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17http://zoobank.org/urn:lsid:zoobank.org:pub:6EF06A74-CD28-4B02-BE0F-566F72D8D06F
DOI: 10.24412/2226-0773-2021-10-1-56-69
A new species of the genus Pogonocherus Dejean, 1821 (Coleoptera: Cerambycidae) from China with a redescription of poorly known P. pilosipes (Pic, 1907) as a bases of a new subgenus P. (Neopogonocherus subgen. n.)
M.A. Lazarev
Free Economic Society of Russia, Department of Scientifics Conferences and
All-Russian Projects
Tverskaya str., 22a, Moscow 125009 Russia
e-mail: cerambycidae@bk.ru, humanityspace@gmail.com
Key words: Coleoptera, Cerambycidae, new subgenus, new species, taxonomy, Gansu, Sichuan, China.
Abstract: Pogonocherus (s. str.) paradimidiatus sp. n. close to P. (s. str.) dimidiatus Blessig, 1873 is described from China (Gansu and Sichuan provinces). The distinguishing characters are discussed. P. (s. str.) dimidiatus Blessig, 1873 is recorded from Gansu and Sichuan. A new subgenus is proposed: P. (Neopogonocherus subgen. n.) with type species Pogonochaerus pilosipes Pic, 1907. Pogonocherus (N.) pilosipes (Pic, 1907) is redescribed.
Introduction
The genus Pogonocherus included only 3 Chinese species up to now. P. (Pityphilus) fasciculatus (DeGeer, 1775) distributed from Europe to Eastern Asia and 2 Pogonocherus (s. str.). P. (s. str.) dimidiatus Blessig, 1873 is very common in Russia (from Amur Region to Kuril Islands), all around Korean Peninsula and widely distributed in China to Sichuan Province southwards. P. (Neopogonocherus subgen. n.) pilosipes (Pic, 1907) is a poorly known species described after a single specimen from “Chine Orientale” without exact locality designation and sex determination. Two specimens of new species Pogonocherus (s. str.) paradimidiatus sp. n. described bellow were collected in Gansu and Sichuan by I. Belousov and I. Kabak in 2006 and 2012, one specimen was collected in Sichuan by S. Murzin in 2001.
Materials and methods
All photographs were taken with Canon PowerShot G10 digital camera equipped with Cannon Zoom lens 5X IS 6.1 - 30.5 mm 1:2.8 - 4.5 and microscope AmScope SM745NTP. The illustration were edited with Adobe Photoshop 7.0 and Helicon Focus 3.2.
Results
Pogonocherus (s. str.) paradimidiatus sp. n.
Figs 1-2
Three females only are available; body black, strongly attenuated posteriorly; head with very fine, short recumbent pubescence, without erect setae; frons slightly exposed, strongly transverse; genae wide, about as wide as width of lover eye-lobes; eyes strongly emarginated, about totally divided; the distance between dorsal eye-lobes a little more than apical width of the 1st antennal joint; antennae with numerous oblique setae; antennae reaching elytral apex; 1st antennal joint widened at middle; 2nd joint elongated; 2nd - 11th joints reddish basally; 4th joint half red; 3rd antennal joint about as long as 4th or a little shorter; 5th joint nearly 2 times shorter.
Prothorax transverse, about 1.25 times wider than long, as wide anteriorly as posteriorly, with large lateral tubercles; pronotum with a pair of oblique short central protuberances, depressed in the middle; scutellum semicircular, with black pubescence and contrast white stroke.
Elytra about 1.8 times longer than basal width with long outer apical spines typical for Pogonocherus (s. str.); internal angles of elytral apices totally obliterated; dorsal elytral surface with large irregular punctation along middle; large white transverse bend reaching epipleurae does not reach anterior elytral margin; a pair of central anterior tubercles with several short black setae; humeral and external elytral carinae anteriorly distinct; a pair of central elongated protuberances beyond the middle reddish, bear 2 or 3 black setae tufts each; elytral apices with bright white setae spots before apical spines.
Legs without erect setae; all tibiae with white setae bands and reddish bases.
Abdomen with exposed 5th visible sternite bearing deep small hollow near posterior margin.
Body length: 5.0-7.0 mm, width: 1.9-2.7 mm.
Differential diagnosis. The new species is very close to P. (s.str.) dimidiatus Blessig, 1873 widely distributed in East Asia because of long elytral spines and bright white elytral band, but in P. dimidiatus white elytral band reaches anterior elytral margin; white small spots of fine setae scattered over elytra and legs; hollow of the last visible abdominal sternite in females is reduced to emargination of its posterior border.
