Gratshev & Zherikhin, 1996
Megabrenthorrhinus grandis Gratshev & Zherikhin, 1996: 115
Distribution. Late Jurassic (Kazakhstan: Karatau). Megabrenthorrhinus longicornis Gratshev & Zherikhin, 1996
Megabrenthorrhinus longicornis Gratshev & Zherikhin, 1996: 115
Distribution. Late Jurassic (Kazakhstan: Karatau).
Genus Microbrenthorrhinus Gratshev & Zherikhin, 2000
Microbrenthorrhinus Gratshev & Zherikhin, 2000b: 40 Type species: Microbrenthorrhinus martinezi Gratshev & Zherikhin, 2000
Microbrenthorrhinus martinezi Gratshev & Zherikhin, 2000
Microbrenthorrhinus martinezi Gratshev & Zherikhin, 2000b: 41
Distribution. Early Cretaceous (Spain: Montsec Range).
ACKNOWLEDGEMENTS
The author is thankful to his colleagues for the help with the work.
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