P. (s. str.) paradimidiatus sp. n. is similar to European P. (s.str.) hispidulus (Piller, Mitterpacher, 1783), distributed to the East up to Orenburg Region, as white elytral band in P hispidulus does not reach elytral bases; but elytral spines in P. hispidulus much shorter, hollow of the last visible abdominal sternite in females is reduced to emargination of its posterior border; internal angle of elytral apices well developed.
Another China species of Pogonocherus - P. pilosipes (Pic, 1907) is known on the bases of holotype and original description. It has strongly elongated pale-brown body similar to P. perroudi Mulsant, 1839, but without elytral setae tufts.
Type material. Holotype, female, China, Southern Gansu, NNW Kaba vill., 34°11'06"N, 103°22'52"E / 34°11'17"N, 103°22'49"E, 3640-3680 m, 15.06.2006, I. Belousov and I. Kabak leg. - author’s collection; 2 paratypes: female, China, Northern Sichuan, WNW Jiuzhalgou, 33°22'26"N, 103°48'08"E, 3320 m, 22.06.2012, I. Kabak leg. - author’s collection; female, China, Sichuan prov., 40 km W Zhangla, 3600-3700 m, 12-14.7.2001, S. Murzin leg. - collection of S. Murzin (Moscow).
Additional material used for comparison. Pogonocherus dimidiatus: 1 female (fig. 8), Gansu, SSW Woshar 3000 m 34°30'23"N, 104°49'48"E, 17.6.2005, S. Murzin leg. - author’s collection; 1 female (fig. 7), S. Gansu, Tochizi, S. Wudu, 2400 m, 21-24.5.1997, S. Murzin leg. - collection of S. Murzin (Moscow); 1 male (fig. 5), Sichuan, W Heishui, 2500 m, 32°2'47.40"N, 103°1'0.48"E, 3-10.6.2012, S. Murzin leg. - author’s collection; 1 male (fig. 6), Shaanxi, Zhouzhi, Taibaishan nat. park, 1350 m, 30.5.1999, M. Murzin leg. - collection of S.Murzin (Moscow); 1 ex., Primorsky Region, Kamenushka, 25.5.1992, S. Khvylya leg. - collection of V. Ustinov (Moscow); 1 ex., Primorsky Region, Kavalerovo, N. Luzhki 1.7.1997 - collection of V. Ustinov (Moscow); 1 male (fig. 3), Primorsky Region, Lazo, 9.5.2001, D. Kochetkov leg. - collection of M. Smirnov (Ivanovo); 28 specimens from the collection of Zoological Museum of Moscow State University (Moscow): 1 ex., Sibir or.; 4 ex., Primorsky Region, The upper reaches of the Lyanchikhe (new name Bogataya) River, 12.5.1947, from A. Kurentsova; 1 ex., Primorsky Region, Shkotovo, V-VI.1932; 1 ex., Suchansk distr., Freyzovka, from T. Samoylov; 2 ex., Ussuri, Ossinovka, 30.5.1917, P. Elsky; ex., Ussuri, Okeanskaia, 4.6.1925; 1 ex., Ussuri, Ossinovka, 7.6.1917, P. Elsky; 1 ex., Ussuri, Ossinovka, 15.6.1917, P. Elsky; 1 ex., Ussuri, Ossinovka, 17.6.1917, P. Elsky; 1 ex., Ussuri, Ossinovka, 1.7.1917, P. Elsky; 1 ex., Ussuri, Ossinovka, 4.7.1917, P. Elsky; 2 ex., Ussuri, Ossinovka, 6.7.1917, P. Elsky; 1 ex., Ussuri, Ossinovka, 8.7.1917, P. Elsky; 1 ex., Ussuri, Ossinovka, 6.9.1917, P. Elsky; 4 ex., Mandshuria, circ. Charbin, Siaolin, 5.5.1940, V. Alin; 1 ex., Japan, Sapporo, Tamanuki; 40 specimens from the collection of M. Danilevsky (Moscow): 1 male, 1 female, Primorsky Region, Gorno-Tayozhnaya Station, 19-20.7.1958, Filippov leg.; 1 male, Vladivostok; 4 males, 1 female, Vladivostok env. 26 km, 6.6.1958, Filippov leg.; 1 female, Primorsky Region, Ugolnaya Station, 27.5.1960, Anufriev leg.; 4 males, 5 females, Suchan [Partizansk] 10.6.1934; 5 males, 4 females, Khabarovsk env., Bychikha, 9.7.1975 ex larv. Aralia, M.Danilevsky leg.; 1 female (fig. 4), Khabarovsk env., Bychikha, 24.5.1976, Aralia, A. Kompantsev leg.; 1 female, Suputinsky Nat. Res. 22.5.1947; 2 males, 1 female, Sakhalin Is., Kholmsk Distr., Kalinino, 24.9.1984, M. Nesterov leg.; 1 female, Kunashir Is., Mendeleevo, 15.9.1976, A. Kompantsev leg.; 1 male, 1 female, Lyanchihe, 12.5.1947; 1 male, 3 females (fig. 9), South Korea (GW), Cheorwon-gun, Munhye-ri, 127.37°E / 38.17°N, 5.4.2010, S.H. Oh leg.; 1 male, Japan, Sapporo, Tamanuki leg.; 12 specimens from the collection of S. Murzin (Moscow): 3 males, 3 female, Primorsky Region, Kamenushka River, 15.6.1990, S. Khvylya leg.; 1 male, Primorsky Region, Suputinsky Nat. Res., 31.5.1969, M. Chernyakhovsky leg.; 3 males, 2 females, Primorsky Region, Osinovka, 17.6.1917, P. Elsky leg.
Figs 1-2. Pogonocherus (s. str.) paradimidiatus sp. n.:
1 - Holotype, female; 2 - Paratype, female, China, Northern Sichuan, WNW Jiuzhalgou, 33°22'26"N, 103°48'08"E, 3320 m, 22.06.2012, I. Kabak leg.
Figs 3-6. Pogonocherus (s. str.) dimidiatus Blessig, 1873: 3 - male, Primorsky Region, Lazo, 9.5.2001, D. Kochetkov leg. (photo by M. Smirnov); 4 - female, Khabarovsk env., Bychikha, 24.5.1976, Aralia, A. Kompantsev leg.; 5 - male, Sichuan, W Heishui, 2500 m, 32°2'47.40"N, 103°1'0.48"E, 3-10.6.2012, S. Murzin leg.; 6 - male, Shaanxi, Zhouzhi, Taibaishan nat. park, 1350 m, 30.5.1999, M. Murzin leg.
Figs 7-9. Pogonocherus (s. str.) dimidiatus Blessig, 1873: 7 - female, S. Gansu, Tochizi, S. Wudu, 2400 m, 21-24.05.1997, S. Murzin leg.; 8 - female, Gansu, SSW Woshar 3000 m 34°30'23"N, 104°49'48"E, 17.06.2005, S. Murzin leg.; 9 - female, South Korea (GW), Cheorwon-gun, Munhye-ri, 127.37°E / 38.17°N, 5.4.2010, S.H. Oh leg.
Pogonocherus (Neopogonocherus subgen. n.)
Type species. Pogonochaerus pillosipes Pic, 1907
Diagnosis. The taxon is characterized by elongated body, elytra without setae tufts; elytral apices without spines, but roundly emarginated with distinct outer angles sharpened; female antennae longer than body; prothorax with big lteral tubercles sharpened apically; elytra without distinct transverse or onlique bands, but with diffused pale pubescent areas. All other known Pogonocherus have more or less distinct elytral setae tufts.
The new taxon includes a single Chinese species P. (N.) pilosipes (Pic, 1907).
Etymology. the name is composed of two words: Latin neo (new) and Pogonocherus; gender masculine.
Pogonocherus (Neopogonocherus) pilosipes (Pic, 1907)
Figs 10-11
Pogonochaerus pillosipes Pic, 1907: 21 [incorrect original spelling] - “Chine orientale”.
Pogonocherus (s. str.) pilosipes, Aurivillius, 1923: 332 - “Ostchina”; Winkler, 1929: 1210 - China orientalis; Hua, 1982: 112; Lin, Tavakilian, 2019: 361 - “China”; Danilevsky, 2020: 448 - “Chine orientale”.
Pogonocherus (s. str.) pillosipes, Plavilstshikov, 1926: 155, 161 - West-China [incorrect original spelling].
Ponogocherus (s. str.) pillosipes, Gressitt, 1951: 516 - “E.China” [misprint in the genus name; incorrect original spelling for species name].
Pogonocherus (Eupogonocherus) pilosipes, Breuning, 1963: 519 - Chine or.; 1975: 28 - Chine orientale.
Pogonocherus pilosipes, Hua, 2002: 225 (attribute to “Pic, 1923”) - China: E. China; Hua et al., 2009: 465 (attribute to “Pic, 1923”); Löbl, Smetana, 2010: 31 - “Chine orientale”.
Remark. Pogonocherus pilosipes (Pic, 1907) was a poorly known species. The holotype was never depicted before. A photo of the holotype-male (fig. 11) from Pic’s collection was kindly sent to me by G. Tavakilian.
Fig. 10. Original description of Pogonochaerus pilosipes Pic, 1907.
Body relatively narrow, attenuated posteriorly, covered by dense, brownish recumbent pubescence, with numerous pale long erected setae; antennae much longer than body, reaching elytral apices by 8th joint; joints 1-3 dark brown, other joints reddish-brown, darkened apically; 4th antennal joint is the longest, longer than 1st; 3rd joint shorter than 1st; 3rd and 4th joints with very long erected pale setae.
Prothorax about as wide anteriorly as posteriorly, about 1.2 times shorter than basal width; with anterior and posterior constrictions; with big lateral tubercles; pronotum with a pair of central carinae; scutellum small, transverse, with pale-grey pubescence.
Elytra about 2 times longer than basal width; with numerous pale erect setae and wide lateral yellowish stripes, without elytral setae tufts (a unique character in the genus); yellowish oblique stripes are situated before middle and near apices; very narrow dark-brown short strokes are distinct at middle; elytral apices with shallow emargination, with distinct outer angles, but without spines.
Legs also with numerous pales, long erect setae, with strongly clavate femora; apical joint of posterior tarsi longer than 2nd-3rd joints combined.
Body length: 7.5 mm.
Material. Holotype, male with a label: “Chine Orient. / (ou Tonkin ?)” - Pic’s collection (Muséum National d'Histoire Naturelle, Paris).
Taxonomy remark. The genus Pogonocherus Dejean, 1821 (type species: Cerambyx hispidus Linnaeus, 1758) was traditionally regarded consisting of two subgenera: the nominative and subgenus Pityphilus Mulsant, 1862 (type species: Cerambyx ovatus Goeze, 1777). Subgenus Pogonocherus s. str. was characterized by elytral epical spines, while in subgenus Pityphilus Mulsant, 1862 elytral apices were rounded or truncted, but without spines. In reality this division was not exact as several species demonstrated intermediate situation and were arbitrary placed to one subgenus or another. For example, P. sturanii Sama, Schurmann, 1982 with emarginated elytral apex and distinct outer apical elytral angle was accepted by Vives, Alonso-Zarazaga (2000) in P. (s. str.), while P. ehdenensis Sama, Rapuzzi, 2000 with about same elytral apex was placed to Pogonocherus (Pityphilus Muls.) by Cocquempot et al. (2016). That is why many of modern authors gave up the divisions of Pogonocherus into subgenera (Sama, 2003; Hasegawa, 2007; Miroshnikov, 2009; Löbl, Smetana, 2010). While others accept two subgenera up to now (Vives, Alonso-Zarazaga, 2000; Vitali et al., 2011; Shapovalov, 2012; Cocquempot et al., 2016; Doychev et al., 2017, Lin, Yang, 2019; Danilevsky, 2020). Sometimes both subgenera were upgraded to genus rank (Villiers, 1978; Bílý, Mehl, 1989).
Here I accept three subgenera; 4 species with truncated (or feebly emarginated) elytral apices must be included in subgenus Pityphilus Mulsant, 1862: P. (Pityphilus) ehdenensis Sama & Rapuzzi, 2000; P. (Pityphilus) ovatoides Rapuzzi & Sama, 2014; P. (Pityphilus) ressli Holzschuh, 1977; P. (Pityphilus) sturanii Sama & Schurmann, 1982.
Note. Wang (2014: 983) used the name “Pogonocherus pilosipes Pic, 1923” for another depicted taxon (without lateral thoracic tubercles), which does not belong to the genus Pogonocherus.
Fig. 11. Pogonocherus (Neopogonocherus) pilosipes (Pic, 1907):
Holotype, male (photo by G. Tavakilian).
Acknowledgements. I am very grateful to my friend S. Murzin (Moscow) for supplying me with the specimens of new species, to A. Gusakov (Zoological Museum of Moscow State University), M. Danilevsky (Moscow), S. Murzin and V. Ustinov (Moscow) for the comparative materials, to G. Tavakilian for the holotype photo of Pogonochaerus pilosipes Pic, 1907, to M. Smirnov (Ivanovo) for exclusevly perfect photo of Pogonocherus dimidiatus, to J. Kurzawa (Tomaszów Mazowiecki) and P. Rapuzzi (Ronchi di Cialla) for valuable consultations.
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Received: 06.12.2020
Accepted: 16.01.2021
